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An annotated list of the mayflies, stoneflies, and caddisflies of the Sand Creek basin, Great Sand Dunes National Park and Preserve, Colorado, 2004 and 2005

The U.S. Geological Survey, in conjunction with the Great Sand Dunes National Park and Preserve and its cooperators, did an extensive inventory of certain targeted aquatic-insect groups in the Sand Creek Basin, Great Sand Dunes National Park and Preserve, to establish a species list for future monitoring efforts. Study sites were established to monitor these groups following disturbance events. Such potential disturbances may include, but are not limited to, chemical treatment of perennial stream reaches to remove nonnative fishes and the subsequent reintroduction of native fish species, increased public use of backcountry habitat (such as hiking and fishing), and natural disturbances such as fire. This report is an annotated list of the mayflies, stoneflies, and caddisflies found in the Sand Creek Basin, Great Sand Dunes National Park and Preserve, 2004 and 2005. The primary objective of the study was to qualitatively inventory target aquatic-insect groups in perennial streams, and selected unique standing-water habitats, such as springs, and wetlands associated with the Sand Creek Basin. Efforts focused on documenting the presence of aquatic-insect species within the following taxonomic groups: Ephemeroptera (mayflies), Plecoptera (stoneflies), and Trichoptera (caddisflies). These insect orders were chosen because published species accounts, geographic distribution, and identification keys exist for many Colorado species. Given the extent of available information for these groups, there existed a potential for identifying new species and documenting range extensions of known species.

Data Series↗

Ground-water quality data in the Central Sierra study unit, 2006— Results from the California GAMA Program

Ground-water quality in the approximately 950 square kilometer (370 square mile) Central Sierra study unit (CENSIE) was investigated in May 2006 as part of the Priority Basin Assessment project of the Groundwater Ambient Monitoring and Assessment (GAMA) Program. The GAMA Priority Basin Assessment project was developed in response to the Ground-Water Quality Monitoring Act of 2001, and is being conducted by the U.S. Geological Survey (USGS) in cooperation with the California State Water Resources Control Board (SWRCB). This study was designed to provide a spatially unbiased assessment of the quality of raw ground water used for drinking-water supplies within CENSIE, and to facilitate statistically consistent comparisons of ground-water quality throughout California. Samples were collected from thirty wells in Madera County. Twenty-seven of the wells were selected using a spatially distributed, randomized grid-based method to provide statistical representation of the study area (grid wells), and three were selected to aid in evaluation of specific water-quality issues (understanding wells). Ground-water samples were analyzed for a large number of synthetic organic constituents (volatile organic compounds [VOCs], gasoline oxygenates and degradates, pesticides and pesticide degradates), constituents of special interest (N-nitrosodimethylamine, perchlorate, and 1,2,3-trichloropropane), naturally occurring inorganic constituents [nutrients, major and minor ions, and trace elements], radioactive constituents, and microbial indicators. Naturally occurring isotopes [tritium, and carbon-14, and stable isotopes of hydrogen, oxygen, nitrogen, and carbon], and dissolved noble gases also were measured to help identify the sources and ages of the sampled ground water. In total, over 250 constituents and water-quality indicators were investigated. Quality-control samples (blanks, replicates, and samples for matrix spikes) were collected at approximately one-sixth of the wells, and the results for these samples were used to evaluate the quality of the data for the ground-water samples. Results from field blanks indicated contamination was not a noticeable source of bias in the data for ground-water samples. Differences between replicate samples were within acceptable ranges, indicating acceptably low variability. Matrix spike recoveries were within acceptable ranges for most constituents. This study did not attempt to evaluate the quality of water delivered to consumers; after withdrawal from the ground, water typically is treated, disinfected, or blended with other waters to maintain water quality. Regulatory thresholds apply to water that is served to the consumer, not to raw ground water. However, to provide some context for the results, concentrations of constituents measured in the raw ground water were compared with health-based thresholds established by the U.S. Environmental Protection Agency (USEPA) and California Department of Public Health (CDPH), and thresholds established for aesthetic concerns (Secondary Maximum Contaminant Levels, SMCL-CA) by CDPH. Therefore, any comparisons of the results of this study to drinking-water standards only is for illustrative purposes and is not indicative of compliance or non-compliance to those standards. Most constituents that were detected in ground-water samples were found at concentrations below drinking-water standards or thresholds. Six constituents? fluoride, arsenic, molybdenum, uranium, gross-alpha radioactivity, and radon-222?were detected at concentrations higher than thresholds set for health-based regulatory purposes. Three additional constituents?pH, iron and manganese?were detected at concentrations above thresholds set for aesthetic concerns. Volatile organic compounds (VOCs) and pesticides, were detected in less than one-third of the samples and generally at less than one one-hundredth of a health-based threshold.

California↗

Summary of the major water-quality findings from the Eastern Iowa Basins study unit of the National Water-Quality Assessment Program

An integrated assessment of the water quality in streams and aquifers in the Wapsipinicon, Iowa, Cedar, and Skunk River basins was conducted in 1996 through 1998 as part of the Eastern Iowa Basins (EIWA) study unit of the U.S. Geological Survey's National Water-Quality Assessment Program (NAWQA). The EIWA study unit is one of 59 study units across the Nation designed to assess the status and trends in the quality of the Nation's ground- and surface-water resources and to link the status and trends with an understanding of the natural and human factors that affect the quality of water. Over 90 percent of the land in the EIWA study unit is used for agricultural purposes, while forested areas account for only 4 percent and urban areas about 2 percent of the land. Surface-water samples were collected monthly and during selected storm events from six sites in medium-sized basins (125 to about 400 mi2) and five sites in large river basins (2,300 to 12,500 mi2). The medium-sized basins were selected to be representative of various physical features, hydrogeology, and agricultural landuse (row crops and concentrated animal feeding operations) that may affect water quality. The large river sites were selected to determine the integrated effects of combinations of landuse and hydrogeology on river-water quality. Ground-water samples were collected primarily from the alluvial aquifers because of the aquifers' direct hydraulic connection with rivers and streams and because alluvial aquifers are one of the most important sources for domestic, municipal, and industrial water supplies in the study area. Monitoring wells were installed in agricultural and urban areas of the alluvial aquifers to assess the quality of the most recently recharged water in relation to land use. Existing domestic wells screened in alluvial aquifers and the Silurian/Devonian aquifer were sampled to assess deeper and older ground water. Surface- and ground-water samples were analyzed for a wide variety of chemical constituents (major ions, nutrients, and pesticides) commonly associated with agricultural and urban activities. Because they were not expected to occur in rivers and streams, volatile organic compounds (VOC's), commonly comprising fuels, solvents, and other industrial compounds were only analyzed in ground-water samples. The age of the ground water, important information needed to relate ground-water quality to land use, was determined using both tritium and chlorofluorocarbons (Freon?) age-dating methods. Results from the EIWA NAWQA study build on previous and ongoing research and water-quality monitoring programs in Iowa and provide new insights into the relation between the quality of the State's water resources and human activities. The major findings from the study are listed below.

Iowa↗

Hybridization between historically allopatric Chinook salmon populations in the White Salmon River, WA

Chinook Salmon spawning in the White Salmon River consist of members of three historically distinct populations: spring Chinook Salmon, Tule fall Chinook Salmon and Upriver Bright (URB) fall Chinook Salmon. Previous work examined juveniles captured in 2006-2008 and reported hybridization between introduced URBs, and the native threatened Tules. Recent increases in nearby hatchery URB release numbers raised the question of whether hybridization rates were increasing. We estimated hybrid frequencies among juveniles collected in the lower White Salmon River between 2016 and 2019. We also evaluated the frequencies at which non-target fish and hybrids were incorporated into the broodstocks of adjacent hatcheries. We observed that frequencies of hybrids in juvenile samples from the White Salmon River were greater in 2017-2019 (17-32%) than they had been in 2006-2008 (4-15%), but that a few (2/9) comparisons exhibited overlapping confidence intervals, suggesting that the rate has increased over time, but also that more sampling is needed to understand the importance of year-to-year variation. Further, differences in the habitat following dam removal and in the sampling sites complicated interpretation of our results. Examination of broodstocks of nearby hatcheries revealed low rates (< 0.5%) of incorporation of non-target populations and higher rates (< 9.0%) of incorporation of hybrids into those broodstocks. The relative compositions of all hatchery and natural-origin collections were similar: most individuals were one of the two parental stocks, a very small fraction were F1 hybrids, and a larger minority fraction were back-crosses. This pattern, in the context of hybridization which we know has been happening for several generations, is consistent with a hypothesis of selection against hybrids in which F1 hybrids are less fit than backcross hybrids.

Washington↗

Age and growth of the kiyi, Leucichthys kiyi Koelz, in Lake Michigan

Ages were determined and individual growth histories were calculated from the examination and measurement of the scales of 1,649 kiyis captured at seven localities in Lake Michigan in 1931 and 1932. The numbers of individuals employed for the investigation of other phases of the life history (such as length-frequency distributions, length-weight relationship, and sex ratio) varied according to the amount of data available or required. Age-group IV was dominant in the 1931 collections from Racine, Port Washington, and Kewaunee, Wisconsin, and age-group V dominated the 1932 samples from the Fox Islands and from three localities southward of Manistique, Michigan. A trend was noticeable toward an increase in average age from south to north. Among the explanations suggested for the observed differences in age composition were: Variation with latitude in the natural span of life; differences in fishing intensity; fluctuations in the strength of year classes (to account possibly for the shift in the dominant age group from 1931 to 1932). The oldest male kiyi belonged to the VII group and the oldest female was a member of the X group. The possible distorting effects of such factors as gear selection traceable to differences in the mesh sizes of nets fished in 1931 and 1932, selection by nets on the basis of the condition (K) of the fish, and local variations in fishing intensity and hence in the selective destruction of rapidly growing individuals in the fishery were held to be sufficiently great to render doubtful the significance of most of the observed local differences in growth rate. Kiyis from all samples were combined to determine the general growth in length. The growth in weight of the Fox Islands fish, however, was considered separately as these fish were consistently lighter than kiyis of corresponding length from other localities. The Lake Michigan kiyi grows slowly, with the females growing slightly more rapidly than the males. The grand average calculated lengths indicated, for example, that the females did not attain a total length of 10 inches until the fifth year of life or the males until the sixth. Similarly, the calculated weight of 4 ounces was not reached until the fifth or sixth year (with the actual time varying with sex and locality). The season's growth of the kiyi probably begins sometime in May and most or all of the growth is completed by the end of August. The calculated lengths of the age groups exhibited large discrepancies that differed from “Lee's phenomenon” as ordinarily observed in that the data for the later rather than the earlier years of life were affected most severely. Chief among the factors held responsible for these discrepancies were gear selection and the selective destruction of the more rapidly growing individuals in the fishery. Errors inherent in the (direct-proportion) method of computing growth from scale measurements were considered to have been unimportant. The Lake Michigan kiyi exhibits growth compensation–the tendency for the smaller of the young fish to have the more rapid growth in the later years of life. Comparisons with the average lengths and weights of the age groups of the Lake Ontario kiyi given by Pritchard (1931) indicated the Lake Michigan fish to be the larger at the earlier ages (age-groups II and III) and the smaller at the later ages (age-groups IV to VI). The length-frequency distributions of the age groups exhibited extensive overlap. As many as eight age groups were represented in a single centimeter interval of length. The length frequencies and average lengths of all fish collected, arranged according to the mesh sizes of the gill nets by which they were captured, revealed that the selective action of these nets in the taking of kiyis was much more obvious in the numbers of fish in the catch than in their average size. As an illustration, in 1930–1931, the 2 3/4-inch mesh nets took fish that were only 0.1 inch longer than those in 2 1/2-inch meshes but captured less than one fourth as many. Gill nets fished in northern Lake Michigan in 1932 captured kiyis that averaged 0.2 to 0.4 inch longer than those taken in the same meshes in southern Lake Michigan in 1930–1931. Because of the more slender form of kiyis from the northeastern island region of Lake Michigan, data on the general length weight relationship were compiled separately for fish of that area and for those of the great central basins of the lake. In both regions the weight increased to a power slightly greater than the cube of the length. Available information on condition indicated that the coefficient (K) was higher in August and early September than in May, June, and July. Condition declined from early September to October and early November–the latter period the time of most active spawning. Spawning itself was accompanied by an additional loss of about 12 per cent of the body weight of females and of somewhat less than 2 per cent of the weight of males. Analysis of the variations of K within a group that was homogeneous with respect to age, sex, maturity, and time of collection revealed that a net of a particular mesh size tends to take the heavier of the shorter fish and the lighter of the longer fish within its range of effectiveness. Among fish of the same length the values of K tended to increase with increase in the mesh size of the nets employed for their capture. Practically all fish in the samples were mature (only 11 immature in more than 6,000). These “immature” fish were probably “non-functional” since all of them approached or exceeded the average length of the mature kiyis. Females were strongly predominant in the collections at all seasons but were relatively more plentiful during the summer (90 per cent of the total) than during the spawning period (75 per cent). Possible factors contributing to this predominance of females and to the change in the sex ratio at the spawning season were discussed. A decrease in the relative abundance of males with increase in age appears to be characteristic of the kiyi. This decrease indicates a differential mortality of th sexes (greater relative destruction of males in the spawning period when they are unusually abundant or a greater natural mortality rate for the males). Current fishery regulations on mesh size and closed seasons afford the kiyi good protection but offer no guarantee against depletion from too intensive fishing.

Transactions of the American Fisheries Society↗

Characterization of and relations among precipitation, streamflow, suspended-sediment, and water-quality data at the U.S. Army Garrison Fort Carson and Piñon Canyon Maneuver Site, Colorado, water years 2016–18

Frequent and prolonged military training maneuvers are an intensive type of land use that may disturb land cover, compact soils, and have lasting effects on adjacent stream hydrology and ecosystems. To better understand the potential effect of military training on hydrologic and environmental processes, the U.S. Geological Survey in cooperation with the U.S. Army established hydrologic and water-quality data-collection networks at the U.S. Army Garrison Fort Carson (AGFC) in 1978 and at the Piñon Canyon Maneuver Site (PCMS) in 1982. The purpose of this report is to present precipitation, streamflow, suspended-sediment, and water-quality data collected by the U.S. Geological Survey at the AGFC and PCMS for water years (WYs) 2016–18 and to evaluate those data in relation to long-term data from the AGFC and PCMS. In WYs 2016–18, the U.S. Geological Survey monitored 26 sites on the AGFC and 17 sites on the PCMS for precipitation amount, streamflow, suspended sediment, and (or) water quality. On the AGFC, total annual precipitation in WYs 2016–18 was larger than the long-term mean for all 3 years at Rod and Gun Meteorologic Station at Fort Carson, CO (Rod and Gun). There were statistically significant upward trends in annual precipitation at Rod and Gun and Young Hollow Meteorologic Station at Fort Carson, CO (Young Hollow) with slopes of 1.25 and 0.66 inches per year (in/yr), respectively. The precipitation totals for WY 2017 were either the largest on record or in the top three for both sites and at Sullivan Park Meteorologic Station at Fort Carson, CO. On the PCMS, total annual precipitation was larger than the long-term mean in WYs 2016–18 at Brown Sheep Camp Meteorologic Station near Tyrone, CO; CIG Pipeline South Meteorologic Station near Simpson, CO; Bear Springs Hills Meteorologic Station near Houghton, CO (Bear Springs); and Upper Red Rock Canyon Meteorologic Station near Houghton, CO (Upper Red Rock). There were statistically significant upward trends in precipitation at Bear Springs and Upper Red Rock with slopes of 0.16 and 0.19 in/yr, respectively. The precipitation totals for WY 2017 were the largest on record for all sites except for Upper Bent Canyon Meteorological Station near Delhi, CO. Streamflow was calculated at 18 sites on the AGFC and 7 sites on the PCMS in at least 1 of WYs 2016–18. At AGFC, mean annual (or seasonal) streamflow in WYs 2016–18 was less than the long-term mean at 7 sites and greater than the long-term mean at 3 sites. There were statistically significant downward trends in mean annual or seasonal streamflow at Womack Ditch from Little Fountain Creek near Fort Carson, CO, and Ripley Ditch from Little Fountain Creek at Fort Carson, CO, with slopes of −0.036 and −0.028 cubic feet per second per year (ft 3 /s/y), respectively; and a significant upward trend in streamflow at Turkey Creek West Seepage below Teller Reservoir near Stone City, CO, with a slope of less than 0.001 ft 3 /s/y. Unlike for precipitation, the mean annual or seasonal streamflow for WY 2017 was not in the top 3 for any of the 12 sites with measured flow. At the PCMS, mean annual (or seasonal) streamflow was less than the long-term mean streamflow in WYs 2016–18 at the Taylor Arroyo below Rock Crossing near Thatcher, CO, and Bent Canyon Creek at Mouth near Timpas, CO, sites; and in WYs 2016 and 2018 at the Purgatoire River near Thatcher, CO (Purgatoire Thatcher), and Purgatoire River at Rock Crossing near Timpas, CO (Purgatoire Rock Crossing). There were no statistically significant trends in mean annual (or seasonal) streamflow at sites on the PCMS, and unlike for precipitation, the mean streamflow for WY 2017 was not in the top three for any sites except Purgatoire Rock Crossing. In WYs 2016–18, streamflow from sites on the AGFC and PCMS represented only a small fraction of streamflow in Fountain Creek or the Purgatoire River, and changes in streamflow that resulted from military maneuvers on the AGFC and PCMS were not likely to be detected in the downstream receiving waters. Suspended-sediment concentrations, loads, and yields for WYs 2016–18, were analyzed at two sites on the AGFC and five sites on the PCMS. On the AGFC, mean seasonal suspended-sediment concentrations ranged from 3.10 to 155 milligrams per liter (mg/L), mean seasonal suspended-sediment loads ranged from 0.04 to 27.1 tons per day (t/d), and seasonal suspended-sediment yields ranged from 0.28 to 216 tons per season per square mile (t/s/mi 2 ). Suspended-sediment yields at the two AGFC sites in WYs 2016–18 were all less than the long-term means. On the PCMS, mean seasonal suspended-sediment concentrations (at sites with some streamflow during a WY) ranged from 1.12 to 41.8 mg/L, mean suspended-sediment loads ranged from 0.01 to 13.1 t/d, and seasonal suspended-sediment yields ranged from 0.06 to 57.4 t/s/mi 2 . Suspended-sediment yields at the five PCMS sites in WYs 2016–18 were all less than the long-term means. In WYs 2016–18, mean daily suspended-sediment loads at Little Fountain were 1.3, 2.5, and 7.6 percent, respectively, of the mean daily suspended-sediment load at Fountain Creek at Security, Colorado. Likewise, the total of mean daily suspended-sediment loads from the five tributary sites to the Purgatoire River in WYs 2016–18 were about 0.25, 0.17, and 3.2 percent, respectively, of the historical mean daily suspended-sediment load at Purgatoire Thatcher. Spearman’s rank correlation coefficient was used to evaluate the strength and form of the relations between daily total precipitation and daily mean streamflow and between daily mean streamflow and suspended-sediment concentration and load for WYs 2016–18. For the sites on the AGFC and PCMS, there were weak or statistically insignificant positive correlations between precipitation and streamflow at nearby streamgauges, but strong statistically significant positive correlations between streamflow and suspended-sediment concentration and load. The ephemeral nature of the streams, quantity and timing of precipitation, air temperature, seasonal soil-moisture deficits, and effective runoff detention in erosion-control ponds could all contribute to inconsistent conversion of precipitation to streamflow. Water-quality data were analyzed for as many as 43 parameters from 9 samples collected from 3 sites on the AGFC and from 37 samples collected from 4 sites on PCMS during WYs 2016–18. The concentrations of selected water-quality parameters were compared to regulatory standards for aquatic life from the Colorado Department of Public Health and Environment (CDPHE) or aquatic-life criteria from the U.S. Environmental Protection Agency (EPA). There is at least 1 CDPHE standard or EPA criterion for 30 of the 43 water-quality parameters. For all samples from both the AGFC and the PCMS in WYs 2016–18, the concentrations of most water-quality parameters were compliant with the associated standards or criteria. However, there were some exceedances of standards or criteria: 11 samples exceeded the CDPHE recreational class standard for Escherichia coli concentration, 9 samples exceeded the CDPHE chronic unfiltered phosphorus aquatic-life standard, 36 samples exceeded the CDPHE chronic sulfate aquatic-life standard, 5 samples exceeded the EPA criterion for selenium, 7 samples exceeded the EPA criterion for aluminum, 2 samples exceeded the CDPHE chronic standard for iron, and 15 samples exceeded the CDPHE chronic standard for manganese. Identifying potential effects of military training on water quality in adjacent streams on the AGFC and PCMS is difficult due to the ephemeral nature of streamflow, limited number of sampling locations and samples, and limited access to the study areas. At the PCMS, pairs of water-quality samples were collected in March and May 2017 before and after an April–May 2017 military training event. At the Purgatoire Rock Crossing site, streamflow at the time of the May sample was approximately 35 times larger than streamflow for the March sample. The absolute percent differences of concentrations for 27 parameters ranged from −71.7 to 183 percent, and 7 parameters had increases in concentration whereas 22 parameters had no change or decreases in concentrations. The absolute percent differences of loads for 24 parameters ranged from 141 to 198 percent. The generally lower concentrations and higher loads were expected given the higher streamflows at the time of collection of the May compared to the March samples.

Colorado↗

Temporal invariance of social-ecological catchments

Natural resources such as waterbodies, public parks, and wildlife refuges attract people from varying distances on the landscape, creating "social-ecological catchments." Catchments have provided great utility for understanding physical and social relationships within specific disciplines. Yet, catchments are rarely used across disciplines, such as its application to understand complex spatiotemporal dynamics between mobile human users and patchily distributed natural resources. We collected residence ZIP codes from 19,983 angler parties during 2014–2017 to construct seven angler–waterbody catchments in Nebraska, USA. We predicted that sizes of dense (10% utilization distribution) and dispersed (95% utilization distribution) angler–waterbody catchments would change across seasons and years as a function of diverse resource selection among mobile anglers. Contrary to expectations, we revealed that catchment size was invariant. We discuss how social (conservation actions) and ecological (low water quality, reduction in species diversity) conditions are expected to impact landscape patterns in resource use. We highlight how this simple concept and user-friendly technique can inform timely landscape-level conservation decisions within coupled social-ecological systems that are currently difficult to study and understand.

Ecological Applications↗

Temporal variation in capture efficiency underrepresents spring out-migrating Bull Trout in a trap-and-haul program

Trap-and-haul programs can maintain connection among habitats for migratory salmonids in fragmented systems. To conserve diversity within and among life history strategies, downstream trap and transport of juvenile salmonids could ideally mimic the natural, underlying out-migration dynamics of the population. A two-way trap-and-haul program is implemented in the lower Clark Fork River, Montana, to conserve adfluvial Bull Trout Salvelinus confluentus. We used PIT technology to assess whether downstream trapping efforts are effectively capturing variation in the out-migration dynamics of juvenile Bull Trout in Graves Creek, a key spawning and rearing tributary in the system. We tagged 821 juvenile Bull Trout in Graves Creek and used these tagged Bull Trout in conjunction with stationary PIT antennas to monitor out-migration and evaluate efficiency of the downstream trapping program. Capture efficiency in Graves Creek varied substantially from autumn to spring, with 89–96% of autumn out-migrating Bull Trout captured and 5–10% of spring out-migrating Bull Trout captured. Overall, we found that Bull Trout transported during the autumn out-migration periods generally reflect the natural out-migration dynamics of the population; however, Bull Trout that out-migrate in the spring are currently underrepresented in the downstream transport program. By understanding the underlying out-migration dynamics of the Bull Trout population in Graves Creek, management of the downstream trapping efforts can focus on minimizing potential selection for or against out-migrants based on timing and age at out-migration. Minimizing selection will conserve variation within the adfluvial life history strategy and therefore maximize resilience of the adfluvial Bull Trout populations.

Idaho, Montana↗

A demonstration of the instream flow incremental methodology, Shenandoah River, Virginia

Current and projected demands on the water resources of the Shenandoah River have increased concerns for the potential effect of these demands on the natural integrity of the Shenandoah River system. The Instream Flow Incremental Method (IFIM) process attempts to integrate concepts of water-supply planning, analytical hydraulic engineering models, and empirically derived habitat versus flow functions to address water-use and instream-flow issues and questions concerning life-stage specific effects on selected species and the general well being of aquatic biological populations. The demonstration project also sets the stage for the identification and compilation of the major instream-flow issues in the Shenandoah River Basin, development of the required multidisciplinary technical team to conduct more detailed studies, and development of basin specific habitat and flow requirements for fish species, species assemblages, and various water uses in the Shenandoah River Basin. This report presents the results of an IFIM demonstration project, conducted on the main stem Shenandoah River in Virginia, during 1996 and 1997, using the Physical Habitat Simulation System (PHABSIM) model. Output from PHABSIM is used to address the general flow requirements for water supply and recreation and habitat for selected life stages of several fish species. The model output is only a small part of the information necessary for effective decision making and management of river resources. The information by itself is usually insufficient for formulation of recommendations regarding instream-flow requirements. Additional information, for example, can be obtained by analysis of habitat time-series data, habitat duration data, and habitat bottlenecks. Alternative-flow analysis and habitat-duration curves are presented.

Virginia↗

Diurnal habitat selection of migrating Whooping Crane in the Great Plains

Available stopover habitats with quality foraging opportunities are essential for migrating waterbirds, including Whooping Crane ( Grus americana ). Several studies have evaluated habitats used by Whooping Crane for roosting throughout its migration corridor; however, habitats associated with foraging and other diurnal activities have received less attention. We used data collected from 42 Whooping Crane individuals that included 2169 diurnal use locations within 395 stopover sites evaluated during spring 2013 to fall 2015 to assess diurnal habitat selection throughout the U.S. portion of the migration corridor. We found that Whooping Crane selected wetland land-cover types (i.e., open water, riverine, and semipermanent wetlands) and lowland grasslands for diurnal activities over all other land-cover types that we evaluated, including croplands. Whooping Crane generally avoided roads, and avoidance varied based on land-cover class. There has been considerable alteration and destruction of natural wetlands and rivers that serve as roosting and foraging sites for migrating Whooping Crane. Given recent droughts and the likelihood of future landscape changes within the migration corridor, directing conservation efforts toward protecting and enhancing wetland stopover areas may prove critical for continued growth of the last remaining wild population of Whooping Crane. Future studies of this Whooping Crane population should focus on specific wetland complexes and riverine sites throughout the migration corridor to identify precise management actions that could be taken to enhance and protect these imperilled land-cover types.

Avian Conservation and Ecology↗

Preliminary review of adaptation options for climate-sensitive ecosystems and resources. A report by the U.S. Climate Change Science Program and the Subcommittee on Global Change Research

Climate variables are key determinants of geographic distributions and biophysical characteristics of ecosystems, communities, and species. Climate change is therefore affecting many species attributes, ecological interactions, and ecosystem processes. Because changes in the climate system will continue into the future regardless of emissions mitigation, strategies for protecting climate-sensitive ecosystems through management will be increasingly important. While there will always be uncertainties associated with the future path of climate change, the response of ecosystems to climate impacts, and the effects of management, it is both possible and essential for adaptation to proceed using the best available science. This report provides a preliminary review of adaptation options for climate-sensitive ecosystems and resources in the United States. The term “adaptation” in this document refers to adjustments in human social systems (e.g., management) in response to climate stimuli and their effects. Since management always occurs in the context of desired ecosystem conditions or natural resource management goals, it is instructive to examine particular goals and processes used by different organizations to fulfill their objectives. Such an examination allows for discussion of specific adaptation options as well as potential barriers and opportunities for implementation. Using this approach, this report presents a series of chapters on the following selected management systems: National Forests, National Parks, National Wildlife Refuges, Wild and Scenic Rivers, National Estuaries, and Marine Protected Areas. For these chapters, the authors draw on the literature, their own expert opinion, and expert workshops composed of resource management scientists and representatives of managing agencies. The information drawn from across these chapters is then analyzed to develop the key synthetic messages presented below.

Synthesis and Assessment Product↗

Conspecific reproductive success and breeding habitat selection: Implications for the study of coloniality

Habitat selection is a crucial process in the life cycle of animals because it can affect most components of fitness. It has been proposed that some animals cue on the reproductive success of conspecifics to select breeding habitats. We tested this hypothesis with demographic and behavioral data from a 17-yr study of the Black-legged Kittiwake ( Rissa tridactyla ), a cliff-nesting seabird. As the hypothesis assumes, the Black-legged Kittiwake nesting environment was patchy, and the relative quality of the different patches (i.e., breeding cliffs) varied in time. The average reproductive success of the breeders of a given cliff was predictable from one year to the next, but this predictability faded after several years. The dynamic nature of cliff quality in the long term is partly explained by the autocorrelation of the prevalence of an ectoparasite that influences reproductive success. As predicted by the performance-based conspecific attraction hypothesis, the reproductive success of current breeders on a given cliff was predictive of the reproductive success of new recruits on the cliff in the following year. Breeders tended to recruit to the previous year’s most productive cliffs and to emigrate from the least productive ones. Consequently, the dynamics of breeder numbers on the cliffs were explained by local reproductive success on a year-to-year basis. Because, on average, young Black-legged Kittiwakes first breed when 4 yr old, such a relationship probably results from individual choices based on the assessment of previous-year local quality. When breeders changed breeding cliffs between years, they selected cliffs of per capita higher reproductive success. Furthermore, after accounting for the potential effects of age and sex as well as between-year variations, the effect of individual breeding performance on breeding dispersal was strongly influenced by the average reproductive success of other breeders on the same cliff. Individual breeding performance did not appear to influence the probability of dispersing for birds breeding on cliffs with high local reproductive success, whereas individual breeding performance did have a strong effect on dispersal for birds that bred on cliffs with lower local reproductive success. This suggests that the reproductive success of locally breeding conspecifics may be sufficient to override an individual’s own breeding experience when deciding whether to emigrate. These results, which are supported by behavioral observations of the role of prospecting in recruitment, suggest that both first breeders and adults rely on the reproductive success of conspecifics as “public information” to assess their own chances of breeding successfully in a given patch and to make settling decisions. A corollary prediction is that individuals should attempt to breed near successful conspecifics (a form of social attraction) in order to benefit from the same favorable local environmental conditions. Such a performance-based conspecific attraction mechanism can thus lead to an aggregative distribution of nests and may have played a role in the evolution of coloniality.

Ecology↗

Hydrologic models and analysis of water availability in Cuyama Valley, California

Changes in population, agricultural development practices (including shifts to more water-intensive crops), and climate variability are placing increasingly larger demands on available water resources, particularly groundwater, in the Cuyama Valley, one of the most productive agricultural regions in Santa Barbara County. The goal of this study was to produce a model capable of being accurate at scales relevant to water management decisions that could be considered in the evaluation of the sustainable water supply. The Cuyama Valley Hydrologic Model (CUVHM) was designed to simulate the most important natural and human components of the hydrologic system, including components dependent on variations in climate, thereby providing a reliable assessment of groundwater conditions and processes that can inform water users and help to improve planning for future conditions. Model development included a revision of the conceptual model of the flow system, construction of a precipitation-runoff model using the Basin Characterization Model (BCM), and construction of an integrated hydrologic flow model with MODFLOW-One-Water Hydrologic Flow Model (MF-OWHM). The hydrologic models were calibrated to historical conditions of water and land use and, then, used to assess the use and movement of water throughout the Valley. These tools provide a means to understand the evolution of water use in the Valley, its availability, and the limits of sustainability. The conceptual model identified inflows and outflows that include the movement and use of water in both natural and anthropogenic systems. The groundwater flow system is characterized by a layered geologic sedimentary sequence that—in combination with the effects of groundwater pumping, natural recharge, and the application of irrigation water at the land surface—displays vertical hydraulic-head gradients. Overall, most of the agricultural demand for water in the Cuyama Valley in the initial part of the growing season is supplied by groundwater, which is augmented by precipitation during wet winter and spring seasons. In addition, the amount of groundwater used for irrigation varies from year to year in response to climate variation and can increase dramatically in dry years. Model simulation results, however, also indicated that irrigation may have been less efficient during wet years. Agricultural pumpage is a major component to simulated outflow that is often poorly recorded. Therefore, an integrated, coupled farm-process model is used to estimate historical pumpage for water-balance subregions that evolved with the development of groundwater in the Valley from 1949 through 2010. The integrated hydrologic model includes these water-balance subregions and delineates natural, municipal, and agricultural land use; streamflow networks; and groundwater flow systems. The redefinition of the geohydrologic framework (including the internal architecture of the sedimentary units) and incorporation of these units into the simulation of the regional groundwater flow system indicated that faults have compartmentalized the alluvial deposits into subregions, which have responded differently to regional groundwater flow, locations of recharge, and the effects of development. The Cuyama Valley comprises nine subregions grouped into three regional zones, the Main, Ventucopa Uplands, and Sierra Madre Foothills, which are fault bounded, represent different proportions of the three alluvial aquifers, and have different water quality. The CUVHM uses MF-OWHM to simulate and assess the use and movement of water, including the evolution of land use and related water-balance regions. The model is capable of being accurate at annual to interannual time frames and at subregional to valley-wide spatial scales, which allows for analysis of the groundwater hydrologic budget for the water years 1950–2010, as well as potential assessment of the sustainable use of groundwater. Simulated changes in storage over time showed that significant withdrawals from storage generally occurred not only during drought years (1976–77 and 1988–92) but also during the early stages of industrial agriculture, which was initially dominated by alfalfa production. Since the 1990s, agriculture has shifted to more water-intensive crops. Measured and simulated groundwater levels indicated substantial declines in selected subregions, mining of groundwater that is thousands to tens of thousands of years old, increased groundwater storage depletion, and land subsidence. Most of the recharge occurs in the upland regions of Ventucopa and Sierra Madre Foothills, and the largest fractions of pumpage and storage depletion occur in the Main subregion. The long-term imbalance between inflows and outflows resulted in simulated overdraft (groundwater withdrawals in excess of natural recharge) of the groundwater basin over the 61-year period of 1949–2010. Changes in storage varied considerably from year to year, depending on land use, pumpage, and climate conditions. Climatically driven factors can greatly affect inflows, outflows, and water use by more than a factor of two between wet and dry years. Although precipitation during inter-decadal wet years previously replenished the basin, the water use and storage depletion have lessened the effects of these major recharge events. Simulated and measured water-level altitudes indicated the presence of large areas where depressed water levels have resulted in large desaturated zones in the younger and Older Alluvium layers in the Main-zone subregions. The results of modeled projection of the base-case scenario 61 years into the future indicated that current supply-and-demand are unsustainable and will result in additional groundwater-level declines and related storage depletion and land subsidence. The reduced-supply and reduced-demand projections reduced groundwater storage depletion but may not allow for sustainable agriculture under current demands, agricultural practices, and land use.

California↗

Geologic and hydrologic characterization and evaluation of the Basin and Range Province relative to the disposal of high-level radioactive waste: Part I, Introduction and guidelines

The U.S. Geological Survey's program for geologic and hydrologic evaluation of physiographic provinces to identify areas potentially suitable for locating repository sites for disposal of high-level nuclear wastes was announced to the Governors of the eight States in the Basin and Range Province on May 5, 1981. Representatives of Arizona, California, Idaho, New Mexico, Nevada, Oregon, Texas, and Utah, were invited to cooperate with the Federal Government in the evaluation process. Each Governor was requested to nominate an Earth scientist to represent the State in a province working group composed of State and U.S. Geological Survey representatives. This report, Part I of a three-part report, provides the background, introduction and scope of the study. This part also includes a discussion of geologic and hydrologic guidelines that will be used in the evaluation process and illustrates geohydrologic environments and the effect of individual factors in providing multiple natural barriers to radionuclide migration. Part II is a reconnaissance characterization of the geologic and hydrologic factors to be used in the initial screening of the Basin and Range Province. Part III will be the initial evaluation of the Province and will identify regions that appear suitable for further study. The plan for study of the Province includes a stepwise screening process by which successively smaller land units are considered in increasing detail. Each step involves characterization of the geology and hydrology and selection of subunits for more intensive characterization. Selection of subunits for further study is by evaluation of geologic and hydrologic conditions following a set of guidelines. By representation on the Province Working Group, the States participate in a consultation and review role in: (1) Establishing geologic and hydrologic guidelines, and (2) characterizing and evaluating the Province. The States also participate in compilation of geologic and hydrologic data used in characterizing the Province. The current (1983) needs for a high-level radioactive waste repository include: (1) Disposal in a mined repository; (2) retrievability of the waste for as much as 50 years; and (3) confidence of isolation of the waste from the accessible environment. Isolation of the waste needs to be assured using geologic and hydrologic conditions that: (1) Minimize risk of inadvertent future intrusions by man; (2) minimize the possibility of disturbance by processes that would expose the waste or increase its mobility; and (3) provide a system of natural barriers to the migration of waste by ground water. The guidelines adopted by the Province Working Group are designed to provide a standard with which these conditions can be compared. The guidelines can be grouped into four principal categories: (1) Potential host media, (2) ground-water conditions, (3) tectonic conditions, and. (4) occurrence of natural resources. Ideally the host medium constitutes the first natural barrier to migration of radionculides. The host medium ideally should be a rock type that prevents or retards dissolution and transport of radionuclides. Rocks in both the saturated and unsaturated zones may have desirable characteristics for host media. Rocks-other than the host-in the ground-water flow path from the repository ideally should be major barriers to radionuclide migration. Confining beds of low permeability might be present to retard the rate of flow between more permeable beds. Additionally, sorption of radionuclides by materials such as clays and zeolites in the flow path can further retard the flow of radionuclides by several orders of magnitude. Tectonic conditions in an area should not present a probable cause for exhumation or increased mobility of radioactive waste. Natural resources are a factor for consideration because of the problem of future human intrusion and exposure to radioactivity in the quest for minerals, oil, gas, water, and geothermal resources. The ultimate evaluation of the suitability of a geohydrologic environment for developing a mined repository needs to assess all geologic and hydrologic characteristics and their interaction in providing confidence that a geohydrologic environment will effectively isolate radionuclides from human access. Several hypothetical settings with typical geohydrologic conditions in the Basin and Range Province are used to illustrate the effect of multiple barriers in the isolation of radionuclides.

Arizona, California, Idaho, Nevada, New Mexico, Or↗

Northern bobwhite predator avoidance behavior in response to varying types of threat

The flight behavior and cover use of northern bobwhite (Colinus virginianus) have been examined in several studies, but the current data lack quantitative measures of how bobwhites respond to natural threats. We examined aspects of bobwhite behavior in response to 4 threat categories: researcher, hunter, raptor, and mammal. We found that bobwhite flight distance is best predicted by threat type and covey size, and bobwhite flight speed is best predicted by threat type. Bobwhites flushed by the hunter threat and the raptor threat selected for significantly taller obstruction at landing sites than was randomly available, with average heights of 6.2cm taller (P=0.034), and 38.1cm taller (P<0.001), respectively. Raptor-flushed bobwhites also selected for significantly denser shrub cover (42.2%, P<0.001) and a lower angle of obstruction (70.4 degrees, P<0.001) at landing points than was randomly available. In the process of data collection, we also observed bobwhite roost locations have lower visual height obstruction (7.3cm, P=0.03), lower shrub intercept (10.1%, P=0.02), and greater angles of obstruction (18.5 degrees, P=0.005) than bobwhite diurnal locations; this may facilitate escape from nocturnal mammalian predators. Our results suggest that bobwhite escape strategies and cover use vary among threat types. These results support current management recommendations of creating a patchwork of vegetation covers for bobwhites but also suggest how a more complete understanding of bobwhite behavior would improve management and conservation efforts.

Journal of Wildlife Management↗

Wild juvenile salmonid abundance in Wisconsin tributaries indicates limited contributions to Lake Michigan fisheries

Natural reproduction of salmonids occurs in many Lake Michigan tributaries, yet little is known about abundance and the potential contribution of wild fish hatching in Wisconsin tributaries. The objectives of our study were to determine if: 1) abundance of wild juvenile salmonids (primarily adfluvial rainbow trout, Oncorhynchus mykiss , referred to as steelhead) varied among selected Wisconsin streams based on available spawning and age-0 habitat; 2) stream temperature regimes could limit survival of juvenile salmonids, and 3) wild juvenile salmonids outmigrate from Wisconsin tributaries into Lake Michigan or larger tributaries. In 2016 and 2017, juvenile salmonid abundance was estimated in six Wisconsin tributaries to Lake Michigan by multiple-pass depletion sampling using backpack electrofishing. Habitat assessments included steelhead redd surveys, age-0 habitat surveys, and stream temperatures were monitored using in-stream loggers. Passive integrated transponder (PIT) tagging and PIT antennas were used to detect outmigration from three streams (Willow, Stony and Hibbard creeks). Population estimates for individual streams ranged from 75-2,276 for juvenile steelhead and from 0-243 for juvenile coho salmon, Oncorhynchus kisutch. No correlation was detected between juvenile steelhead abundance and quality age-0 habitat. Stream temperatures rarely exceeded the thermal limit for steelhead (27°C). Outmigration rates for three streams ranged from 0.6%-3.1%, but these estimates were considered minimum values. Low abundance of wild juvenile steelhead and coho salmon alone suggest that the contributions of these tributaries to Lake Michigan fisheries are likely small. Furthermore, relying on returns of wild steelhead produced in these streams is probably insufficient to maintain stream fisheries.

Wisconsin↗

Identification of conservation and restoration priority areas in the Danube River based on the multi-functionality of river-floodplain systems

Large river-floodplain systems are hotspots of biodiversity and ecosystem services but are also used for multiple human activities, making them one of the most threatened ecosystems worldwide. There is wide evidence that reconnecting river channels with their floodplains is an effective measure to increase their multi-functionality, i.e., ecological integrity, habitats for multiple species and the multiple functions and services of river-floodplain systems, although, the selection of promising sites for restoration projects can be a demanding task. In the case of the Danube River in Europe, planning and implementation of restoration projects is substantially hampered by the complexity and heterogeneity of the environmental problems, lack of data and strong differences in socio-economic conditions as well as inconsistencies in legislation related to river management. We take a quantitative approach based on best-available data to assess biodiversity using selected species and three ecosystem services (flood regulation, crop pollination , and recreation), focused on the navigable main stem of the Danube River and its floodplains. We spatially prioritize river-floodplain segments for conservation and restoration based on (1) multi-functionality related to biodiversity and ecosystem services, (2) availability of remaining semi-natural areas and (3) reversibility as it relates to multiple human activities (e.g. flood protection, hydropowerand navigation). Our approach can thus serve as a strategic planning tool for the Danube and provide a method for similar analyses in other large river-floodplain systems.

Danube River↗

The earliest seeds

Lagenostomalean-type seeds in bifurcating cupule systems have been discovered in the late Devonian Hampshire Formation of Randolph County, West Virginia, USA (Fig. 1). The associated megaflora, plants from coal balls, and vertebrate and invertebrate faunas demonstrate that the material is Famennian; the microflora indicates a more specific Fa2c age. Consequently, these seeds predate Archaeosperma arnoldii1 from the Fa2d of northeastern Pennsylvania, the oldest previously reported seed. By applying precision fracture, transfer, de??gagement, and thin-section techniques to selected cupules from the more than 100 specimens on hand, we have determined the three-dimensional morphology and histology of the seeds (Fig. 2a-h, k) and cupule systems. A comparison with known late Devonian to early Carboniferous seeds reveals that ours are more primitively organized than all except Genomosperma2,3. ?? 1981 Nature Publishing Group.

Nature↗