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Bokan Mountain peralkaline granitic complex, Alexander terrane (southeastern Alaska): evidence for Early Jurassic rifting prior to accretion with North America

The circular Bokan Mountain complex (BMC) on southern Prince of Wales Island, southernmost Alaska, is a Jurassic peralkaline granitic intrusion about 3 km in diameter that crosscuts igneous and metasedimentary rocks of the Alexander terrane. The BMC hosts significant rare metal (rare earth elements, Y, U, Th, Zr, and Nb) mineralization related to the last stage of BMC emplacement. U–Pb (zircon) and 40 Ar/ 39 Ar (amphibole and whole-rock) geochronology indicates the following sequence of intrusive activity: ( i ) a Paleozoic basement composed mainly of 469 ± 4 Ma granitic rocks; ( ii ) intrusion of the BMC at 177 ± 1 Ma followed by rapid cooling through ca. 550 °C at 176 ± 1 Ma that was synchronous with mineralization associated with vertical, WNW-trending pegmatites, felsic dikes, and aegirine–fluorite veins and late-stage, sinistral shear deformation; and ( iii ) intrusion of crosscutting lamprophyre dikes at >150 Ma and again at ca. 105 Ma. The peralkaline nature of the BMC and the WNW trend of associated dikes suggest intrusion during NE–SW rifting that was followed by NE–SW shortening during the waning stages of BMC emplacement. The 177 Ma BMC was synchronous with other magmatic centres in the Alexander terrane, such as (1) the Dora Bay peralkaline stock and (2) the bimodal Moffatt volcanic suite located ∼30 km north and ∼100 km SE of the BMC, respectively. This regional magmatism is interpreted to represent a regional extensional event that precedes deposition of the Late Jurassic – Cretaceous Gravina sequence that oversteps the Wrangellia and Alexander exotic accreted terranes and the Taku and Yukon–Tanana pericratonic terranes of the Canadian–Alaskan Cordillera.

Alaska↗

Effects of fine sediment, hyporheic flow, and spawning site characteristics on survival and development of bull trout embryos

Successful spawning is imperative for the persistence of salmonid populations, but relatively little research has been conducted to evaluate factors affecting early life-stage survival for bull trout ( Salvelinus confluentus ), a threatened char. We conducted a field experiment to assess the relationship between site-specific environmental factors and bull trout embryo survival and fry emergence timing. Survival from egg to hatch was negatively related to percent fine sediment (<1 mm) in the redd and positively related to the strength of downwelling at spawning sites. Survival of eggs to fry emergence was also negatively related to fine sediment, and the best statistical models included additional variables that described the rate of downwelling and intragravel flow within the incubation environment. Fry emerged at an earlier stage in development from redds with high percentages of fine sediment. Increased hydraulic conductivity via redd construction and selection of spawning sites with strong downwelling appear to enhance hyporheic flow rates and bull trout egg survival, but early life-stage success may ultimately be limited by intrusion of fine sediment into the incubation environment.

Oregon↗

Geology and petroleum potential of the Arctic Alaska petroleum province

The Arctic Alaska petroleum province encompasses all lands and adjacent continental shelf areas north of the Brooks Range–Herald Arch orogenic belt and south of the northern (outboard) margin of the Beaufort Rift shoulder. Even though only a small part is thoroughly explored, it is one of the most prolific petroleum provinces in North America with total known resources (cumulative production plus proved reserves) of c . 28 BBOE. The province constitutes a significant part of a displaced continental fragment, the Arctic Alaska microplate, that was probably rifted from the Canadian Arctic margin during formation of the Canada Basin. Petroleum prospective rocks in the province, mostly Mississippian and younger, record a sequential geological evolution through passive margin, rift and foreland basin tectonic stages. Significant petroleum source and reservoir rocks were formed during each tectonic stage but it was the foreland basin stage that provided the necessary burial heating to generate petroleum from the source rocks. The lion's share of known petroleum resources in the province occur in combination structural–stratigraphic traps formed as a consequence of rifting and located along the rift shoulder. Since the discovery of the super-giant Prudhoe Bay accumulation in one of these traps in the late 1960s, exploration activity preferentially focused on these types of traps. More recent activity, however, has emphasized the potential for stratigraphic traps and the prospect of a natural gas pipeline in this region has spurred renewed interest in structural traps. For assessment purposes, the province is divided into a Platform assessment unit (AU), comprising the Beaufort Rift shoulder and its relatively undeformed flanks, and a Fold-and-Thrust Belt AU, comprising the deformed area north of the Brooks Range and Herald Arch tectonic belt. Mean estimates of undiscovered, technically recoverable resources include nearly 28 billion barrels of oil (BBO) and 122 trillion cubic feet (TCF) of nonassociated gas in the Platform AU and 2 BBO and 59 TCF of nonassociated gas in the Fold-and-Thrust Belt AU.

Alaska↗

Magnetostratigraphy susceptibility for the Guadalupian Series GSSPs (Middle Permian) in Guadalupe Mountains National Park and adjacent areas in West Texas

Here we establish a magnetostratigraphy susceptibility zonation for the three Middle Permian Global boundary Stratotype Sections and Points (GSSPs) that have recently been defined, located in Guadalupe Mountains National Park, West Texas, USA. These GSSPs, all within the Middle Permian Guadalupian Series, define (1) the base of the Roadian Stage (base of the Guadalupian Series), (2) the base of the Wordian Stage and (3) the base of the Capitanian Stage. Data from two additional stratigraphic successions in the region, equivalent in age to the Kungurian–Roadian and Wordian–Capitanian boundary intervals, are also reported. Based on low-field, mass specific magnetic susceptibility (χ) measurements of 706 closely spaced samples from these stratigraphic sections and time-series analysis of one of these sections, we (1) define the magnetostratigraphy susceptibility zonation for the three Guadalupian Series Global boundary Stratotype Sections and Points; (2) demonstrate that χ datasets provide a proxy for climate cyclicity; (3) give quantitative estimates of the time it took for some of these sediments to accumulate; (4) give the rates at which sediments were accumulated; (5) allow more precise correlation to equivalent sections in the region; (6) identify anomalous stratigraphic horizons; and (7) give estimates for timing and duration of geological events within sections.

Texas↗

Embryonic and larval development and early behavior in grass carp, Ctenopharyngodon idella : implications for recruitment in rivers

With recent findings of grass carp Ctenopharyngodon idella in tributaries of the Great Lakes, information on developmental rate and larval behavior is critical to efforts to assess the potential for establishment within the tributaries of that region. In laboratory experiments, grass carp were spawned and eggs and larvae reared at two temperature treatments, one "cold" and one "warm", and tracked for developmental rate, egg size, and behavior. Developmental rate was quantified using Yi's (1988) developmental stages and the cumulative thermal units method. Grass carp had a thermal minimum of 13.5&deg;C for embryonic stages and 13.3&deg;C for larval stages. Egg size was related to temperature and maternal size, with the largest eggs coming from the largest females, and eggs were generally larger in warmer treatments. Young grass carp larvae exhibited upward and downward swimming interspersed with long periods of lying on the bottom. Swimming capacity increased with ontogeny, and larvae were capable of horizontal swimming and position holding with gas bladder emergence. Developmental rates, behavior, and egg attributes can be used in combination with physical parameters of a river to assess the risk that grass carp are capable of reproduction and recruitment in rivers.

PLoS ONE↗

Differential responses of dinitrogen fixation, diazotrophic cyanobacteria and ammonia oxidation reveal a potential warming-induced imbalance of the N-cycle in biological soil crusts

N 2 fixation and ammonia oxidation (AO) are the two most important processes in the nitrogen (N) cycle of biological soil crusts (BSCs). We studied the short-term response of acetylene reduction assay (ARA) rates, an indicator of potential N 2 fixation, and AO rates to temperature (T, -5°C to 35°C) in BSC of different successional stages along the BSC ecological succession and geographic origin (hot Chihuahuan and cooler Great Basin deserts). ARA in all BSCs increased with T until saturation occurred between 15 and 20°C, and declined at 30–35°C. Culture studies using cyanobacteria isolated from these crusts indicated that the saturating effect was traceable to their inability to grow well diazotrophically within the high temperature range. Below saturation, temperature response was exponential, with Q 10 significantly different in the two areas (~ 5 for Great Basin BSCs; 2–3 for Chihuahuan BSCs), but similar between the two successional stages. However, in contrast to ARA, AO showed a steady increase to 30–35°C in Great Basin, and Chihuhuan BSCs showed no inhibition at any tested temperature. The T response of AO also differed significantly between Great Basin (Q 10 of 4.5–4.8) and Chihuahuan (Q 10 of 2.4–2.6) BSCs, but not between successional stages. Response of ARA rates to T did not differ from that of AO in either desert. Thus, while both processes scaled to T in unison until 20°C, they separated to an increasing degree at higher temperature. As future warming is likely to occur in the regions where BSCs are often the dominant living cover, this predicted decoupling is expected to result in higher proportion of nitrates in soil relative to ammonium. As nitrate is more easily lost as leachate or to be reduced to gaseous forms, this could mean a depletion of soil N over large landscapes globally.

PLoS ONE↗

The annual migration cycle of emperor geese in western Alaska

Most emperor geese ( Chen canagica ) nest in a narrow coastal region of the Yukon-Kuskokwim Delta (YKD) in western Alaska, but their winter distribution extends more than 3000 km from Kodiak Island, Alaska, to the Commander Islands, Russia. We marked 53 adult female emperor geese with satellite transmitters on the YKD in 1999, 2002, and 2003 to examine whether chronology of migration or use of seasonal habitats differed among birds that wintered in different regions. Females that migrated relatively short distances (650–1010 km) between the YKD and winter sites on the south side of the Alaska Peninsula bypassed autumn staging areas on the Bering Sea coast of the Alaska Peninsula or used them for shorter periods (mean = 57 days) than birds that made longer migrations (1600–2640 km) to the western Aleutian Islands (mean = 97 days). Alaska Peninsula migrants spent more days at winter sites (mean = 172 days, 95% CI: 129–214 days) than western Aleutian Island migrants (mean = 91 days, 95% CI: 83–99 days). Birds that migrated 930–1610 km to the eastern Aleutian Islands spent intermediate intervals at fall staging (mean = 77 days) and wintering areas (mean = 108 days, 95% CI: 95–119 days). Return dates to the YKD did not differ among birds that wintered in different regions. Coastal staging areas on the Alaska Peninsula may be especially important in autumn to prepare Aleutian migrants physiologically for long-distance migration to winter sites, and in spring to enable emperor geese that migrate different distances to reach comparable levels of condition before nesting.

Alaska↗

Frequent vocalizing is negatively associated with brood parasitism in a host of the brown-headed cowbird

Brood parasitism by the Brown-headed Cowbird ( Molothrus ater ) can substantially affect host species' reproductive success. The "host-activity" hypothesis suggests that parasites eavesdrop on conspicuous behaviors to locate and parasitize hosts, and several studies of cowbird hosts support this hypothesis. In contrast, a recent study of the Least Bell's Vireo ( Vireo bellii pusillus ) reported a negative association between the host's vocalization rate near the nest and brood parasitism. This contradictory pattern is intriguing because Bell's Vireo is a common cowbird host and vocalizes near and on its nests. We tested a key assumption of the host-activity hypothesis in a different subspecies ( V. b. arizonae ) to determine whether the contradictory pattern reported in V. b. pusillus is an anomaly or could be generalized to other subspecies. Unparasitized vireos vocalized more frequently than parasitized birds, confirming that the pattern in Bell's Vireos is the opposite of that reported for other cowbird hosts. Nesting stage played a role: unparasitized birds vocalized more than parasitized birds only during the nest-building and incubation stages. Given that vocalization rate and other behaviors change through the breeding season, future tests of the host-activity hypothesis should control for nesting stage. Moreover, future efforts to identify the underlying cause for the association between vocalization rate and probability of parasitism should consider the possibility of reciprocal causal relationships between them. We propose five additional hypotheses to explain why in Bell's Vireo the pattern between these two traits is opposite of what has been reported in other birds.

The Condor↗

Intercontinental migratory connectivity and population structuring of Dunlins from western Alaska

The Dunlin (Calidris alpina) is a polytypic shorebird with complex patterns of distribution and migration throughout its holarctic range. We analyzed mark-re sighting data obtained between 1977 and 2010 from birds captured at two major staging areas in western Alaska to test the hypothesis that the migration patterns of Alaskan populations are a mixture of parallel and chain, similar to those of Dunlin populations in the western Palearctic. Birds marked on the Yukon—Kuskokwim Delta were found wintering in both Asia and North America, which documented the unexpected mixing of C. a. arcticola from northern Alaska and C. a. pacifica from western Alaska and contradicted our initial prediction of parallel migration pathways for these two subspecies. In its North American winter range C. a. pacifica segregated according to location of marking, confirming our prediction of a chain migration pattern within this population. Individuals of C. a. pacifica marked on the delta were resighted significantly farther north, mostly in southern British Columbia and Washington, than birds marked on the second, more southerly staging area on the Alaska Peninsula, which were resighted primarily in the San Francisco Bay area of northern California. We recommend additional studies use a combination of intrinsic and extrinsic markers to quantify the strength of migratory connectivity between breeding, staging, and wintering areas. Such information is needed to guide conservation efforts because the Dunlin and other waterbirds are losing intertidal habitats at an unprecedented rate and scale, particularly in the Yellow Sea and other parts of Asia.

Alaska↗

Larval American shad: Effects of age and group size on swimming and feeding behavior

We analyzed the behavior of 3–4‐d‐old prolarval and 28–33‐d‐old metalarval American shad Alosa sapidissima in groups of 3–1,000 fish per 22‐L glass tank, to determine whether (1) previously described juvenile behavior patterns first develop in larvae, (2) group size or density alters the behavior of larvae, and (3) schooling or other forms of cohesive behavior develop in larvae to promote social interactions. Twelve discrete behaviors or modal action patterns (MAPS) oflarvae were observed at all group sizes; halfthese patterns are unique to larval stages. Conversely, larvae do not develop five previously described juvenile MAPS. Stereotyped metalarval feeding sequences were absent or poorly developed in prolarvae. Group size was directly related to duration of free swimming in water column (metalarvae only) and to frequencies of “proximity to another fish” (all larvae), “contact another fish” (all larvae), and “escape or flee” (all larvae). Age or larval stage significantly affected all swimming‐related activities and three feeding behaviors. Larvae foraged and fed independently of one another and used MAPs typical of other larval fishes (“fixate,” “sigmoid,” “lunge,” and “capture”). With one exception (a direct relationship between frequency of food capture and metalarval size), group size and individual size did not significantly affect larval feeding success. Neither schooling nor forms of behavior leading to coordinated group swimming were observed at either larval stage. Larval behavior differed from juvenile behavior in a way that suggests survival in riverine habitats is promoted by behavior that disperses larvae and enables them to function nonsocially.

Transactions of the American Fisheries Society↗

The effects of dissolved gas supersaturation on white sturgeon larvae

Spill at dams has caused supersaturation of atmospheric gas in waters of the Columbia and Snake rivers and raised concerns about the effects of dissolved gas supersaturation (DGS) on white sturgeons Acipenser transmontanus. The timing and location of white sturgeon spawning and the dispersal of white sturgeon larvae from incubation areas makes the larval stage potentially vulnerable to the effects of DGS. To assess the effects of DGS on white sturgeon larvae, we exposed larvae to mean total dissolved gas (TDG) levels of 118% and 131% saturation in laboratory bioassay tests. Gas bubble trauma (GBT) was manifested as a gas bubble in the buccal cavity, nares, or both and it first occurred at developmental stages characterized by the formation of the mouth and gills. Exposure times of 15 min were sufficient to elicit these signs in larvae in various stages of development. No mortality was observed in larvae exposed to 118% TDG for 10 d, but 50% mortality occurred after a 13-d exposure to 131% TDG. The signs of GBT we observed resulted in positive buoyancy and alterations in behavior that may affect the dispersal and predation vulnerability of white sturgeon larvae. The exact depth distribution of dispersing white sturgeon larvae in the Columbia River currently is unknown. Thus, our results may represent a worst-case scenario if white sturgeon larvae are dispersed at depths with insufficient hydrostatic pressure to compensate for high TDG levels.

Transactions of the American Fisheries Society↗

Post-mortem sporulation of Ceratomyxa shasta (Myxozoa) after death in adult Chinook salmon

Ceratomyxa shasta (Myxozoa) is a common gastrointestinal pathogen of salmonid fishes in the Pacific Northwest of the United States. We have been investigating this parasite in adult Chinook salmon ( Oncorhynchus tshawytscha ) in the Willamette River, Oregon. In prior work, we observed differences in the pattern of development of C. shasta in adult salmon compared to juvenile salmon. Adult salmon consistently had large numbers of prespore stages in many of the fish that survived to spawn in the fall. However, myxospores were rarely observed, even though they were exposed and presumably infected for months before spawning. We evaluated the ability of C. shasta to sporulate following fish death because it is reported that myxosores are common in carcasses of Chinook salmon. We collected the intestine from 30 adult salmon immediately after artificial spawning and death (T 0 ). A total of 23 fish were infected with C. shasta based on histology, but only a few myxospores were observed in 1 fish by histology. Intestines of these fish were examined at T 0 and T 7 (latter held at 17 C for 7 days) using quantified wet mount preparations. An increase in myxospore concentrations was seen in 39% of these fish, ranging between a 1.5- to a 14.5-fold increase. The most heavily infected fish exhibited a 4.6-fold increase from 27,841 to 129,352 myxospores/cm. This indicates, supported by various statistical analyses, that under certain conditions presporogonic forms are viable and continue to sporulate after death in adult salmon. Considering the life cycle of C. shasta and anadromous salmon, the parasite may have evolved 2, non-mutually exclusive developmental strategies. In young fish (parr and smolts), the parasite sporulates shortly after infection and is released into freshwater from either live or dead fish before their migration to seawater, where the alternate host is absent. The second strategy occurs in adult salmon, particularly spring Chinook salmon, which become infected upon their return to freshwater in the spring or early summer. For several months throughout the summer, only prespore stages are observed in most fish, even at the time of spawning. But once the fish dies, environmental conditions experienced by C. shasta change and viable presporogonic stages are induced to sporulate. As the post-spawned fish occur in the upper reaches of rivers, the myxospores would be released in a freshwater environment that would provide a reasonable opportunity for them to encounter their freshwater polychaete hosts, which reside downstream.

Journal of Parasitology↗

Sensitivity of two salamander (Ambystoma) species to ultraviolet radiation

Increased ultraviolet-B (UV-B) radiation reaching the Earth's surface has been implicated in amphibian declines. Recent studies have shown that many amphibian species have differences in sensitivity depending on developmental stage. Embryos and larvae of Ambystoma maculatum (Spotted Salamander) and larvae of Ambystoma talpoideum (Mole Salamander) were exposed to five simulated UV-B treatments in controlled laboratory experiments to determine the relative sensitivity of different lifestages. Hatching success of the embryos exceeded 95% in all treatments; however, the larvae of both species exhibited greater sensitivity to UV-B exposure. Older larvae of A. maculatum that were not exposed to UV-B as embryos were more sensitive than larvae that had hatched during exposure to UV-B. Growth of surviving larvae of A. maculatum was significantly reduced as UV-B intensity increased, whereas growth of A. talpoideum was unaffected. These results were compared to ambient UV-B conditions in natural environments. It appears that the embryo stage is relatively unaffected by UV-B levels observed in natural habitats, probably because of protection from vegetation, organic matter in the water column, oviposition depth, and egg jelly. The larval stage of these species may be at greater risk, particularly if there is an increase in UV-B radiation exposure caused by increases in water clarity and/or decreases in dissolved organic carbon.

Journal of Herpetology↗

When can efforts to control nuisance and invasive species backfire?

Population control through harvest has the potential to reduce the abundance of nuisance and invasive species. However, demographic structure and density-dependent processes can confound removal efforts and lead to undesirable consequences, such as overcompensation (an increase in abundance in response to harvest) and instability (population cycling or chaos). Recent empirical studies have demonstrated the potential for increased mortality (such as that caused by harvest) to lead to overcompensation and instability in plant, insect, and fish populations. We developed a general population model with juvenile and adult stages to help determine the conditions under which control harvest efforts can produce unintended outcomes. Analytical and simulation analyses of the model demonstrated that the potential for overcompensation as a result of harvest was significant for species with high fecundity, even when annual stage-specific survivorship values were fairly low. Population instability as a result of harvest occurred less frequently and was only possible with harvest strategies that targeted adults when both fecundity and adult survivorship were high. We considered these results in conjunction with current literature on nuisance and invasive species to propose general guidelines for assessing the risks associated with control harvest based on life history characteristics of target populations. Our results suggest that species with high per capita fecundity (over discrete breeding periods), short juvenile stages, and fairly constant survivorship rates are most likely to respond undesirably to harvest. It is difficult to determine the extent to which overcompensation and instability could occur during real-world removal efforts, and more empirical removal studies should be undertaken to evaluate population-level responses to control harvests. Nevertheless, our results identify key issues that have been seldom acknowledged and are potentially generic across taxa. ?? 2009 by the Ecological Society ot America.

Ecological Applications↗

Ecosystem development in roadside grasslands: Biotic control, plant-soil interactions, and dispersal limitations

Roadside grasslands undergoing secondary succession are abundant, and represent ecologically meaningful examples of novel, human‐created ecosystems. Interactions between plant and soil communities (hereafter plant–soil interactions) are of major importance in understanding the role of biotic control in ecosystem functioning, but little is known about these links in the context of ecosystem restoration and succession. The assessment of the key biotic communities and interactions driving ecosystem development will help practitioners to better allocate the limited resources devoted to roadside grassland restoration. We surveyed roadside grasslands from three successional stages (0–2, 7–9, and >20 years) in two Mediterranean regions of Spain. Structural equation modeling was used to evaluate how interactions between plants, biological soil crusts (BSCs), and soil microbial functional diversity (soil microorganisms) affect indicators of ecosystem development and restoration: plant similarity to the reference ecosystem, erosion control, and soil C storage and N accumulation. Changes in plant community composition along the successional gradient exerted the strongest influence on these indicators. High BSC cover was associated with high soil stability, and high soil microbial functional diversity from late‐successional stages was associated with high soil fertility. Contrary to our expectations, the indirect effects of plants, mediated by either BSCs or soil microorganisms, were very weak in both regions, suggesting a minor role for plant–soil interactions upon ecosystem development indicators over long periods. Our results suggest that natural vegetation dynamics effectively improved ecosystem development within a time frame of 20 years in the grasslands evaluated. They also indicate that this time could be shortened if management actions focus on: (1) maintaining well‐conserved natural areas close to roadsides to enhance plant compositional changes towards late‐successional stages, (2) increasing BSC cover in areas under strong erosion risk, to avoid soil loss, and (3) enhancing soil microbial functional diversity in resource‐limited areas, to enhance soil C and N accumulation.

Ecological Applications↗

Interactions of tissue and fertilizer nitrogen on decomposition dynamics of lignin-rich conifer litter

High tissue nitrogen (N) accelerates decomposition of high-quality leaf litter in the early phases of mass loss, but the influence of initial tissue N variation on the decomposition of lignin-rich litter is less resolved. Because environmental changes such as atmospheric N deposition and elevated CO 2 can alter tissue N levels within species more rapidly than they alter the species composition of ecosystems, it is important to consider how within-species variation in tissue N may shape litter decomposition and associated N dynamics. Douglas-fir (Pseudotsuga menziesii ) is a widespread lignin-rich conifer that dominates forests of high carbon (C) storage across western North America, and displays wide variation in tissue and litter N that reflects landscape variation in soil N. We collected eight unique Douglas-fir litter sources that spanned a two-fold range in initial N concentrations (0.67&ndash;1.31%) with a narrow range of lignin (29&ndash;35%), and examined relationships between initial litter chemistry, decomposition, and N dynamics in both ambient and N fertilized plots at four sites over 3 yr. High initial litter N slowed decomposition rates in both early (0.67 yr) and late (3 yr) stages in unfertilized plots. Applications of N fertilizer to litters accelerated early-stage decomposition, but slowed late-stage decomposition, and most strongly affected low-N litters, which equalized decomposition rates across litters regardless of initial N concentrations. Decomposition of N-fertilized litters correlated positively with initial litter manganese (Mn) concentrations, with litter Mn variation reflecting faster turnover of canopy foliage in high N sites, producing younger litterfall with high N and low Mn. Although both internal and external N inhibited decomposition at 3 yr, most litters exhibited net N immobilization, with strongest immobilization in low-N litter and in N-fertilized plots. Our observation for lignin-rich litter that high initial N can slow decomposition yet accelerate N release differs from findings where litter quality variation across species promotes coupled C and N release during decomposition. We suggest reevaluation of ecosystem models and projected global change effects to account for a potential decoupling of ecosystem C and N feedbacks through litter decomposition in lignin-rich conifer forests.

Ecosphere↗

Anatomy and dynamics of a floodplain, Powder River, Montana, U.S.A.

Centimeter-scale measurements on several Powder River floodplains provide insights into the nature of overbank depositional processes that created the floodplains; during a 20-year period after a major flood in 1978. Rising stages initially entered across a sill at the downriver end of the floodplains. Later, as stages continued to rise, water entered the floodplains through distinct low saddles along natural levees. The annual maximum depth of water over the levee crest averaged 0.19 in from 1983 through 1996, and the estimated flow velocities were approximately 0.15 m s-1. Water ponded in the floodplain trough, a topographic low between the natural levee and the pre-flood riverbank, and mud settled as thin layers of nearly constant thickness. Mud layers alternated with sand layers, which were relatively thick near the channel. Together, these beds created a distinctive natural levee. In some locations, individual flood deposits began as a thin mud layer that gradually coarsened upwards to medium-grained sand. Coarsening-upwards sequences form initially as mud because only the uppermost layers of water in the channel supply the first overbank flows, which are rich in mud but starved of sand. At successively higher stages, fine sands and then medium sands increase in concentration in the floodwater and are deposited as fine- and medium-sand layers overlying the initial mud layer. Theoretical predictions from mathematical models of sediment transport by advection and diffusion indicate that these processes acting alone are unlikely to create the observed sand layers of nearly uniform thickness that extend across much of the floodplain. We infer that other transport processes, notably bedload transport, must be important along Powder River. Even with the centimeter-scale measurements of floodplain deposits, daily hydraulic data, and precise annual surface topographic surveys, we were unable to determine any clear correspondence between the gauged flow record of overbank floods and the depositional layers mapped in the floodplain. These results provide a detailed example of floodplain deposits and depositional processes that should prove useful for interpreting natural levee deposits in a variety of geologic settings. Copyright ?? 2008, SEPM (Society for Sedimentary Geology).

Journal of Sedimentary Research↗

Defoliation effects on Bromus tectorum seed production: Implications for grazing

Cheatgrass (Bromus tectorum L.) is an invasive annual grass that creates near-homogenous stands in areas throughout the Intermountain sagebrush steppe and challenges successful native plant restoration in these areas. A clipping experiment carried out at two cheatgrass-dominated sites in eastern Oregon (Lincoln Bench and Succor Creek) evaluated defoliation as a potential control method for cheatgrass and a seeding preparation method for native plant reseeding projects. Treatments involved clipping plants at two heights (tall = 7.6 cm, and short = 2.5 cm), two phenological stages (boot and purple), and two frequencies (once and twice), although purple-stage treatments were clipped only once. Treatments at each site were replicated in a randomized complete block design that included a control with no defoliation. End-of-season seed density (seeds??m-2) was estimated by sampling viable seeds from plants, litter, and soil of each treatment. Undipped control plants produced an average of approximately 13 000 and 20 000 seeds??m-2 at Lincoln Bench and Succor Creek, respectively. Plants clipped short at the boot stage and again 2 wk later had among the lowest mean seed densities at both sites, and were considered the most successful treatments (Lincoln Bench: F 6,45 = 47.07, P < 0.0001; Succor Creek: F6,40 = 19.60, P < 0.0001). The 95% confidence intervals for seed densities were 123-324 seeds??m-2 from the Lincoln Bench treatment, and 769-2256 seeds??m-2 from the Succor Creek treatment. Literature suggests a maximum acceptable cheatgrass seed density of approximately 330 seeds??m-2 for successful native plant restoration through reseeding. Thus, although this study helped pinpoint optimal defoliation parameters for cheatgrass control, it also called into question the potential for livestock grazing to be an effective seed-bed preparation technique in native plant reseeding projects in cheatgrass-dominated areas.

Rangeland Ecology and Management↗