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Arctic Refuge coastal plain terrestrial wildlife research summaries
In 1980, when the U.S. Congress enacted the Alaska National Interest Lands Conservation Act (ANILCA), it also mandated a study of the coastal plain of the Arctic National Wildlife Refuge. Section 1002 of ANILCA stated that a comprehensive inventory of fish and wildlife resources would be conducted on 1.5 million acres of the Arctic Refuge coastal plain (1002 Area). Potential petroleum reserves in the 1002 Area were also to be evaluated from surface geological studies and seismic exploration surveys. Results of these studies and recommendations for future management of the Arctic Refuge coastal plain were to be prepared in a report to Congress. In 1987, the Department of the Interior published the Arctic National Wildlife Refuge, Alaska, Coastal Plain Resource Assessment - Report and Recommendations to the Congress of the United States and Final Environmental Impact Statement. This report to Congress identified the potential for oil and gas production (updated* most recently by the U.S. Geological Survey in 2001), described the biological resources, and evaluated the potential adverse effects to fish and wildlife resources. The 1987 report analyzed the potential environmental consequences of five management alternatives for the coastal plain, ranging from wilderness designation to opening the entire area to lease for oil and gas developement. The report's summary recommended opening the 1002 Area to an orderly oil and gas leasing program, but cautioned that adverse effects to some wildlife populations were possible. Congress did not act on this recommendation nor any other alternative for the 1002 Area, and scientists continued studies of key wildlife species and habitats on the coastal plain of the Arctic Refuge and surrounding areas. This report contains updated summaries of those scientific investigations of caribou, muskoxen, predators (grizzly bears, wolves, golden eagles), polar bears, snow geese, and their wildlife habitats. Contributions to this report were made by scientists affiliated with the U.S. Geological Survey; U.S. Fish and Wildlife Service; Alaska Department of Fish and Game; University of Alaska-Fairbanks; Canadian Wildlife Service; Yukon Department of Renewable Resources; and the Northwest Territories Department of Resources, Wildlife, and Economic Development. Sections of the report presenting new information on caribou and forage plants were peer-reviewed by three independent, non-affiliated scientists. The remaining sections summarize previously published peer-reviewed scientific papers and were reviewed by a single independent scientist. The U.S. Geological Survey and the U.S. Fish and Wildlife Service collaborated in the publication of this report.
Community Structure and Trophic Ecology of Demersal Fishes on the Pinnacles Reef Tract, Final Synthesis Report
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Introduction
The Arctic National Wildlife Refuge in northeastern Alaska is one of 16 refuges in Alaska and 539 refuges nationwide within the National Wildlife Refuge System administered by the U.S. Fish and Wildlife Service. First established as the Arctic National Wildlife Range in 1960 by Public Land Order 2214, it initially had a three-fold purpose to preserve unique wildlife, wilderness, and recreation values on 8.9 million acres. In 1980, the Arctic National Wildlife Range was expanded to the southwest and renamed the Arctic National Wildlife Refuge (also called the Arctic Refuge in this report) when the U.S. Congress passed the Alaska National Interest Lands Conservation Act (ANILCA), Public Law 96-487 (94 Stat. 2371). This legislation also designated almost all of the original Arctic National Wildlife Range as wilderness, and it directed the Secretary of the Interior to conduct studies evaluating both the biological resources and the potential petroleum reserves of 1.5 million acres (titled the 1002 Area) on the coastal plain of the Arctic Refuge. In April 1982, the Arctic Refuge staff completed a report summarizing the then current state of knowledge on the fish, wildlife, and their habitats present on the coastal plain of the Arctic Refuge (U.S. Fish and Wildlife Service 1982). From 1982 to 1985, field investigations of biological resources of the 1002 Area were carried out by a number of investigators, and annual reports summarized the results (Garner and Reynolds 1983, 1984, 1985, 1986, 1987). These reports and other resources were used to prepare a Department of the Interior report to Congress: Arctic National Wildlife Refuge, Alaska Coastal Plain Resource Assessment - Report and Recommendation to the Congress of the United States and Final Environmental Impact Satement (Clough et al., 1987).
Cumulative index to chemicals and to common and scientific names of species listed in Contaminant Hazard Reviews 1 through 34
The Contaminant Hazard Review (CHR) series--sponsored by the U.S. Geological Survey, Patuxent Wildlife Research Center--synthesizes ecotoxicological data for selected environmental contaminants, with emphasis on hazards to native species of flora and fauna. From 1985 through 1998, 34 reviews were published in various report series of the U.S. Department of the Interior on agricultural pesticides (acrolein, atrazine, carbofuran, chlordane, chlorpyrifos, diazinon, diflubenzuron, famphur, fenvalerate, mirex, paraquat, toxaphene), metals and metalloids (arsenic, boron, cadmium, chromium, copper, lead, mercury, molybdenum, nickel, selenium, silver, tin, zinc), mammalian biocides (sodium monofluoroacetate), organic industrial and municipal wastes (dioxins, pentachlorophenol, polycyclic aromatic hydrocarbons, polychlorinated biphenyls), minin wastes (cyanide), and ionizing radiations. This current report is a cumulative index to the common and scientific names of all biological species listed in the first 34 reports in the CHR series, with individual species cross-referenced with contaminant hazard review and corresponding page numbers. A similar index for chemicals is included.
Coefficients of productivity for Yellowstone's grizzly bear habitat
This report describes methods for calculating coefficients used to depict habitat productivity for grizzly bears in the Yellowstone ecosystem. Calculations based on these coefficients are used in the Yellowstone Grizzly Bear Cumulative Effects Model to map the distribution of habitat productivity and account for the impacts of human facilities. The coefficients of habitat productivity incorporate detailed information that was collected over a 20-year period (1977-96) on the foraging behavior of Yellowstone's bears and include records of what bears were feeding on, when and where they fed, the extent of that feeding activity, and relative measures of the quantity consumed. The coefficients also incorporate information, collected primarily from 1986 to 1992, on the nutrient content of foods that were consumed, their digestibility, characteristic bite sizes, and the energy required to extract and handle each food. Coefficients were calculated for different time periods and different habitat types, specific to different parts of the Yellowstone ecosystem. Stratifications included four seasons of bear activity (spring, estrus, early hyperphagia, late hyperphagia), years when ungulate carrion and whitebark pine seed crops were abundant versus not, areas adjacent to (< 100 m) or far away from forest/nonforest edges, and areas inside or outside of ungulate winter ranges. Densities of bear activity in each region, habitat type, and time period were incorporated into calculations, controlling for the effects of proximity to human facilities. The coefficients described in this report and associated estimates of grizzly bear habitat productivity are unique among many efforts to model the conditions of bear habitat because calculations include information on energetics derived from the observed behavior of radio-marked bears.
Biogeographical profiles of shorebird migration in midcontinental North America
The biogeographic information described here will help identify the uniqueness of different regions of the plains to migrating shorebirds. Although shorebirds migrating along Atlantic and Pacific coastal areas are capable of long jumps between refueling stops, there is evidence that some species move short rather than long distances between refueling sites. Maps of distribution patterns and chronology accounts can lend insight towards understanding migration strategies of the different shorebird species. This report focuses on the distribution patterns of enroute migrants that refuel in interior wetlands during migration. We provide information on the spatial and temporal occurrence and habitat requirements for individual species and groups of species with the intent that this information be used in guiding management efforts.
Mississippi/Alabama pinnacle trend ecosystem monitoring Final Synthesis Report
This Final Synthesis Report summarizes a four-year program to characterize and monitor carbonate mounds on the Mississippi/Alabama outer continental shelf (OCS). The study area is shown in Fig.ES.1. The study was conducted by Continental Shelf Associates, Inc. and the Geochemical and Environmental Research Group (GERG) of Texas A&M University (TAMU), for the U.S. Geological Survey (USGS), Biological Resources Division.
Biomonitoring of Environmental Status and Trends (BEST) Program: Environmental contaminants and their effects on fish in the Mississippi River Basin
We collected, examined, and analyzed 1378 fish of 22 species from 47 sites in the Mississippi River basin (MRB) during 1995 and from a reference site in 1996. The sampling sites in the MRB represented National Contaminant Biomonitoring Program (NCBP) stations situated at key points on major rivers and National Water- Quality Assessment Program (NAWQA) stations located on lower-order rivers and streams in the Eastern Iowa Basins (EIB) and Mississippi Embayment (MSE) Study Units. The reference site was the water supply system of the USGS-Leetown Science Center in rural Jefferson County, WV. Common carp (Cyprinus carpio; carp) and black basses (Micropterus spp.; bass), the targeted species, together represented 82% of the fish collected. Each fish was examined in the field for externally and internally visible gross lesions, selected organs were weighed to compute various ponderal and organo-somatic indices, and selected tissues and fluids were obtained and preserved for analysis of biomarkers. Fish health indicators included splenic macrophage aggregates, lysozyme activity, and hispathological analysis of liver, kidney, and other tissues. Reproductive biomarkers included analysis of plasma concentrations of vitellogenin (vtg) and the sex steroid hormones 17-estradiol (E2) and 11-ketotestosterone (11- kt); and the histological determination of percent oocyte atresia (in female fish) and gonadal stage. Hepatic ethoxyresorufin O-deethylase (EROD) activity was also measured. Composite samples of whole fish from each station were grouped by species and gender and analyzed for persistent organochlorine and elemental contaminants and for dioxin-like activity (TCDD-EQ) using the H4IIE rat hepatoma cell bioassay. Organochlorine and inorganic contaminant concentrations in fish were generally low relative to historical levels at most sites, but remained present at concentrations representing threats to piscivorous wildlife in some locations. Toxaphene and DDT (mostly as p,p?-DDE) concentrations remained elevated in fish from the cottongrowing regions of the lower Mississippi valley, and were generally greater in the smaller streams draining agricultural areas (that is, in the MSE Study Unit) than at large river sites. Cyclodiene pesticide concentrations were also greatest in the EIB Study Unit and elsewhere in the corn-growing regions of the mid-MRB. Former point-sources of organochlorine pesticides also remained evident, especially in the Mississippi River near Memphis, TN. Consistent with previous findings, total PCB concentrations tended to be greatest (1-3 g/g) in the industrialized and urbanized Ohio River and Upper Mississippi sub-basins and at Memphis, TN, and were generally correlated with TCDD-EQ and EROD activity. Conversely, PCB concentrations were low (<0.1 g/g) in the more agricultural parts of the MRB. Concentrations of inorganic contaminants were also relatively low and stable or declining relative to past levels at most sites. Exceptions were Hg and Se; Hg concentrations were slightly elevated (>0.3 g/g) in bass from the Mississippi River at Memphis and several other sites and in carp from one MSE site. Concentrations of Se were also great enough to constitute a hazard to piscivorous wildlife (>0.6 g/g) at several MRB sites in the western parts of the MRB and were especially high (4-5 g/g) in fish from John Martin Reservoir, CO, where elevated concentrations were reported previously. Biomarker results indicated that fish from many stations had been exposed to contaminants, but at no sites did findings indicate exposure to high concentrations of toxic chemicals. Noteworthy among biomarker findings was that 73% of the male smallmouth bass (Micropterus dolomieui) from the Mississippi River at Lake City, MN (Lake Pepin) were intersex as indicated by the histological detection of ovotestes; and the combined EROD and H4IIE results indicated that fish from several rural sites in the
Wetland restoration in the Prairie Pothole Region of North America: A literature review
The landscape of the prairie pothole region (PPR), a grassland biome of the northern U.S. Great Plains and parts of Canada, has been greatly altered by land use since the 1800's. Conversion of grassland to cropland and drainage of wetlands has resulted in wetland losses of up to 90% in some areas. Besides the area providing critical habitat to various wildlife, breeding waterfowl, and migratory birds, its seasonal wetlands support diverse plant and invertebrate communities, play a role in flood attenuation, act as traps for nutrients. store and recharge groundwater. and are valued recreational lands. Most of the restoration of prairie potholes has only occurred since the 1980's, with few follow-up studies performed and little postrestoration monitoring of these restorations. Monitoring and research of wetland restoration in the PPR is less common relative to the number of postrestoration studies done on other wetland types in the United States. This report is a synthesis of current knowledge of restored prairie pothole wetlands and makes suggestions for future wetland restoration-related research. In order to determine the benefits of restored wetlands, it is important to better understand how closely restored wetlands in the PPR resemble their natural analogues in terms of functions and values. The report categorizes PPR literature into five general sections: wildlife, vegetation, invertebrates, fish, and physical and chemical characteristics of restored wetlands. Each of these five sections has a summary of research and is divided into two parts: an overview of research and findings and regional case studies. Most PPR studies have focused on bird and plant communities, whereas research done on the functions of restored wetlands and studies concerning less visible fauna and physical and chemical characteristics are scarce. In addition, there is a scarcity of research in the western and northern portions or the PPR; most studio to date have been conducted in Iowa. Minnesota, or South Dakota. Key Words: wetland restoration, prairie pothole region, PPR, literature review, wetland functions and values. postrestoration studies
A national survey of Conservation Reserve Program (CRP) participants on environmental effects, wildlife issues, and vegetation management on program lands
A national survey of Conservation Reserve Program (CRP) contractees was completed to obtain information about Abstract environmental and social effects of the program on participants, farms, and communities. Of interest were observations concerning wildlife, attitudes about long-term management of program lands, and effectiveness of U.S. Department of Agriculture (USDA) assistance in relation to these issues. Surveys were delivered to 2,189 CRP participants with a resultant response rate of 64.5%. Retired farmers represented the largest category of respondents (52%). Enhanced control of soil erosion was the leading benefit of the CRP reported. Over 73% of respondents observed increased numbers of wildlife associated with lands enrolled in the program. The majority of respondents reported CRP benefits, including increased quality of surface and ground waters, improved air quality, control of drifting snow, and elevated opportunities to hunt or simply observe wildlife as part of daily activities. Income stability, improved scenic quality of farms and landscapes, and potential increases in property values and future incomes also were seen as program benefits. Negative aspects, reported by a smaller number of respondents, included seeing the CRP as a source of weeds, fire hazard, and attracting unwanted requests for trespass. Over 75% of respondents believed CRP benefits to wildlife were important. A majority of respondents (82%) believed the amount of assistance furnished by USDA related to planning and maintaining wildlife habitat associated with CRP lands was appropriate. Nearly 51% of respondents would accept incorporation of periodic management of vegetation into long-term management of CRP lands to maintain quality of wildlife habitats. Provision of funds to address additional costs and changes in CRP regulations would be required to maximize long-term management of program lands. Additional, on-ground assistance related to management of CRP, and other agricultural lands, to maintain wildlife habitats was commonly identified as a need by survey respondents.
Physical setting: Chapter 1 in Initial biotic survey of Lisbon Bottom, Big Muddy National Fish and Wildlife Refuge
Lisbon Bottom consists of approximately 875 ha of river bottom along the Missouri River in Howard County, Missouri, from approximately river mile (RM) 213 to RM 219. As used in this report, the Lisbon Bottom area also includes the main channel of the Missouri River adjacent to the Lisbon Bottom and portions of the adjacent flood plain in Saline County, Missouri. Before regulation and structuring of the Missouri River, riverine areas like Lisbon Bottom were shifting mosaics of dynamic habitat patches that were created and maintained by hydrologic and geomorphic processes. Flow regulation, navigation structures, and bank-stabilization projects isolated Lisbon Bottom from the river by decreasing the magnitude and frequency with which hydrologic and geomorphic processes could alter habitat characteristics. The flood of 1993 breached agricultural levees around Lisbon Bottom, thereby reestablishing a connection to the Missouri River. Although still affected by flow regulation and the requirement to maintain navigation in the main channel, Lisbon Bottom presents the opportunity to study ecosystem processes and dynamic geomorphology in a setting that more closely mimics the natural riverine system than any other site on the navigable Lower Missouri River. The purpose of this chapter is to describe the physical setting and processes that structure the riverine ecosystem of Lisbon Bottom and similar areas of the Lower Missouri River. The description compiles available information on temporal and spatial changes at the site, including the history of channel changes and hydrology.
Evaluation of Metal Toxicity in Streams Affected by Abandoned Mine Lands, Upper Animas River Watershed, Colorado
Acid drainage from abandoned mines and from naturally-acidic rocks and soil in the upper Animas River watershed of Colorado generates elevated concentrations of acidity and dissolved metals in stream waters and deposition of metal-contaminated particulates in streambed sediments, resulting in both toxicity and habitat degradation for stream biota. High concentrations of iron (Fe), aluminum (Al), zinc (Zn), copper (Cu), cadmium (Cd), and lead (Pb) occur in acid streams draining headwaters of the upper Animas River watershed, and high concentrations of some metals, especially Zn, persist in circumneutral reaches of the Animas River and Mineral Creek, downstream of mixing zones of acid tributaries. Seasonal variation of metal concentrations is reflected in variation in toxicity of stream water. Loadings of dissolved metals to the upper Animas River and tributaries are greatest during summer, during periods of high stream discharge from snowmelt and monsoonal rains, but adverse effects on stream biota may be greater during winter low-flow periods, when stream flows are dominated by inputs of groundwater and contain greatest concentrations of dissolved metals. Fine stream-bed sediments of the upper Animas River watershed also contain elevated concentrations of potentially toxic metals. Greatest sediment metal concentrations occur in the Animas River upstream from Silverton, where there are extensive deposits of mine and mill tailings, and in mixing zones in the Animas River and lower Mineral Creek, where precipitates of Fe and Al oxides also contain high concentrations of other metals. This report summarizes the findings of a series of toxicity studies in streams of the upper Animas River watershed, conducted on-site and in the laboratory between 1998 and 2000. The objectives of these studies were: (1) to determine the relative toxicity of stream water and fine stream-bed sediments to fish and invertebrates; (2) to determine the seasonal range of toxicity in stream water; (3) to develop site-specific thresholds for toxicity of Zn and Cu in stream water; and (4) to develop models of the contributions of Cu and Zn to toxicity of stream water, which may be used to characterize toxicity before and after planned remediation efforts. We evaluated the toxicity of metal-contaminated sediments by conducting sediment toxicity tests with two species of benthic invertebrates, the midge, Chironomus tentans. and the amphipod, Hyalella azteca. Laboratory toxicity tests with both taxa, exposed to fine stream-bed sediments collected in September 1997, showed some evidence of sediment toxicity, as survival of midge larvae in sediments from Cement Creek (C48) and lower Mineral Creek (M34), and growth of amphipods in sediments from these sites and three Animas River sites (A68, Animas at Silverton; A72, Animas below Silverton, and A73, Animas at Elk Park) were significantly reduced compared to a reference site, South Mineral Creek (SMC) . Amphipods were also exposed to site water and fine stream-bed sediment, separately and in combination, during the late summer low flow period (August-September) of 1998. In these studies, stream water, with no sediment present, from all five sites tested (same sites as above, except C48) caused 90% to 100% mortality of amphipods. In contrast, significant reductions in survival of amphipods occurred at two sites (A72 and SMC) in exposures with field-collected sediment plus stream water, and at only one site (A72) in exposures with sediments and clean overlying water. Concentrations of Zn, Pb, Cu, and Cd were high in both sediment and pore water (interstitial water) from most sites tested, but greatest sediment toxicity was apparently associated with greater concentrations of Fe and/or Al in sediments. These results suggest that fine stream-bed sediments of the more contaminated stream reaches of the upper Animas River watershed are toxic to benthic invertebrates, but that these impacts are less serious than tox
Waterfowl migration on Klamath Basin National Wildlife Refuges 1953-2001
The Klamath Basin National Wildlife Refuge (NWR) complex, located in northeastern California and southern Oregon, is situated on a major Pacific Flyway migration corridor connecting waterfowl breeding grounds in the north with major wintering grounds in California and Mexico. The complex comprises five waterfowl refuges including Lower Klamath NWR, Tule Lake NWR, Upper Klamath NWR, Klamath Marsh NWR, and Clear Lake NWR, and one bald eagle refuge, Bear Valley NWR. Lower Klamath and Tule Lake NWRs are the largest refuges in the complex; historically, they supported some of the greatest autumn and spring concentrations of migrating waterfowl in North America. Starting in 1953, standardized waterfowl surveys from small aircraft have been conducted in autumn through spring. This report summarizes waterfowl migration activity (i.e., abundance, species composition, distribution on refuges, and chronology) over four time periods—the long-term (1953-2001), early (1953-76), recent (1977-2001), and the most recent (1998-2001)—to describe changing patterns of migration on Klamath Basin refuges from autumn 1953 to spring 2001. Over the long term, waterfowl abundance (birds per day) on the refuge complex averaged about 1.0 million in autumn and about 360,000 in spring. A record peak count of 5.8 million waterfowl was recorded September 24-25, 1958. Average abundance of autumn staging waterfowl for the refuge complex, after reaching record levels in the 1950s and early 1960s, began a decline that lasted until the 1980s. A gradual recovery occurred during the 1990s, but autumn abundance has not recovered to pre-1970 levels. In contrast to autumn, average spring abundance was generally lower in the early decades but has gradually increased through the 1990s, particularly on Lower Klamath NWR. Dabbling ducks represented an average of 68% of all waterfowl in autumn and 55% in spring for the long term. Northern pintail ( Anas acuta ) was dominant, representing 62% of all dabblers in autumn and 51% in spring. A significant decline in pintail abundance starting in the late 1950s altered waterfowl composition on Klamath Basin refuges. As pintail declined, other species such as mallard ( Anas platyrhynchos ) and green-winged teal ( Anas crecca ) increased in abundance. Although Arctic nesting geese, including white-fronted ( Anser albifrons ), cackling Canada ( Branta canadensis minima ), white geese (lesser snow [ Chen caerulescens caerulescens ], and Ross’s [ Chen rossii ]) have become less prominent in recent decades, they reached an historically high abundance during autumn in the 1960s and 1970s, particularly on Tule Lake NWR. Tule Lake NWR supported the highest average autumn waterfowl populations until surpassed by Lower Klamath NWR around 1980. During the recent period (1977-2001), Lower Klamath NWR accounted for 60% of all waterfowl using the refuge complex in autumn and 61% in spring. Habitat diversity and wetland productivity contributed to its greater waterfowl abundance. Tule Lake NWR supported the most geese over the long term, 79% in autumn and 66% in spring; however, total waterfowl abundance on this refuge in autumn has been in decline, likely because of reduced diversity and productivity of sumps in the refuge. Upper Klamath, Klamath Marsh, and Clear Lake NWRs accounted for less than 8% of total waterfowl use in autumn and spring but provided diverse habitats for migrants. Waterfowl use-days on Klamath Basin refuges typically peaked in mid-autumn, decreased as migrants passed through the basin, and then reached a lesser peak during spring passage. Waterfowl abundance reached a pronounced peak in autumn during the early period (1953-76), but spring peak buildup was much less pronounced. For the recent period the autumn peak was more subdued. Waterfowl abundance, species composition, and distribution on Klamath Basin refuges have fluctuated over the decades and have been influenced by events such as productivity on breeding grounds and habitat conditions on wintering grounds that cause shifts in migration patterns. A major challenge for the future appears to be the availability of adequate water for wetland management on Klamath Basin refuges.
Field guide to malformations of frogs and toads: with radiographic interpretations
In 1995, students found numerous malformed frogs on a field trip to a Minnesota pond. Since that time, reports of malformed frogs have increased dramatically. Malformed frogs have now been reported in 44 states in 38 species of frogs, and 19 species of toads. Estimates as high as 60% of the newly metamorphosed frog populations have had malformations at some ponds (NARCAM, ’99). The wide geographic distribution of malformed frogs and the variety of malformations are a concern to resource managers, research scientists and public health officials. The potential for malformations to serve as a signal of ecosystem disruption, and the affect this potential disruption might have on other organisms that share those ecosystems, has not been resolved. Malformations represent an error that occurred early in development. The event that caused the developmental error is temporally distant from the malformation we see in the fully developed animal. Knowledge of normal developmental principles is necessary to design thoughtful investigations that will define the events involved in abnormal development in wild frog populations. Development begins at the time an egg is fertilized and progresses by chemical communication between cells and cell layers. This communication is programmed through gene expression. Malformations represent primary errors in development, errors in chemical communication or translation of genetic information. Deformations arise later in development and usually result from the influence of mechanical factors (such as amputation) that alter shape or anatomy of a structure that has developed normally. The occurrence and the type of malformations are influenced by the type of error or insult as well as the timing of the error (the developmental stage at which the error occurred). The appearance of the malformation can therefore provide clues that suggest when the error may have occurred. If the malformation is an incomplete organ, such as an incomplete limb, the factor or insult acted during a susceptible period prior to organ completion. Although defining the anatomy of the malformed metamorphosed frog can give us an idea of the approximate window during which the developmental insult was initiated, and might even suggest the type of insult that may have occurred, the morphology of the malformation does not define the cause. To define causes and mechanisms of frog malformations we need to use well designed investigations that are different from traditional tests used in acute toxicity or disease pathogenicity studies. When investigating malformations in metamorphosed frogs, we are looking at the affect of exposure to an agent that occurred early in tadpole development. Therefore investigations to determine causes of malformations need to look at agents that are present in the tadpoles or their environments at these early developmental times. Laboratory experiments need to expose embryos and tadpoles to suspect agents at appropriate developmental stages and look at acute results, such as toxicity and death, as well as following the developmental process to completion to determine the impact of the agent on the developing tadpole and the fully developed frog. This means holding animals past metamorphic climax to assure that the anatomy and physiology of the adult have developed normally. As we look at field collections of abnormal frogs, we need to keep in mind that these collections reflect survivors only. We are looking at malformations that were not fatal to tadpoles. We cannot assume that because we do not collect other malformations, they did not exist. More work needs to be done on the developing tadpole, in the field and in the laboratory, to better elucidate the range, frequency, character and causes of anuran malformations.
Seasonal bathymetric distributions of 16 fishes in Lake Superior, 1958-75
The bathymetric distributions of fishes in Lake Superior, which is one of the largest and deepest lakes in the world, has not been studied on a lakewide scale. Knowledge about the bathymetric distributions will aid in designing fish sampling programs, estimating absolute abundances, and modeling energy flow in the lake. Seasonal bathymetric distributions were determined , by 10-m depth intervals, for 16 fishes collected with bottom trawls and bottom-set gill nets within the upper 150 m of Lake Superior during 1958-75. In spring trawl catches, maximum abundance occurred at these depths: 15 m for round whitefish (Prosopium cylindraceum); 25m for longnose sucker (Catostomus catostomus); 35 m for lake whitefish (Coregonus clupeaformis) and rainbow smelt (Osmerus mordax); 45 m for lake trout (Salvelinus namaycush); 65 m for pygmy whitefish (Prospoium coulteri) and bloater (Coregonus hoyi); 75 m for trout- perch (Percopsis omiscomaycus); 105 m for shortjaw cisco (Coregonus zenithicus); and 115 m for ninespine stickleback (Pungitius pungitius), burbot (Lota lota), slimy sculpin (Cottus cogantus), spoonhead sculpin (Cottus ricei), and deepwater sculpin (Myoxcephalus thompsoni). Bathymetric distributions in spring gill nets were similar to those in trawls, except that depths of maximum abundances in gill nets were shallower than those in trawls for lake trout, rainbow smelt, longnose sucker, and burbot. Lake herring (Coregonus artedi) and kiyi (Coregonus kiyi) were rarely caught in trawls, and their maximum abundances in spring gill net collections were at depths of 25 and 145 m, respectively. In summer, pygmy whitefish, shortjaw cisco, lake herring, kiyi, longnose sucker, burbot, ninespine stickleback, trout-perch, slimy sculpin, and spponhead sculpin were at shallower depths than in spring, whereas rainbow smelt were found in deeper water; there was no change for other species. In fall, shortjaw cisco was at shallower depths than in summer, whereas the remaining species were found deeper, except for lake whitefish and lake trout whose modal depths did not change. Distributions of lake trout and lake whitefish were analyzed by age group, and the young (ages 1-3) of both species were often found in shallower water than were older fish. The shallow-water species exhibited little seasonal changes in bathymetric distributions, whereas the species that inhabited the middepths of deeper water generally moved shallower as the seasons progressed. Most of the more pronounced seasaonl changes in bathymetric distribution were associated with spawning movements.
Polar Bears
Polar bears ( Ursus maritimus ) are hunted throughout most of their range. In addition to hunting polar bears of the Beaufort Sea region are exposed to mineral and petroleum extraction and related human activities such as shipping road-building, and seismic testing (Stirling 1990). Little was known at the start of this project about how polar bears move about in their environment, and although it was understood that many bears travel across political borders, the boundaries of populations had not been delineated (Amstrup 1986, Amstrup et al. 1986, Amstrup and DeMaster 1988, Garner et al. 1994, Amstrup 1995, Amstrup et al. 1995, Amstrup 2000). As human populations increase and demands for polar bears and other arctic resources escalate, managers must know the sizes and distributions of the polar bear populations. Resource managers also need reliable estimates of breeding rates, reproductive intervals, litter sizes, and survival of young and adults. Our objectives for this research were 1) to determine the seasonal and annual movements of polar bears in the Beaufort Sea, 2) to define the boundaries of the population(s) using this region, 3) to determine the size and status of the Beaufort Sea polar bear population, and 4) to establish reproduction and survival rates (Amstrup 2000).
Historical land cover changes in the Great Lakes Region
Two different methods of reconstructing historical vegetation change, drawing on General Land Office (GLO) surveys and fossil pollen deposits, are demonstrated by using data from the Great Lakes region. Both types of data are incorporated into landscape-scale analyses and presented through geographic information systems. Results from the two methods reinforce each other and allow reconstructions of past landscapes at different time scales. Changes to forests of the Great Lakes region during the last 150 years were far greater than the changes recorded over the preceding 1,000 years. Over the last 150 years, the total amount of forested land in the Great Lakes region declined by over 40%, and much of the remaining forest was converted to early successional forest types as a result of extensive logging. These results demonstrate the utility of using GLO survey data in conjunction with other data sources to reconstruct a generalized 'presettlement' condition and assess changes in landcover.