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At least 145 records · Page 8Linked to original sources

Effects of invasive American bullfrogs and their removal on Northwestern pond turtles

The American bullfrog ( Rana catesbeiana ) is an invasive species globally significant for its role as a generalist predator in freshwater systems. Native turtles are among the species eaten by bullfrogs, and turtle populations are slow to recover from this impact. We examined the effects of bullfrogs and their removal on Northwestern pond turtles ( Actinemys marmorata ) at four sites in Yosemite National Park. From 2016 to 2022, we monitored turtle populations in two sites where bullfrogs were present and two where they have been absent. We removed 12,317 bullfrogs, larvae, and whole egg masses from one site and 4067 from the other, reaching near complete eradication by 2019. We captured just large adult turtles where bullfrogs were present compared with all sizes where bullfrogs were absent. Prior to near complete eradication, juvenile turtles were only found with bullfrogs when they were recovered from bullfrog stomachs. Turtles at bullfrog present sites were 26–36 % larger and 76–97 % heavier than turtles from bullfrog absent sites. Turtle abundance and densities were also 2–100 times higher at bullfrog absent sites. We captured the first juvenile turtles at bullfrog present sites only after reaching near complete bullfrog eradication in 2019. Altogether, our study shows a prolonged lack of juvenile turtle recruitment where bullfrogs were present but offers hope that bullfrog control may succeed in recovering turtle populations by easing predation pressure on hatchlings and juveniles. Our results indicate that bullfrog eradication efforts may be necessary to ensure persistence of at-risk species like native turtles.

California

Predicted macroinvertebrate response to water diversion from a montane stream using two-dimensional hydrodynamic models and zero flow approximation

We used two-dimensional hydrodynamic models for the assessment of water diversion effects on benthic macroinvertebrates and associated habitat in a montane stream in Yosemite National Park, Sierra Nevada Mountains, CA, USA. We sampled the macroinvertebrate assemblage via Surber sampling, recorded detailed measurements of bed topography and flow, and coupled a two-dimensional hydrodynamic model with macroinvertebrate indicators to assess habitat across a range of low flows in 2010 and representative past years. We also made zero flow approximations to assess response of fauna to extreme conditions. The fauna of this montane reach had a higher percentage of Ephemeroptera, Plecoptera, and Trichoptera (%EPT) than might be expected given the relatively low faunal diversity of the study reach. The modeled responses of wetted area and area-weighted macroinvertebrate metrics to decreasing discharge indicated precipitous declines in metrics as flows approached zero. Changes in area-weighted metrics closely approximated patterns observed for wetted area, i.e., area-weighted invertebrate metrics contributed relatively little additional information above that yielded by wetted area alone. Loss of habitat area in this montane stream appears to be a greater threat than reductions in velocity and depth or changes in substrate, and the modeled patterns observed across years support this conclusion. Our models suggest that step function losses of wetted area may begin when discharge in the Merced falls to 0.02 m 3 /s; proportionally reducing diversions when this threshold is reached will likely reduce impacts in low flow years.

California

Impacts of temporal revisit designs on the power to detect trend with a linear mixed model: An application to long-term monitoring of Sierra Nevada lakes

Long-term ecological monitoring programs often use linear mixed models to estimate trend in an ecological indicator sampled across large landscapes. A linear mixed model is versatile for estimating a linear trend in time as well as components of spatial and temporal variationin the case of unbalanced data structures , which are common in complex monitoring designs where limited sampling effort must be optimized over time and space. A power analysis was used to inform a lake chemistry monitoring design, including selecting the most appropriate temporal revisit design. Pilot data from surveys of lakes across large wilderness national parks (Sequoia, Kings Canyon, and Yosemite national parks) were used to obtain variance components for a Monte Carlo power simulation. Using a linear mixed model for a range of temporal revisit designs, sample sizes, and trend magnitudes, we evaluated the power to detect trend, the trend test size, and the relative bias of trend coefficient estimates for four continuous and normally distributed indicators. Contrary to prior research based on large-sample approximations that identified a single panel of sites visited annually as the revisit design generating the highest power, we found that the power to detect a 12-year trend based on the Wald t-test from a linear mixed model may be optimized by obtaining unbalanced data sets with limited to no annual replication. We emphasize the importance of examining variance composition, sample size, and the power and size of the trend test with Monte Carlo simulation when allocating sampling effort over time and space.

Ecological Indicators

Snag dynamics and surface fuel loads in the Sierra Nevada: Predicting the impact of the 2012–2016 drought

Forest die-backs linked to extreme droughts are expected to increase as the climate dries and warms. An example is the 2012-2016 hotter drought in California that induced widespread tree mortality in the Sierra Nevada, California. The sudden increase in snags (i.e., standing dead trees) raised immediate concerns about their impact on wildfire hazard and longer-term questions about their impact on ecosystem structure and function. We quantified the likely progression of snag fall and fuel succession following the recent extensive mortality event in the southern Sierra Nevada mixed conifer forest. Our results used data from a long-term demography study to project trends in surface fuel loads at three study sites in Yosemite and Sequoia Kings Canyon National Parks. In the short term (2017-2021), fine woody debris and litter + duff significantly increased across all three sites (>145% and >55%, respectively); coarse woody debris increased significantly at one site (48.6%); and total fuel loads increased significantly at two of the three sites (38% and 69%). Snag longevity increased with size, with the relationship varying by species. Yellow pine was a notable outlier: size played a small role in influencing its fall rates. Overall, species-specific snag fall rates in the southern Sierra Nevada were 20% to 40% slower than previously reported. By 2040, projected median cumulative inputs of biomass from future snag fall range from 49.4 Mg ha-1 to 136.1 Mg ha-1across our three sites, which exceeds the amounts currently present (47.17-89.97 Mg ha-1) and is well above estimates of historical coarse woody debris amounts in the Sierra Nevada (17.7 Mg ha -1). These results provide a robust empirical basis to refine the snag fall algorithm in vegetation simulation models. Options to manage the impact of extreme number of snags and their large surface combustible biomass include salvage operations and prescribed burning, with both methods having operational, financial, and legal limitations that need to be considered.

Forest Ecology and Management

Forest cover influences fish mercury concentrations in national parks of the western U.S.

The global prevalence of mercury (Hg) contamination and its complex biogeochemical cycling has resulted in elevated Hg concentrations in biota in remote and pristine environments. However, there is uncertainty in the relative importance of Hg deposition and landscape factors that control Hg cycling and bioaccumulation. To address this, we measured total mercury (THg) concentrations in 1344 fish across 60 subalpine lakes from 12 national parks (NPs). These parks represent three distinct high-elevation regions across the western U.S.: Cascades and Olympic Peninsula, Sierra Nevada and Great Basin, and Rocky Mountains. Within these regions, three NPs (Mount Rainier, Yosemite, and Rocky Mountain) were intensively studied representatives of each region. This study aimed to (1) assess the magnitude of mercury contamination in a collection of remote, small catchment lakes; (2) quantify the variability of fish THg concentrations among and within parks; and (3) test the relative importance of Hg inputs in comparison to landscape characteristics on lake-specific fish THg concentrations. The spatial variability in fish THg concentrations was 2.6-fold higher than variation in deposition to watersheds, suggesting that factors other than Hg delivery are important determinants of Hg accumulation in these environments. Spatially, fish THg concentrations (ng/g ww ± standard error) were lower in the Rockies (46.2 ± 5.0) and Sierra (56.5 ± 5.8) compared to the Cascades (67.8 ± 6.1). Additionally, fish THg concentrations increased with increasing conifer forest cover (Intensive parks: P < 0.0001, R 2 = 0.43; All parks: P = 0.0001, R 2 = 0.23) but were not correlated with wet Hg deposition across the catchment. These findings suggest that forest composition is likely an important aspect of Hg delivery to lake food webs, and although the mechanisms are unclear, could be tied to some combination of forest influences on catchment organic carbon and increased surface area for dry Hg deposition.

western United States

Chemical weathering of a soil chronosequence on granitoid alluvium: II. Mineralogic and isotopic constraints on the behavior of strontium

The use of strontium isotopes to evaluate mineral weathering and identify sources of base cations in catchment waters requires an understanding of the behavior of Sr in the soil environment as a function of time. Our approach is to model the temporal evolution of 87 Sr/ 86 Sr of the cation exchange pool in a soil chronosequence developed on alluvium derived from central Sierra Nevada granitoids during the past 3 Ma. With increasing soil age, 87 Sr/ 86 Sr of ammonium-acetate extractable Sr initially decreases from values typical of K-feldspar to those of plagioclase and hornblende and then remains constant, even though plagioclase and hornblende are absent from the soils after approximately 1 Ma of weathering. The temporal variation of 87 Sr/ 86 Sr of exchangeable Sr is modeled by progressively equilibrating Sr derived from mineral weathering and atmospheric deposition with Sr on exchange sites as waters infiltrate a soil column. Observed decreases in quartz-normalized modal abundances of plagioclase, hornblende, and K-feldspar with time, and the distinct 87 Sr/ 86 Sr values of these minerals can be used to calculate Sr flux from weathering reactions. Hydrobiotites in the soils have nearly constant modal abundances, chemistry, and 87 Sr/ 86 Sr over the chronosequence and provide negligible Sr input to weathering solutions. The model requires time and soil horizon-dependent changes in the amount of exchangeable Sr and the efficiency of Sr exchange, as well as a biologic cycling term. The model predicts that exchangeable Sr initially has 87 Sr/ 86 Sr identical to that of K-feldspar, and thus could be dominated by Sr leached from K-feldspar following deposition of the alluvium. The maximum value of 87 Sr/ 86 Sr observed in dilute stream waters associated with granitoids of the Yosemite region is likewise similar to that of the K-feldspars, suggesting that K-feldspar and not biotite may be the dominant source of radiogenic Sr in the streams. This study reveals that, when attempting to use Strontium isotopes to identify sources of base cations in catchment waters and biomass, both preferential leaching of Sr from minerals during incipient soil development and changing Sr exchange efficiency must be considered along with chemical contributions due to mineral dissolution.

Geochimica et Cosmochimica Acta

Pesticides in amphibian habitats of Central and Northern California, USA

Previous studies have indicated that toxicity from pesticide exposure may be contributing to amphibian declines in California and that atmospheric deposition could be a primary pathway for pesticides to enter amphibian habitats. We report on a survey of California wetlands sampled along transects associated with Lassen Volcanic National Park, Lake Tahoe, Yosemite National Park, and Sequoia National Park. Each transect ran from the Pacific coast to the Cascades or Sierra Nevada mountains. Pacific chorus frogs (Pseudacris regilla), water, and sediment were collected from wetlands in 2001 and 2002. Twenty-three pesticides were found in frog, water, or sediment samples. Six contaminants including trifluralin, &alpha;-endosulfan, chlordanes, and trans-nonachlor were found in adult P. regilla. Seventeen contaminants were found in sediments, including endosulfan sulfate, chlordanes, 1-chloro-4-[2,2-dichloro-1-(4-chlorophenyl)ethenyl]benzene (4,4&prime;-DDE), and chlorpyrifos. The mean number of chemicals detected per pond in sediments was 2.4 (2.5, standard deviation). In water, 17 chemicals were detected, with &beta;-endosulfan being present in almost all samples. Trifluralin, chlordanes, and chlorpyrifos were the next most common. The mean number of chemicals in water per pond was 7.8 (2.9). With the possible exception of chlorpyrifos oxon in sediments and total endosulfans, none of the contaminants exceeded known lethal or sublethal concentrations in P. regilla tissue. Endosulfans, chlorpyrifos, and trifluralin were associated with historic and present day population status of amphibians. Cholinesterase, an essential neurological enzyme that can be depressed by certain pesticides, was reduced in tadpoles from areas with the greatest population declines.

Book chapter

Chemistry of selected high-elevation lakes in seven national parks in the western United States

A chemical survey of 69 high-altitude lakes in seven national parks in the western United States was conducted during the fallof 1999; the lakes were previously sampled during the fall of 1985, as part of the Western Lake Survey. Lakes in parks in the Sierra/southern Cascades (Lassen Volcanic, Yosemite, Sequoia/Kings Canyon National Parks) and in the southern RockyMountains (Rocky Mountain National Park) were very dilute; medianspecific conductance ranged from 4.4 to 12.2 μS cm -1 andmedian alkalinity concentrations ranged from 32.2 to 72.9 μeqL -1 . Specific conductances and alkalinity concentrations were substantially higher in lakes in the central and northernRocky Mountains parks (Grand Teton, Yellowstone, and GlacierNational Parks), probably due to the prevalence of more reactivebedrock types. Regional patterns in lake concentrations of NO 3 and SO 4 were similar to regional patterns in NO 3 and SO 4 concentrations in precipitation, suggestingthat the lakes are showing a response to atmospheric deposition.Concentrations of NO 3 were particularly high in Rocky Mountain National Park, where some ecosystems appear to be undergoing nitrogen saturation.

Water, Air, and Soil Pollution: Focus

Rockfall triggering by cyclic thermal stressing of exfoliation fractures

Exfoliation of rock deteriorates cliffs through the formation and subsequent opening of fractures, which in turn can lead to potentially hazardous rockfalls. Although a number of mechanisms are known to trigger rockfalls, many rockfalls occur during periods when likely triggers such as precipitation, seismic activity and freezing conditions are absent. It has been suggested that these enigmatic rockfalls may occur due to solar heating of rock surfaces, which can cause outward expansion. Here we use data from 3.5 years of field monitoring of an exfoliating granite cliff in Yosemite National Park in California, USA, to assess the magnitude and temporal pattern of thermally induced rock deformation. From a thermodynamic analysis, we find that daily, seasonal and annual temperature variations are sufficient to drive cyclic and cumulative opening of fractures. Application of fracture theory suggests that these changes can lead to further fracture propagation and the consequent detachment of rock. Our data indicate that the warmest times of the day and year are particularly conducive to triggering rockfalls, and that cyclic thermal forcing may enhance the efficacy of other, more typical rockfall triggers.

California

Detection of rock bridges by infrared thermal imaging and modeling

Characterization of rock discontinuities and rock bridges is required to define stability conditions of fractured rock masses in both natural and engineered environments. Although remote sensing methods for mapping discontinuities have improved in recent years, remote detection of intact rock bridges on cliff faces remains challenging, with their existence typically confirmed only after failure. In steep exfoliating cliffs, such as El Capitan in Yosemite Valley (California, USA), rockfalls mainly occur along cliff-parallel exfoliation joints, with rock bridges playing a key role in the stability of partially detached exfoliation sheets. We employed infrared thermal imaging (i.e., thermography) as a new means of detecting intact rock bridges prior to failure. An infrared thermal panorama of El Capitan revealed cold thermal signatures for the surfaces of two granitic exfoliation sheets, consistent with the expectation that air circulation cools the back of the partially detached sheets. However, we also noted small areas of warm thermal anomalies on these same sheets, even during periods of nocturnal rock cooling. Rock attachment via rock bridges is the likely cause for the warm anomalies in the thermal data. 2-D model simulations of the thermal behavior of one of the monitored sheets reproduce the observed anomalies and explain the temperature differences detected in the rock bridge area. Based on combined thermal and ground-based lidar imaging, and using geometric and rock fracture mechanics analysis, we are able to quantify the stability of both sheets. Our analysis demonstrates that thermography can remotely detect intact rock bridges and thereby greatly improve rockfall hazard assessment.

California

Contrasting prescription burning and wildfires in California Sierra Nevada national parks and adjacent national forests

History of prescription burning and wildfires in the three Sierra Nevada National Park Service (NPS) parks and adjacent US Forest Service (USFS) forests is presented. Annual prescription (Rx) burns began in 1968 in Sequoia and Kings Canyon National Parks, followed by Yosemite National Park and Lassen Volcanic National Park. During the last third of the 20th century, USFS national forests adjacent to these parks did limited Rx burns, accounting for very little area burned. However, in 2004, an aggressive annual burn program was initiated in these national forests and in the last decade, area burned by planned prescription burns, relative to area protected, was approximately comparable between these NPS and USFS lands. In 1968, the NPS prescription burning program was unique because it coupled planned Rx burns with managing many lightning-ignited fires for resource benefit. From 1968 to 2017, these natural fires managed for resource benefit averaged the same total area burned as planned Rx burns in the three national parks; thus, they have had a substantial impact on total area burned by prescription. In contrast, on USFS lands, most lightning-ignited fires have been managed for suppression, but increasing attention is being paid to managing wildfires for resource benefit.

California

Pesticides are involved with population declines of amphibians in the California Sierra Nevadas

Several species of frogs and toads are in serious decline in the Sierra Nevada Mountains of California. These species include the threatened red-legged frog ( Rana aurora ), foothill yellow-legged frog ( R. boylii ), mountain yellow-legged frog ( R. muscosa ), Cascades frog ( Rana cascadae ), western toad ( Bufo boreas ) and Yosemite toad ( B. canorus ). For many of these species current distributions are down to 10% of historical ranges. Several factors including introduced predators, habitat loss, and ultraviolet radiation have been suggested as causes of these declines. Another probable cause is air-borne pesticides from the Central Valley of California. The Central Valley, especially the San Joaquin Valley, is a major agricultural region where millions of pounds of active ingredient pesticides are applied each year (http://www.cdpr.ca.gov/dprdatabase.htm). Prevailing westerly winds from the Pacific Coast transport these pesticides into the into the Sierras.

Scientific World Journal

Climate change refugia and habitat connectivity promote species persistence

Background Climate change refugia, areas buffered from climate change relative to their surroundings, are of increasing interest as natural resource managers seek to prioritize climate adaptation actions. However, evidence that refugia buffer the effects of anthropogenic climate change is largely missing. Methods Focusing on the climate-sensitive Belding’s ground squirrel ( Urocitellus beldingi ), we predicted that highly connected Sierra Nevada meadows that had warmed less or shown less precipitation change over the last century would have greater population persistence, as measured by short-term occupancy, fewer extirpations over the twentieth century, and long-term persistence measured through genetic diversity. Results Across California, U. beldingi were more likely to persist over the last century in meadows with high connectivity that were defined as refugial based on a suite of temperature and precipitation factors. In Yosemite National Park, highly connected refugial meadows were more likely to be occupied by U. beldingi . More broadly, populations inhabiting Sierra Nevada meadows with colder mean winter temperatures had higher values of allelic richness at microsatellite loci, consistent with higher population persistence in temperature-buffered sites. Furthermore, both allelic richness and gene flow were higher in meadows that had higher landscape connectivity, indicating the importance of metapopulation processes. Conversely, anthropogenic refugia, sites where populations appeared to persist due to food or water supplementation, had lower connectivity, genetic diversity, and gene flow, and thus might act as ecological traps. This study provides evidence that validates the climate change refugia concept in a contemporary context and illustrates how to integrate field observations and genetic analyses to test the effectiveness of climate change refugia and connectivity. Conclusions Climate change refugia will be important for conserving populations as well as genetic diversity and evolutionary potential. Our study shows that in-depth modeling paired with rigorous fieldwork can identify functioning climate change refugia for conservation.

California

Visitation to national parks in California shows annual and seasonal change during extreme drought and wet years

This study examines the influence of drought indicators on recreational visitation patterns to National Park Service units in California (USA) from 1980 to 2019. We considered mountain, arid, and coastal park types across a climate gradient where seasonal recreational opportunities are directly or indirectly dependent on water resources. Significant departures from the normal hydroclimate, reflected by drought or unusually wet conditions, can lead visitors to change their behavior, including recreating at a different time or place. Drought conditions can facilitate earlier seasonal access at higher elevation parks, but displace visitors in other seasons and parks. Wetter-than-average conditions can displace visitors due to snowpack or flooding, but also facilitate other activities. We found a decrease in annual visitation at popular mountain parks including Yosemite (-8.6%) and Sequoia and Kings Canyon (-8.2%) during extreme drought years due to lower-than-average attendance in peak summer and fall months. Extreme wet years also had significantly reduced annual visitation in Sequoia and Kings (-8.5%) and Lassen Volcanic (-13.9%) due to declines in spring and summer use as snowpack restricts road access. For arid parks, drought status did not have a statistically significant effect on annual visitation, although extreme drought led to less use during the hottest months of summer at Death Valley, and extreme wet conditions at Pinnacles led to less visitation throughout the year (-16.6%), possibly from impacts to infrastructure associated with flooding. For coastal park units, extreme drought led to year-round higher levels of use at Redwood (+27.7%), which is typically wet, and less year-round use at Channel Islands (-23.6%), which is relatively dry, while extreme wet years led to higher levels of annual use at Channel Islands (+29.4%). Collectively, these results indicate the effect of extreme drought or wet years on park visitation varies by park depending on geography and recreational activities offered.

California

A multi-scale evaluation of pack stock effects on subalpine meadow plant communities in the Sierra Nevada

We evaluated the influence of pack stock (i.e., horse and mule) use on meadow plant communities in Sequoia and Yosemite National Parks in the Sierra Nevada of California. Meadows were sampled to account for inherent variability across multiple scales by: 1) controlling for among-meadow variability by using remotely sensed hydro-climatic and geospatial data to pair stock use meadows with similar non-stock (reference) sites, 2) accounting for within-meadow variation in the local hydrology using in-situ soil moisture readings, and 3) incorporating variation in stock use intensity by sampling across the entire available gradient of pack stock use. Increased cover of bare ground was detected only within “dry” meadow areas at the two most heavily used pack stock meadows (maximum animals per night per hectare). There was no difference in plant community composition for any level of soil moisture or pack stock use. Increased local-scale spatial variability in plant community composition (species dispersion) was detected in “wet” meadow areas at the two most heavily used meadows. These results suggest that at the meadow scale, plant communities are generally resistant to the contemporary levels of recreational pack stock use. However, finer-scale within-meadow responses such as increased bare ground or spatial variability in the plant community can be a function of local-scale hydrological conditions. Wilderness managers can improve monitoring of disturbance in Sierra Nevada meadows by adopting multiple plant community indices while simultaneously considering local moisture regimes.

California

California Spotted Owl (Strix occidentalis occidentalis) habitat use patterns in a burned landscape

Fire is a dynamic ecosystem process of mixed-conifer forests of the Sierra Nevada, but there is limited scientific information addressing wildlife habitat use in burned landscapes. Recent studies have presented contradictory information regarding the effects of stand-replacing wildfires on Spotted Owls ( Strix occidentalis ) and their habitat. While fire promotes heterogeneous forest landscapes shown to be favored by owls, high severity fire may create large canopy gaps that can fragment the closed-canopy habitat preferred by Spotted Owls. We used radio-telemetry to determine whether foraging California Spotted Owls ( S. o. occidentalis ) in Yosemite National Park, California, USA, showed selection for particular fire severity patch types within their home ranges. Our results suggested that Spotted Owls exhibited strong habitat selection within their home ranges for locations near the roost and edge habitats, and weak selection for lower fire severity patch types. Although owls selected high contrast edges with greater relative probabilities than low contrast edges, we did not detect a statistical difference between these probabilities. Protecting forests from stand-replacing fires via mechanical thinning or prescribed fire is a priority for management agencies, and our results suggest that fires of low to moderate severity can create habitat conditions within California Spotted Owls' home ranges that are favored for foraging.

The Condor

Population size, survival, growth, and movements of Rana sierrae

Based on 2431 captures of 757 individual frogs over a 9-yr period, we found that the population of R. sierrae in one meadow–stream complex in Yosemite National Park ranged from an estimated 45 to 115 adult frogs. Rana sierrae at our relatively low elevation site (2200 m) grew at a fast rate (K = 0.73–0.78), had high overwintering survival rates (44.6–95%), lived a long time (up to 16 yr), and tended to be fairly sedentary during the summer (100% minimum convex polygon annual home ranges of 139 m2) but had low year-to-year site fidelity. Even though the amphibian chytrid fungus (Batrachochytrium dendrobatidis, Bd) has been present in the population for at least 13 yr, there was no clear downward trend as might be expected from reports of R. sierrae population declines associated with Bd or from reports of widespread population decline of R. sierrae throughout its range.

Herpetologica

Recent rates of carbon accumulation in montane fens ofYosemite National Park, California, U.S.A.

Little is known about recent rates of carbon storage in montane peatlands, particularly in the western United States. Here we report on recent rates of carbon accumulation (past 50 to 100 years) in montane groundwater-fed peatlands (fens) of Yosemite National Park in central California, U.S.A. Peat cores were collected at three sites ranging in elevation from 2070 to 2500 m. Core sections were analyzed for bulk density, % organic carbon, and 210 Pb activities for dating purposes. Organic carbon densities ranged from 0.026 to 0.065 g C cm -3 . Mean vertical accretion rates estimated using 210 Pb over the 50-year period from &sim;1960 to 2011 and the 100-year period from &sim;1910 to 2011 were 0.28 (standard deviation = &plusmn;0.09) and 0.18 (&plusmn;-0.04) cm yr -1 , respectively. Mean carbon accumulation rates over the 50- and 100-year periods were 95.4 (&plusmn;25.4) and 74.7 (&plusmn;17.2) g C m -2 yr -1 , respectively. Such rates are similar to recent rates of carbon accumulation in rich fens in western Canada, but more studies are needed to definitively establish both the similarities and differences in peat formation between boreal and temperate montane fens.

California