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Jon E. Keeley

Publications and source records attributed to Jon E. Keeley.

At least 73 records · Page 4Linked to original sources
USGS2012

Fire in Mediterranean climate ecosystems-A comparative overview

Four regions of the world share a similar climate and structurally similar plant communities with the Mediterranean Basin. These five areas, known collectively as "mediterranean-type climate (MTC) regions", are dominated by evergreen sclerophyllous-leaved shrublands, semi-deciduous scrub, and woodlands, all of which are prone to widespread crown fires. Summer droughts produce an annual fire hazard that contributes to a highly predictable fire regime. Fire has been an important factor driving the convergence of these systems and is reflected in plant traits such as lignotubers in resprouting shrubs and delayed reproduction that restricts recruitment to a postfire pulse of seedlings. On fertile soils where postfire resprouting is very rapid, opportunities for postfire seedling recruitment are limited and thus these woody taxa have not opted for delaying reproduction. Such fire-independent recruitment is widespread in the floras of MTC regions of the Mediterranean Basin and California and postfire seeding tends to dominate at the more arid end of the gradient. Due to very different geological histories in South Africa and Western Australia, substrates are nutrient poor and thus postfire resprouters do not pose a similar competitive challenge to seedlings and thus postfire seeding is very widespread in these floras. Although circumstantial evidence suggests that the MTC region of Chile had fire-prone landscapes in the Tertiary, these were lost with the late Miocene completion of the Andean uplift, which now blocks summer lightning storms from moving into the region. Today these five regions pose a significant fire management challenge due to the annual fire hazard and metropolitan centers juxtaposed with highly flammable vegetation. This challenge varies across the five MTC landscapes as a function of differences in regional fuel loads and population density.

Israel Journal of Ecology and Evolution↗
USGS2011

The human dimension of fire regimes on Earth

Humans and their ancestors are unique in being a fire‐making species, but ‘natural’ (i.e. independent of humans) fires have an ancient, geological history on Earth. Natural fires have influenced biological evolution and global biogeochemical cycles, making fire integral to the functioning of some biomes. Globally, debate rages about the impact on ecosystems of prehistoric human‐set fires, with views ranging from catastrophic to negligible. Understanding of the diversity of human fire regimes on Earth in the past, present and future remains rudimentary. It remains uncertain how humans have caused a departure from ‘natural’ background levels that vary with climate change. Available evidence shows that modern humans can increase or decrease background levels of natural fire activity by clearing forests, promoting grazing, dispersing plants, altering ignition patterns and actively suppressing fires, thereby causing substantial ecosystem changes and loss of biodiversity. Some of these contemporary fire regimes cause substantial economic disruptions owing to the destruction of infrastructure, degradation of ecosystem services, loss of life, and smoke‐related health effects. These episodic disasters help frame negative public attitudes towards landscape fires, despite the need for burning to sustain some ecosystems. Greenhouse gas‐induced warming and changes in the hydrological cycle may increase the occurrence of large, severe fires, with potentially significant feedbacks to the Earth system. Improved understanding of human fire regimes demands: (1) better data on past and current human influences on fire regimes to enable global comparative analyses, (2) a greater understanding of different cultural traditions of landscape burning and their positive and negative social, economic and ecological effects, and (3) more realistic representations of anthropogenic fire in global vegetation and climate change models. We provide an historical framework to promote understanding of the development and diversification of fire regimes, covering the pre‐human period, human domestication of fire, and the subsequent transition from subsistence agriculture to industrial economies. All of these phases still occur on Earth, providing opportunities for comparative research.

Journal of Biogeography↗
USGS2011

The application of prototype point processes for the summary and description of California wildfires

A method for summarizing repeated realizations of a space‐time marked point process, known as prototyping, is discussed and applied to catalogues of wildfires in California. Prototype summaries are constructed for varying time intervals using California wildfire data from 1990 to 2006. Previous work on prototypes for temporal and space‐time point processes is extended here to include methods for computing prototypes with marks and the incorporation of prototype summaries into hierarchical clustering algorithms, the latter of which is used to delineate fire seasons in California. Other results include summaries of patterns in the spatial‐temporal distribution of wildfires within each wildfire season.

California↗
USGS2011

Long-term effects of prescribed fire on mixed conifer forest structure in the Sierra Nevada, California

The capacity of prescribed fire to restore forest conditions is often judged by changes in forest structure within a few years following burning. However, prescribed fire might have longer-term effects on forest structure, potentially changing treatment assessments. We examined annual changes in forest structure in five 1 ha old-growth plots immediately before prescribed fire and up to eight years after fire at Sequoia National Park, California. Fire-induced declines in stem density (67% average decrease at eight years post-fire) were nonlinear, taking up to eight years to reach a presumed asymptote. Declines in live stem biomass were also nonlinear, but smaller in magnitude (32% average decrease at eight years post-fire) as most large trees survived the fires. The preferential survival of large trees following fire resulted in significant shifts in stem diameter distributions. Mortality rates remained significantly above background rates up to six years after the fires. Prescribed fire did not have a large influence on the representation of dominant species. Fire-caused mortality appeared to be spatially random, and therefore did not generally alter heterogeneous tree spatial patterns. Our results suggest that prescribed fire can bring about substantial changes to forest structure in old-growth mixed conifer forests in the Sierra Nevada, but that long-term observations are needed to fully describe some measures of fire effects.

California↗
USGS2011

A comparison of effects from prescribed fires and wildfires managed for resource objectives in Sequoia and Kings Canyon National Parks

Current goals for prescription burning are focused on measures of fuel consumption and changes in forest density. These benchmarks, however, do not address the extent to which prescription burning meets perceived ecosystem needs of heterogeneity in burning, both for overstory trees and understory herbs and shrubs. There are still questions about how closely prescribed fires mimic these patterns compared to natural wildfires. This study compared burn patterns of prescribed fires and managed unplanned wildfires to understand how the differing burning regimes affect ecosystem properties. Measures of forest structure and fire severity were sampled in three recent prescribed fires and three wildfires managed for resource objectives in Sequoia and Kings Canyon National Parks. Fine scale patterns of fire severity and heterogeneity were compared between fire types using ground-based measures of fire effects on fuels and overstory and understory vegetation. Prescribed fires and wildfires managed for resource objectives displayed similar patterns of overstory and understory fire severity, heterogeneity, and seedling and sapling survival. Variation among plots within the same fire was always greater than between fire types. Prescribed fires can provide burned landscapes that approximate natural fires in many ways. It is recognized that constraints placed on when wildfires managed for resource objectives are allowed to burn freely may bias the range of conditions that might have been experienced under more natural conditions. Therefore they may not exactly mimic natural wildfires. Overall, the similarity in fire effects that we observed between prescribed fires and managed wildfires indicate that despite the restrictions that are often placed on prescribed fires, they appear to be creating post-fire conditions that approximate natural fires when assessed on a fine spatial scale.

California↗
USGS2011

Comparing the role of fuel breaks across southern California national forests

Fuel treatment of wildland vegetation is the primary approach advocated for mitigating fire risk at the wildland–urban interface (WUI), but little systematic research has been conducted to understand what role fuel treatments play in controlling large fires, which factors influence this role, or how the role of fuel treatments may vary over space and time. We assembled a spatial database of fuel breaks and fires from the last 30 years in four southern California national forests to better understand which factors are consistently important for fuel breaks in the control of large fires. We also explored which landscape features influence where fires and fuel breaks are most likely to intersect. The relative importance of significant factors explaining fuel break outcome and number of fire and fuel break intersections varied among the forests, which reflects high levels of regional landscape diversity. Nevertheless, several factors were consistently important across all the forests. In general, fuel breaks played an important role in controlling large fires only when they facilitated fire management, primarily by providing access for firefighting activities. Fire weather and fuel break maintenance were also consistently important. Models and maps predicting where fuel breaks and fires are most likely to intersect performed well in the regions where the models were developed, but these models did not extend well to other regions, reflecting how the environmental controls of fire regimes vary even within a single ecoregion. Nevertheless, similar mapping methods could be adopted in different landscapes to help with strategic location of fuel breaks. Strategic location of fuel breaks should also account for access points near communities, where fire protection is most important.

California↗
USGS2011

Fire and vegetation shifts in the Americas at the vanguard of Paleoindian migration

Across North and South America, the final millennia of the Pleistocene saw dramatic changes in climate, vegetation, fauna, fire regime, and other local and regional paleo-environmental characteristics. Rapid climate shifts following the Last Glacial Maximum (LGM) exerted a first-order influence, but abrupt postglacial shifts in vegetation composition, vegetation structure, and fire regime also coincided with human arrival and transformative faunal extinctions in the Americas. We propose a model of post-glacial vegetation change in response to climatic drivers, punctuated by local fire regime shifts in response to megaherbivore-driven fuel changes and anthropogenic ignitions. The abrupt appearance of humans, disappearance of megaherbivores, and resulting changes in New World fire systems were transformative events that should not be dismissed in favor of climate-only interpretations of post-glacial paleo-environmental shifts in the Americas. Fire is a mechanism by which small human populations can have broad impacts, and growing evidence suggests that early anthropogenic influences on regional, even global, paleo-environments should be tested alongside other potential causal mechanisms.

Quaternary Science Reviews↗
USGS2011

Fire as an evolutionary pressure shaping plant traits

Traits, such as resprouting, serotiny and germination by heat and smoke, are adaptive in fire-prone environments. However, plants are not adapted to fire per se but to fire regimes. Species can be threatened when humans alter the regime, often by increasing or decreasing fire frequency. Fire-adaptive traits are potentially the result of different evolutionary pathways. Distinguishing between traits that are adaptations originating in response to fire or exaptations originating in response to other factors might not always be possible. However, fire has been a factor throughout the history of land-plant evolution and is not strictly a Neogene phenomenon. Mesozoic fossils show evidence of fire-adaptive traits and, in some lineages, these might have persisted to the present as fire adaptations.

Trends in Plant Science↗
USGS2011

Fire and the fire regime framework

A global view of potential vs. actual vegetation distributions points to fire as a major driver of biome distribution and determinant of community structure (Bond et al . 2005). In ecological terms, fire acts much like an herbivore, consuming biomass and competing with biotic consumers for resources, and in this sense is an important part of trophic ecology (Bond & Keeley 2005). As in other competitive interactions, not only can fire competitively exclude herbivores by temporarily eliminating resources, but intensive grazing is known to exclude fire by consuming herbaceous ground fuels (Savage & Swetnam 1990). Coexistence is often enhanced by temporal separation of trophic niches, with herbivores grazing early in the spring on green herbaceous material that is unavailable for burning, whereas later in the season the remaining dry thatch is readily consumed by fire. In many respects fire is a more potent competitor because it is not limited by either toxins or protein deficiency and readily consumes dead woody biomass, but by contrast it is often limited by ignition sources and continuity of fuels. Fire scientists have long symbolized the critical elements of fire in a triangle of fuel, oxygen and heat (Pyne et al . 1996). These are indeed necessary for fire ignition and propagation but are insufficient for predicting the global distribution of fire-prone ecosystems. The conditions both necessary and sufficient to explain the ecological distribution of fire activity can be summarized by four parameters: biomass, seasonality, ignitions and fuel structure (Fig. 2.1). In addition to biomass fuels to spread a fire there must be a dry season that converts potential fuels to available fuels. In mediterranean-type climate (MTC) ecosystems summer drought results in high fire hazard on an annual basis, in contrast to many temperate forests that are only periodically vulnerable to fire in response to decadal or longer oscillations in climate. Vegetation only burns when ignitions are present to initiate the combustion process and landscapes vary markedly in the potential for natural ignitions from lightning, and in the extent of anthropogenic ignition sources. However, understanding the ecosystem distribution of fire requires consideration of a fourth parameter, fuel structure, which is fundamental to recognizing how different fire regimes develop.

Book chapter↗
USGS2011

Fire-related plant traits

As illustrated in Fig. 2.1 there are four environmental parameters that are necessary to determine the distribution of fire-prone ecosystems. However, they are insufficient to predict ecosystem responses to fire without a detailed understanding of the fire regime (see Fig. 2.7). Different fire regimes have very different potentials for recovery and place very different premiums on specific plant traits. For example, those traits contributing to the persistence of species in crown fire regimes will often be very different from those in surface fire regimes. In short, organisms are not adapted to fire per se , but rather to a particular fire regime. Plant traits that are adaptive in fire-prone environments are discussed here. The evolution of such traits and the extent to which they represent adaptations to fire are considered in Chapter 9. Plant populations exhibit four modes of recovery following fire: endogenous regeneration from resprouts or fire-triggered seedling recruitment, delayed seedling recruitment from postfire resprout seed production, delayed seedling recruitment from in situ surviving parent plants, or colonization from unburned metapopulations.

Book chapter↗
USGS2011

Fire in the Mediterranean Basin

The Mediterranean Basin is a meeting point of three continents, Europe, Asia and Africa, and this is responsible for the great diversity of plants, animals and cultures that formed the cradle of Western civilization. It is considered one of the biodiversity hotspots (Myers et al . 2000) because of its high species richness and high proportion of endemisms (Thompson 2005). The total area showing a Mediterranean-type climate (MTC) is about 2.3 million km 2 , with transitions toward temperate forest ecosystems (in the European mountains) and toward arid ecosystems (in North Africa and the Near East). It is not only the largest of the five MTC regions, but also the most geographically complex (with more than 40 000 km of rough coast in different peninsulas and islands) as well as the most socio-economically, culturally and politically varied. Elevations range up to 3756 m in the east (the highest peak in the Taurus mountains, Turkey) and up to 4167 m in the west (the highest peak in the Atlas mountains, Morocco). There are many volcanoes in Italy and the Aegean Islands, with frequent minor eruptions and rare major explosions. The MTC region of the basin corresponds to a narrow rim around the Mediterranean Sea (Fig. 4.1), and includes: (1) in southern Europe, most of the Iberian peninsula (Portugal and Spain), south of France, most of Italy and Greece, the coast of Croatia, Montenegro and Albania; (2) in southwest Asia (the Near East), Cyprus, Lebanon, Palestine, Israel, most of Turkey, and the coast of Syria; and (3) in North Africa (the Magreb), the north of Tunisia, Algeria, Morocco and small coastal areas of Libya. It also includes all the islands in the Mediterranean Sea. In general terms, summers are hot and dry and winters are mild and relatively wet; winters may be cold in the interior areas with a continental climate influence (e.g. central Spain and central Turkey). The configuration of seas, peninsulas and islands, and the topographic complexity of the area, produce a great regional variety of weather and climate. Rainfall ranges from semi-arid conditions (<300 mm) up to over 2000 mm, and peaks in autumn and spring (in the west) and in autumn and winter (in the east). Because of the air masses' trajectories, the wettest parts of the basin are typically the western parts of the peninsulas (Iberian, Italian and Balkan peninsulas). There are also clear gradients from the colder and wetter northwest (southern France and northern Iberia) to the hotter and more arid south and southeast parts of the basin (North Africa and the Near East). The temperature-moderating effect of the sea is highest in the west (Atlantic coast) and lessens toward the east (water temperatures rise from west to east).

Book chapter↗
USGS2011

Fire in California

On the west coast of North America lies the state of California, USA (Fig. 5.1), the bulk of which is dominated by a mediterranean-type climate (MTC). Elevations range from sea level to over 4000 m. Mountain ranges are largely oriented north to south with a major valley between the coastal ranges and the interior Sierra Nevada range. In the rain shadow east of the interior mountain ranges the climate is more continental with much colder winters and increasing proportion of summer precipitation eastward. This easternmost part of the state has steppe climates in the northern portion and desert climates in the south. In Arizona and a few other parts of southwestern USA and northeastern Mexico are disjunct patches of sclerophyllous-leaved vegetation that closely resembles California MTC vegetation. These include evergreen shrublands, broadleaf woodlands and conifer forests and represent mediterranean-type vegetation (MTV) under non-MTCs. Further east at similar latitudes but under different climates are sclerophyll forests with many similarities to MTC conifer forests. The California Floristic Province (Raven & Axelrod 1978) essentially circumscribes the MTC vegetation of North America and extends across the latitudinal range of the state. On the western slopes of the major mountain ranges is a rich diversity of vegetation types that change along the elevational gradient. Ascending the coastal mountains the main vegetation types sort out along gradients of decreasing aridity in the following order: grasslands, semi-deciduous woody sage scrub, evergreen chaparral shrublands, oak woodlands and conifer forests. A similar pattern is evident on the west side of the interior Sierra Nevada except for the absence of sage scrub. These vegetation types exhibit marked differences in fire regime and tolerance to disturbance tied to the different patterns of fuel structure resulting from changes in dominant growth forms along the elevational gradient. Along this gradient there is an interaction between fires and aridity such that lower fire frequency is required to displace shrubland associations with grasslands and other herbaceous vegetation on xeric than on mesic landscapes (Keeley 2002b). Consequently there are complex local mosaics due to differences in aspect and fire history (see Fig. 1.6c).

California↗
USGS2011

Fire in Chile

The mediterranean-type climate (MTC) in Chile (Fig. 6.1) is distributed from La Serena (30° S; Región IV, see Appendix 6.1) in the north to Concepción (37° S; Región X) in the south. It is constrained to the west side of the Andean mountain range, although as the height of this range decreases in the south, a MTC is observed at least as far eastward as Bariloche, Argentina. Although a pattern of winter rains and summer droughts extends northward into the Atacama Desert, this area falls outside our definition of MTC because winter evaporation exceeds rainfall in these areas of extremely low precipitation. The northern border of the MTC region is the transition from desert communities to shrubby matorral, while the southern border corresponds to the point of transition from sclerophyll woodlands to Valdivian evergreen forests (Gajardo 1994; Amigo & Ramirez 1998; Rundel et al . 2007). The landforms of central Chile can be divided into three north–south trending geomorphic zones (Fig. 6.1): the Coastal Cordillera, the Central Valley, and the high Andean Cordillera (Armesto et al . 2007). The Coastal Cordillera rises relatively sharply from the coast, with little extent of coastal terraces, and reaches elevations as high as 2222 m at Cerro Roble and 1880 m at Cerro Campana, which lie between Valparaíso (Región V) on the coast and Santiago (Región Metropolitana) at the base of the Andes. The Central Valley is a structural basin filled to great depth by sediments from the surrounding mountains. North of Santiago, spurs from the Andes extend west across the valley and connect with the Coastal Cordillera, separating individual river basins such as that of the Río Aconcagua (Región V). From Santiago to the south, however, the valley extends uninterrupted for a distance of 900 km to Puerto Montt (Región X), with typical elevations of 400–700 m. The Andean Cordillera marks the eastern boundary of the MTC zone of central Chile. It is the product of complex tectonic activity beginning in the Cretaceous (see Fig. 9.1), but with major uplift in the late Tertiary. Paleobotanical evidence suggests that the central Andes had not attained more than half their current elevation by 10 Ma (Gregory-Wodzicki 2000), but was close to the present height by the end of the Miocene (Reynolds et al . 1990). Elevations reach from 4000 m to nearly 7000 m. To the north of Santiago the Andes are largely composed of metamorphosed sedimentary rock, but a major volcanic zone extends from south of Santiago through the Lake District (Los Lagos Región X).

Book chapter↗
USGS2011

Fire in the Cape Region of South Africa

South Africa's mediterranean-type climate (MTC) region is the smallest of the five MTC regions, centered in the southwestern corner of the Western Cape Province (Fig. 7.1). This Cape region is dominated by fynbos shrublands (see Fig. 1.6e) but this fynbos biome continues eastward far outside the MTC. The Cape region is unusual in that shrublands dominate under climate regimes that also support forests. Entire landscapes can support alternative ecosystem states. Even the semi-arid areas can support entirely different vegetation: fire-prone shrublands or fire-resistant broadleaf thickets. Perhaps more than any other MTC region, fire plays a central role in determining major vegetation patterns of winter rainfall regions of South Africa. Soils are also thought to be of major importance since much of the Cape's MTC region is on nutrient-poor sandy soils (see Fig. 1.5). A complex interplay between soils, fire and climate and, in the east, large mammal herbivory, determines boundaries of major biomes. The Cape Floristic Region is extremely rich in species with very high levels of endemism (Linder 2003). It is the world's richest temperate flora and is largely restricted to fire-prone ecosystems (Cowling et al . 1996; Linder 2003). So, contrary to the widely held popular belief that fires are an anthropogenic disturbance (e.g. Pillans 1924; Axelrod 1980), or merely incidental to this formation (Hopper 2009), a rich endemic flora has evolved in the Cape whose members are overwhelmingly fire dependent, implying a long history of natural fires as a selective force. Major Vegetation Patterns This chapter discusses fire regimes in the Cape region, what little is known of their determinants, and how they influence major vegetation patterns in the region. Though a large number of studies have explored plant responses to fire (reviewed by Bond 1997; Cowling et al . 1997a), these are heavily biased toward fynbos shrublands, the dominant vegetation cover of the region (Fig. 7.2). Fire responses of species belonging to other vegetation types are poorly known. Yet the existence of these other vegetation types is one of the central conundrums of the Cape region. It implies failure of climate alone to explain apparent convergence with other MTC regions (Chapter 1). For example, low shrublands would be expected in deserts replaced, as rainfall progressively increases, by taller shrublands, woodlands and then forests. But this is clearly not the case in the Cape region. The dominant fynbos vegetation shows very little variation in aboveground biomass from arid desert fringes (mean annual precipitation ~250 mm) to rain-drenched high-altitude heathlands (> 3000 mm) (Fig. 7.3). Yet across the entire rainfall gradient fynbos co-occurs with alternative ecosystems with much greater woody biomass. These broadleaf thickets and forests have an entirely different floristic and functional composition and often are restricted to isolated fire-protected refugia (Fig. 7.2; Taylor 1978; Kruger 1979; Cowling et al . 2005; Rebelo et al . 2006). The implication is that apparent convergence of shrubby fynbos growth forms with other MTC plant communities cannot be understood in terms of climate alone and that one needs to think in terms of the climate, fire, geology filter (see Fig. 1.4).

Cape region↗
USGS2011

Fire in southern Australia

The mediterranean-type climate (MTC) in Australia spans from the southwestern part of Western Australia to include much of South Australia and western Victoria (Fig. 8.1), which covers a longitudinal distance second only to the Mediterranean Basin MTC region. As in other MTC regions, the highly fire-prone evergreen sclerophyllous shrub and tree mediterranean-type vegetation (MTV) extends much further east and north into climatic zones that are not MTC. Australia, however, is distinctly unlike other MTC regions in that fire-prone MTV is extensive across the southern part of the continent and transcends climatic boundaries with relatively subtle changes in community structure and composition. Sclerophyllous MTV dominates both the MTC region of the southwestern corner of the continent as well as the southeastern corner under an aseasonal climate. Both regions share a common fire season of summer to early autumn (McArthur 1972); however, the MTC southwest has a potential fire season every summer whereas in the southeast it is tied to weather anomalies that occur once to several times a decade. Mediterranean-type Vegetation Within the southern Australian MTC zone (Fig. 8.1) evergreen sclerophyllous vegetation dominates. Such MTV is sometimes defined as shrub dominated (Specht 1979), and indeed large areas of sclerophyllous heaths (see Fig. 1.6f), shrublands (Fig. 8.2a) and mallee (Fig. 8.3) occur. However, woodlands and forests form integral parts of the MTC biome (Dell et al . 1989; Gill 1994), and thus MTV includes shrublands, woodlands and forests, and in southern Australia they dominate both in the MTC region and outside that climatic zone (Fig. 8.1). MTV is found across the southern temperate latitudes of Australia (Table 8.1) in an arc below about 30° latitude, accounting for dominant vegetation types in infertile habitats throughout temperate Australia. We specifically focus on the various heaths, shrublands and dry sclerophyll forests that constitute the most fire prone communities in these temperate landscapes. Although similar fire-prone MTV heathlands occur extensively within the tropics on the northern end of the continent (Keith et al . 2002; Russell-Smith & Stanton 2002), here we focus on the temperate MTV, but do consider broader relationships with other adjoining vegetation types (e.g. wet sclerophyll forests, rainforests, and various arid and semi-arid woodlands and shrublands).

South Australia↗