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At least 1,369 records · Page 76Linked to original sources

A new parameterization for estimating co‐occurrence of interacting species

Models currently used to estimate patterns of species co‐occurrence while accounting for errors in detection of species can be difficult to fit when the effects of covariates on species occurrence probabilities are included. The source of the estimation problems is the particular parameterization used to specify species co‐occurrence probability. We develop a new parameterization for estimating patterns of co‐occurrence of interacting species that allows the effects of covariates to be specified quite naturally without estimation problems. In our model, the occurrence of one species is assumed to depend on the occurrence of another, but the occurrence of the second species is not assumed to depend on the presence of the first species. This pattern of co‐occurrence, wherein one species is dominant and the other is subordinate, can be produced by several types of ecological interactions (predator–prey, parasitism, and so on). A simulation study demonstrated that estimates of species occurrence probabilities were unbiased in samples of 50–100 locations and three surveys per location, provided species are easily detected (probability of detection ≥ 0.5). Higher sample sizes (>200 locations) are needed to achieve unbiasedness when species are more difficult to detect. An analysis of data from treefrog surveys in southern Florida indicated that the occurrence of Cuban treefrogs, an invasive predator species, was highest near the point of its introduction and declined with distance from that location. Sites occupied by Cuban treefrogs were 9.0 times less likely to contain green treefrogs and 15.7 times less likely to contain squirrel treefrogs compared to sites without Cuban treefrogs. The detection probabilities of native treefrog species did not depend on the presence of Cuban treefrogs, suggesting that the native treefrog species are naive to the introduced species.

Ecological Applications↗

Food webs including parasites, biomass, body sizes, and life stages for three California/Baja California estuaries

This data set presents food webs for three North American Pacific coast estuaries and a “Metaweb” composed of the species/stages compiled from all three estuaries. The webs have four noteworthy attributes: (1) parasites (infectious agents), (2) body-size information, (3) biomass information, and (4) ontogenetic stages of many animals with complex life cycles. The estuaries are Carpinteria Salt Marsh, California (CSM); Estero de Punta Banda, Baja California (EPB); and Bahía Falsa in Bahía San Quintín, Baja California (BSQ). Most data on species assemblages and parasitism were gathered via consistent sampling that acquired body size and biomass information for plants and animals larger than ∼1 mm, and for many infectious agents (mostly metazoan parasites, but also some microbes). We augmented this with information from additional published sources and by sampling unrepresented groups (e.g., plankton). We estimated free-living consumer–resource links primarily by extending a previously published version of the CSM web (which the current CSM web supplants) and determined most parasite consumer–resource links from direct observation. We recognize 21 possible link types including four general interactions: predators consuming prey, parasites consuming hosts, predators consuming parasites, and parasites consuming parasites. While generally resolved to the species level, we report stage-specific nodes for many animals with complex life cycles. We include additional biological information for each node, such as taxonomy, lifestyle (free-living, infectious, commensal, mutualist), mobility, and residency. The Metaweb includes 500 nodes, 314 species, and 11 270 links projected to be present given appropriate species' co-occurrences. Of these, 9247 links were present in one or more of the estuarine webs. The remaining 2023 links were not present in the estuaries but are included here because they may occur in other places or times. Initial analyses have examined and are examining the interrelationships among consumer strategy, body size, abundance, biomass, trophic level, life stages, and food-web structure and dynamics. Further use of these data may enable a more general exploration how infectious processes and parasites impact communities and ecosystems. Additionally, we present the data and metadata in a standardized format, attempting to provide a system-neutral template for future food-web assembly and publication.

Ecology↗

Conditions for the return and simulation of the recovery of burrowing mayflies in western Lake Erie

In the 1950s, burrowing mayflies, Hexagenia spp. ( H. limbata and H. rigida ), were virtually eliminated from the western basin of Lake Erie (a 3300 km 2 area) because of eutrophication and pollution. We develop and present a deterministic model for the recolonization of the western basin by Hexagenia to pre-1953 densities. The model was based on the logistic equation describing the population growth of Hexagenia and a presumed competitor, Chironomus (dipteran larvae). Other parameters (immigration, low oxygen, toxic sediments, competition with Chironomus, and fish predation) were then individually added to the logistic model to determine their effect at different growth rates. The logistic model alone predicts 10–41 yr for Hexagenia to recolonize western Lake Erie. Immigration reduced the recolonization time by 2–17 yr. One low-oxygen event during the first 20 yr increased recovery time by 5–17 yr. Contaminated sediments added 5–11 yr to the recolonization time. Competition with Chironomus added 8–19 yr to recovery. Fish predators added 4–47 yr to the time required for recolonization. The full model predicted 48–81 yr for Hexagenia to reach a carrying capacity of ≈350 nymphs/m 2 , or not until around the year 2038 if the model is started in 1990. The model was verified by changing model parameters to those present in 1970, beginning the model in 1970 and running it through 1990. Predicted densities overlapped almost completely with actual estimated densities of Hexagenia nymphs present in the western basin in Lake Erie in 1990. The model suggests that recovery of large aquatic ecosystems may lag substantially behind remediation efforts.

Michigan, Ohio, Ontario↗

Multi-trophic resilience of boreal lake ecosystems to forest fires

Fires are the major natural disturbance in the boreal forest, and their frequency and intensity will likely increase as the climate warms. Terrestrial nutrients released by fires may be transported to boreal lakes, stimulating increased primary productivity, which may radiate through multiple trophic levels. Using a before‐after‐control‐impact (BACI) design, with pre‐ and postfire data from burned and unburned areas, we examined effects of a natural fire across several trophic levels of boreal lakes, from nutrient and chlorophyll levels, to macroinvertebrates, to waterbirds. Concentrations of total nitrogen and phosphorus were not affected by the fire. Chlorophyll a levels were also unaffected, likely reflecting the stable nutrient concentrations. For aquatic invertebrates, we found that densities of three functional feeding groups did not respond to the fire (filterers, gatherers, scrapers), while two groups increased (shredders, predators). Amphipods accounted for 98% of shredder numbers, and we hypothesize that fire‐mediated habitat changes may have favored their generalist feeding and habitat ecology. This increase in amphipods may, in turn, have driven increased predator densities, as amphipods were the most numerous invertebrate in our lakes and are commonly taken as prey. Finally, abundance of waterbird young, which feed primarily on aquatic invertebrates, was not affected by the fire. Overall, ecosystems of our study lakes were largely resilient to forest fires, likely due to their high initial nutrient concentrations and small catchment sizes. Moreover, this resilience spanned multiple trophic levels, a significant result for ecologically similar boreal regions, especially given the high potential for increased fires with future climate change.

Alaska↗

Amphibians in the climate vise: loss and restoration of resilience of montane wetland ecosystems in the western US

Wetlands in the remote mountains of the western US have undergone two massive ecological “experiments” spanning the 20th century. Beginning in the late 1800s and expanding after World War II, fish and wildlife managers intentionally introduced millions of predatory trout (primarily Oncorhynchus spp) into fishless mountain ponds and lakes across the western states. These new top predators, which now occupy 95% of large mountain lakes, have limited the habitat distributions of native frogs, salamanders, and wetland invertebrates to smaller, more ephemeral ponds where trout do not survive. Now a second “experiment” – anthropogenic climate change – threatens to eliminate many of these ephemeral habitats and shorten wetland hydroperiods. Caught between climate-induced habitat loss and predation from introduced fish, native mountain lake fauna of the western US – especially amphibians – are at risk of extirpation. Targeted fish removals, guided by models of how wetlands will change under future climate scenarios, provide innovative strategies for restoring resilience of wetland ecosystems to climate change.

Western United States↗

Spatial complexity reduces interaction strengths in the meta-food web of a river floodplain mosaic

Theory states that both the spatial complexity of landscapes and the strength of interactions between consumers and their resources are important for maintaining biodiversity and the 'balance of nature.' Spatial complexity is hypothesized to promote biodiversity by reducing potential for competitive exclusion; whereas, models show weak trophic interactions can enhance stability and maintain biodiversity by dampening destabilizing oscillations associated with strong interactions. Here we show that spatial complexity can reduce the strength of consumer-resource interactions in natural food webs. By sequentially aggregating food webs of individual aquatic habitat patches across a floodplain mosaic, we found that increasing spatial complexity resulted in decreases in the strength of interactions between predators and prey, owing to a greater proportion of weak interactions and a reduced proportion of strong interactions in the meta-food web. The main mechanism behind this pattern was that some patches provided predation refugia for species which were often strongly preyed upon in other patches. If weak trophic interactions do indeed promote stability, then our findings may signal an additional mechanism by which complexity and stability are linked in nature. In turn, this may have implications for how the values of landscape complexity, and the costs of biophysical homogenization, are assessed.

Ecology↗

Comparative growth and consumption potential of rainbow trout and humpback chub in the Colorado River, Grand Canyon, Arizona, under different temperature scenarios

We used bioenergetics models for humpback chub, Gila cypha, and rainbow trout, Oncorhynchus mykiss, to examine how warmer water temperatures in the Colorado River, Grand Canyon, Arizona, through a proposed selective withdrawal system (SWS) at Glen Canyon Dam, would affect growth, consumption, and predation rates. Consumption by the rainbow trout population was at least 10 times higher than by the smaller humpback chub population. Water temperature increases of 6??C during autumn increased growth of humpback chub and likely increased their survival by reducing the time vulnerable to predation. Water temperature increases caused by drought in 2005 did not alter humpback chub growth as much as the SWS. Increased temperatures might cause changes to the invertebrate community and the distribution and abundance of other warmwater nonnative fishes. The implications on the entire aquatic community need to be considered before any management action that includes increasing water temperatures is implemented.

Southwestern Naturalist↗

Microhabitat use of native Santa Ana sucker and arroyo chub in an effluent-dominated southern California stream

A significant amount of the base flow of the Santa Ana River, located within California's arid Los Angeles metropolitan region, originates from two wastewater treatment facilities: the Rialto wastewater treatment facility and Rapid Infiltration and Extraction facility. The Santa Ana sucker ( Pantosteus santaanae , syn. Catostomus santaanae ) and arroyo chub ( Gila orcuttii ) are two native species listed in the Upper Santa Ana Habitat Conservation Plan, which aims to balance water supply needs with the ecological needs of native fauna. Consequently, an understanding of the habitat needs of these fishes will play a crucial role in achieving the goals outlined by the conservation plan. We used fish presence during snorkel surveys with habitat availability surveys to quantify habitat selection by both native fish species within a resource selection function analytical framework. We found that both species selected habitat near structures that could serve as refugia from potential predators. In addition, Santa Ana sucker selected habitats with high gravel and cobble substrate composition, presumably to fulfill complementary habitat needs such as foraging and spawning. Our results suggest that habitat alterations due to water management infrastructure and an increasing population size of nonnative predators may also affect native fish habitat selection in the Santa Ana River.

California↗

Trophic relationships of small nonnative fishes in a natural creek and several agricultural drains flowing into the Salton Sea, and their potential, effects on the endangered desert pupfish

This study was conducted to characterize trophic relationships of small nonnative fishes and to determine if predation by these fishes contributes to the decline of desert pupfish (Cyprinodon macularius), an endangered cyprinodont on the verge of extinction. We sampled 403 hybrid Mozambique tilapias (Oreochromis mossambica by O. urolepis), 107 redbelly tilapias (Tilapia zillii), 32 longjaw mudsuckers (Gillkhthys mirabilis), 182 western mosquitofish (Gambusia affinis), 222 sailfin mollies (Poecilia latipinna), 63 shortfin mollies (Poecilia mexicana), and 235 porthole livebearers (Poecilurpsis gracilis) from a natural creek and four agricultural drains during September 1999- December 2001. Evidence of piscivory was in gastrointestinal contents of 14 hybrid Mozambique tilapias, 3 redbelly tilapias, 10 longjaw mudsuckers, 8 western mosquitofish, 2 sailfin mollies, and 8 porthole livebearers. Although digestion often was too advanced for identification of fishes consumed by nonnative fishes, remains of desert pupfish were in gastrointestinal contents of a longjaw mudsucker. Our findings, along with Field evidence from other studies that inverse relationships exist between abundances of desert pupfish and nonnative species, are consistent with the hypothesis that predation by nonnative species is contributing to decline of desert pupfish. We suspect that competitive interactions with nonnative fishes might also adversely affect abundance of desert pupfish.

Southwestern Naturalist↗

Tadpole swimming performance and activity affected by acute exposure to sublethal levels of carbaryl

General activity and swimming performance (i.e., sprint speed and distance) of plains leopard frog tadpoles (Rana blairi) were examined after acute exposure to three sublethal concentrations of carbaryl (3.5, 5.0, and 7.2 mg/L). Both swimming performance and spontaneous swimming activity are important for carrying out life history functions (e.g., growth and development) and for escaping from predators. Measured tadpole activity diminished by nearly 90% at 3.5 mg/L carbaryl and completely ceased at 7.2 mg/L. Sprint speed and sprint distance also decreased significantly following exposure. Carbaryl affected both swimming performance and activity after just 24 h, suggesting that 24 h may be an adequate length of exposure to determine behavioral effects on tadpoles. Slight recovery of activity levels was noted at 24 and 48 h post-exposure; no recovery of swimming performance was observed. Reduction in activity and swimming performance may result in increased predation rates and, because activity is closely associated with feeding, may result in slowed growth leading to a failure to emerge before pond drying or an indirect reduction in adult fitness. Acute exposure to sublethal toxicants such as carbaryl may not only affect immediate survival of tadpoles but also impact critical life history functions and generate changes at the local population level.

Environmental Toxicology and Chemistry↗

Fall Chinook salmon (Oncorhynchus tshawytscha), sand roller (Percopsis transmontana), and smallmouth bass (Micropterus dolomieu) interactions in a Snake River reservoir: A tale of three species

We studied some of the relationships between federally listed fall Chinook Salmon, Oncorhynchus tshawytscha , endemic Sand Roller, Percopsis transmontana , and non-native Smallmouth Bass, Micropterus dolomieu , in Lower Granite Reservoir on the Snake River. Because of its recent reappearance and population increase, the Sand Rollers could be filling the role of a “native invader” in the reservoir food web. We speculated that Sand Rollers could either negatively affect fall Chinook Salmon by potentially competing with them for resources in shoreline habitats or, alternatively, benefit the salmon by providing a buffer against Smallmouth Bass predation. Nighttime beach seining showed that habitat use by fall Chinook Salmon and Sand Rollers overlapped completely in spring when both species were present along shorelines. Diet data from stomach samples also showed high overlap, but data on stable isotopes of 13 C and 15 N suggested that each species could be obtaining much of their dietary energy from different reservoir locations. Although habitat and diet overlap are evidence of competition, diel and spatial partitioning of resource use between fall Chinook Salmon and Sand Rollers may act to reduce potential competition. Analyses of Smallmouth Bass diets showed that fall Chinook Salmon and Sand Rollers comprised the majority of prey fish consumed by bass. Across years, as Smallmouth Bass increased their consumption of Sand Rollers (range 0.219 to 0.392 fish smallmouth -1 day -1 ), they decreased their consumption of fall Chinook Salmon (range 0.114 to 0.050 fish smallmouth -1 day -1 ). The greatest effect Sand Rollers may have on fall Chinook Salmon in Lower Granite Reservoir is to serve as a buffer against Smallmouth Bass predation.

Idaho, Washington↗

Paleomagnetism of the Red Dog Zn-Pb massive sulfide deposit in northern Alaska

Paleomagnetic methods have isolated two ancient magnetizations in and around the Paleozoic shale-hosted Red Dog ore deposit in northern Alaska. A high-latitude, westerly magnetization carried by magnetite, termed characteristic remanent magnetization A, was found in rocks that have barite and/or substantial quartz replacement of barite. An intermediate- to low-latitude, southerly magnetization (characteristic remanent magnetization B) is carried by pyrrhotite and was found in rocks dominated by galena and sphalerite. The ages the two components are constrained by their relationship with geochemistry, radiometric age dating, and hypotheses for the Mesozoic tectonic history of the Brooks Range. Characteristic remanent magnetization A fails the fold test so it must postdate the end of Brookian orogenesis (??? 150 Ma). It is always found with replacement quartz that has a radiometric date (white mica from a vug, 39Ar/40Ar) of 126 Ma. The paleolatitude for characteristic remanent magnetization B is too shallow to be Mesozoic or younger, regardless of the model for the tectonic origin of northern Alaska, and must predate Brookian orogenesis. Geologic mapping suggests that most of the ore is syngenetic, formed at 330 to 340 Ma, and a radiometric date (Re-Os on pyrite) yields an age of 338 Ma. Since characteristic remanent magnetization B predates deformation, is found in mineralized rocks and is carried by pyrrhotite, it was probably acquired during the mineralizing process as well. The combined radiometric ages and paleomagnetic data sets can be best interpreted by assuming that northern Alaska was part of an accreted terrane that was translated northward by about 30?? into its current location relative to the rest of North America and then rotated counterclockwise by 50?? to 70??. This tectonic interpretation yields plausible magnetization ages for both characteristic remanent magnetization A and B. Geologic evidence, isotopic ages, and paleomagnetic data indicate formation of the deposit at a paleolatitude that is much lower than today. ?? 2004 by Economic Geology.

Economic Geology↗

California Gull population growth and ecological impacts in the San Francisco Bay estuary, 1980–2016

The breeding population of California Gulls (Larus californicus) in the San Francisco Bay estuary increased from 24 individuals in 1980 to a peak of over 53,000 in 2014, then declined to 38,040 in 2016. The expansion of the breeding population may be related to the availability of suitable nesting sites in close proximity to anthropogenic food subsidies at landfills. Telemetry data indicate that California Gull movements are largely dictated by the two primary landfills in South San Francisco Bay. The large population of California Gulls has had negative effects on locally breeding shorebirds and terns, especially the Forster’s Tern (Sterna forsteri), American Avocet (Recurvirostra americana), and Western Snowy Plover (Charadrius nivosus nivosus). In South San Francisco Bay, California Gulls were responsible for 13% and 38% of egg predation events at nests of American Avocets and Snowy Plovers, respectively, and 55% and 54% of chick predation events of American Avocets and Forster’s Terns, respectively. The forced relocation of the largest gull colony (~24,000) at Pond A6 in 2010 resulted in increased survival of Forster’s Tern chicks at the adjacent colony at Pond A7 in 2011. The California Gull population and its effects on locally breeding shorebirds and terns are among the most pressing concerns for wetland managers within the San Francisco Bay estuary, especially for the South Bay Salt Pond Restoration Project. Further research is needed to evaluate the gull’s reproductive rates, habitat use, and annual movements and so to clarify its demographics and to quantify its effects on other breeding birds.

Book chapter↗

Northern Pintail

This medium-sized dabbling duck of slender, elegant lines and conservative plumage coloration is circumpolar in distribution and abundant in North America, with core nesting habitat in Alaska and the Prairie Pothole Region of southern Canada and the northern Great Plains. Breeders favor shallow wetlands interspersed throughout prairie grasslands or arctic tundra. An early fall migrant, the species arrives on wintering areas beginning in August, after wing molt, often forming large roosting and feeding flocks on open, shallow wetlands and flooded agricultural fields. The birds consume grains, marsh plant seeds, and aquatic invertebrates throughout fall and winter. Northern Pintails are among the earliest nesting ducks in North America, beginning shortly after ice-out in many northern areas. Individuals form new pair bonds each winter but are highly promiscuous during the nesting season, with mated and unmated males often involved in vigorous, acrobatic Pursuit Flights. Annual nest success and productivity vary with water conditions, predation, and weather. Females build nests on the ground, often far from water. Only the female incubates; her mate leaves shortly after incubation begins. Ducklings hatch together in one day, follow the female to water after a day in the nest, and fledge by July or August. Adults and ducklings consume mainly aquatic invertebrates during the breeding season. Predators and farming operations destroy many thousands of Northern Pintail nests annually; farming has also greatly reduced the amount of quality nesting cover available. Winter habitats are threatened by water shortages, agricultural development, contamination, and urbanization. Periods of extended drought in prairie nesting regions have caused dramatic population declines, usually followed by periods of recovery. Over the long term, however, the continental population of Northern Pintails has declined significantly from 6 million birds in the early 1970s to less than 3 million in the late 1980s and early 1990s. Since then, the population appears to have stabilized; in 2013, the estimate was 3.3 million birds, a large number but below conservation goals despite favorable wetland conditions in much of the prairie breeding region. Ongoing conservation measures, however, such as habitat restoration and enhancement of agricultural lands, as well as prudent harvest management, suggest that Northern Pintails should have a secure future in North America.

The Birds of North America↗

From the Field: Carbofuran detected on weathered raptor carcass feet

The cause of death for raptors poisoned at illegal carbofuran-laced predator baits is often not confirmed because the carcass matrices that are conventionally analyzed are not available due to decomposition and scavenging. However, many such carcasses retain intact feet that may have come into contact with carbofuran. Eastern screech owls ( Otus asio ) were exposed to carbofuran via simulated predator baits. Detection of carbofuran from owl feet weathered for 28 days demonstrated the temporal reliability of using feet during a forensic investigation. Raptor carcasses previously not submitted for residue analysis because of a lack of the conventional matrices may now be salvaged for their feet.

Wildlife Society Bulletin↗

Nesting ecology of boreal forest birds following a massive outbreak of spruce beetles

We studied breeding dark-eyed juncos ( Junco hyemalis ), yellow-rumped warblers ( Dendroica coronata ), and spruce-nesting birds from 1997 to 1998 among forests with different levels of spruce ( Picea spp.) mortality following an outbreak of spruce beetles ( Dendroctonus rufipennis ) in Alaska, USA. We identified species using live and beetle-killed spruce for nest sites and monitored nests to determine how the outbreak influenced avian habitat selection and reproduction. We tested predictions that 1) nesting success of ground-nesting juncos would increase with spruce mortality due to proliferation of understory vegetation available to conceal nests from predators, 2) nesting success of canopy-nesting warblers would decrease with spruce mortality due to fewer live spruce in which to conceal nests, and 3) both species would alter nest-site selection in response to disturbance. Juncos did not benefit from changes in understory vegetation; nesting success in highly disturbed stands (46%) was comparable to that in undisturbed habitats throughout their range. In stands with low spruce mortality, nesting success of juncos was low (5%) and corresponded with high densities of red squirrels ( Tamiasciurus hudsonicus ). Yellow-rumped warblers nested exclusively in spruce, but success did not vary with spruce mortality. As disturbance increased, nesting warblers switched from selecting forest patches with high densities of live white spruce ( Picea glauca ) to patches with beetle-killed spruce. Warblers also placed nests in large-diameter live or beetle-killed spruce, depending on which was more abundant in the stand, with no differences in nesting success. Five of the 12 other species of spruce-nesting birds also used beetle-killed spruce as nest sites. Because beetle-killed spruce can remain standing for >50 years, even highly disturbed stands provide an important breeding resource for boreal forest birds. We recommend that boreal forest managers preserve uncut blocks of infested forest within managed forest landscapes and practice partial harvest of beetle-killed spruce rather than commercial clear-cutting of infested stands in order to sustain breeding bird populations until natural reforestation occurs. Because breeding densities do not always reflect fitness, assessing impacts of a massive natural disturbance should include measuring impacts of changes in vegetation on both reproductive success and predator–prey dynamics.

Journal of Wildlife Management↗

Demographics of an experimentally released population of elk in Great Smoky Mountains National Park

We assessed the potential for reestablishing elk ( Cervus elaphus ) in Great Smoky Mountains National Park (GSMNP), USA, by estimating vital rates of experimentally released animals from 2001 to 2006. Annual survival rates for calves ranged from 0.333 to 1.0 and averaged 0.592. Annual survival for subadult and adult elk (i.e., ≥1 yr of age) ranged from 0.690 to 0.933, depending on age and sex. We used those and other vital rates to model projected population growth and viability using a stochastic individual-based model. The annual growth rate (λ) of the modeled population over a 25-year period averaged 0.996 and declined from 1.059 the first year to 0.990 at year 25. The modeled population failed to attain a positive 25-year mean growth rate in 46.0% of the projections. Poor calf recruitment was an important determinant of low population growth. Predation by black bears ( Ursus americanus ) was the dominant calf mortality factor. Most of the variance of growth projections was due to demographic variation resulting from the small population size ( n = 61). Management actions such as predator control may help increase calf recruitment, but our projections suggest that the GSMNP elk population may be at risk for some time because of high demographic variation.

Great Smoky Mountains National Park↗

Contrasting activity patterns of sympatric and allopatric black and grizzly bears

The distribution of grizzly (Ursus arctos) and American black bears (U. americanus) overlaps in western North America. Few studies have detailed activity patterns where the species are sympatric and no studies contrasted patterns where populations are both sympatric and allopatric. We contrasted activity patterns for sympatric black and grizzly bears and for black bears allopatric to grizzly bears, how human influences altered patterns, and rates of grizzlyblack bear predation. Activity patterns differed between black bear populations, with those sympatric to grizzly bears more day-active. Activity patterns of black bears allopatric with grizzly bears were similar to those of female grizzly bears; both were crepuscular and day-active. Male grizzly bears were crepuscular and night-active. Both species were more night-active and less day-active when ???1 km from roads or developments. In our sympatric study area, 2 of 4 black bear mortalities were due to grizzly bear predation. Our results suggested patterns of activity that allowed for intra- and inter-species avoidance. National park management often results in convergence of locally high human densities in quality bear habitat. Our data provide additional understanding into how bears alter their activity patterns in response to other bears and humans and should help park managers minimize undesirable bearhuman encounters when considering needs for temporal and spatial management of humans and human developments in bear habitats. ?? 2010 The Wildlife Society.

Journal of Wildlife Management↗