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At least 757 records · Page 42Linked to original sources

Landscape-scale drivers of endangered Cape Sable Seaside Sparrow (Ammospiza maritima mirabilis) presence using an ensemble modeling approach

The Florida Everglades is a vast and iconic wetland ecosystem in the southern United States that has undergone dramatic changes from habitat degradation, development encroachment, and water impoundment. Starting in the past few decades, large restoration projects have been undertaken to restore the landscape, including improving conditions for threatened and imperiled taxa. One focus of restoration has been the marl prairie ecosystem, where the federally endangered Cape Sable Seaside Sparrow ( Ammospiza maritima mirabilis ; CSSS) resides. The CSSS is endemic to the Everglades where populations have been steadily declining, signaling the importance of decision support tools for natural resource managers for evaluating water management and restoration scenarios. Here we developed an ensemble logistic regression, combining a frequentist and Bayesian approach, to model CSSS presence and measure how environmental factors such as hydrometrics, fire occurrence, and vegetation structure impact CSSS habitat suitability. This is the first analysis to quantitatively assess the interdependent relationships between a broad range of environmental factors and CSSS presence across the landscape. Our results show that the probability of CSSS presence was highest in areas with dry conditions, hydroperiods between 80 and 120 days, percentages of canopy cover and woody vegetation less than 10%, and more than six years post-fire where 75% or more of the area was burned. Because the frequentist and Bayesian models had nearly identical spatial outputs with the Bayesian model having slightly higher validation metrics, we used the Bayesian approach as our final model (EverSparrow). The results from our analysis can provide a valuable decision support tool as natural resource managers work to restore the Everglades landscape.

Florida↗

Estimating soil turnover rate from tree uprooting during hurricanes in Puerto Rico

Soil turnover by tree uprooting in primary and secondary forests on the island of Puerto Rico was measured in 42 study plots in the months immediately after the passage of a Category 3 hurricane. Trunk basal area explained 61% of the variability of mound volume and 53% of the variability of mound area. The proportion of uprooted trees, the number of uprooted trees, or the proportion of uprooted basal area explained 84-85% of the variation in hurricane-created mound area. These same variables explain 79-85% of the variation in mound volume. The study indicates that the soil turnover period from tree uprooting by Puerto Rican hurricanes is between 1600 and 4800 years. These rates are faster than soil turnover by landslides and background treefall in the same area and provide a useful age constraint on soil profile development and soil carbon sequestration in these dynamic landscapes. ?? 2009 Elsevier B.V.

Forest Ecology and Management↗

Savanna tree density, herbivores, and the herbaceous community: Bottom-up vs. top-down effects

Herbivores choose their habitats both to maximize forage intake and to minimize their risk of predation. For African savanna herbivores, the available habitats range in woody cover from open areas with few trees to dense, almost‐closed woodlands. This variation in woody cover or density can have a number of consequences for herbaceous species composition, cover, and productivity, as well as for ease of predator detection and avoidance. Here, we consider two alternative possibilities: first, that tree density affects the herbaceous vegetation, with concomitant “bottom‐up” effects on herbivore habitat preferences; or, second, that tree density affects predator visibility, mediating “top‐down” effects of predators on herbivore habitat preferences. We sampled sites spanning a 10‐fold range of tree densities in an Acacia drepanolobium‐dominated savanna in Laikipia, Kenya, for variation in (1) herbaceous cover, composition, and species richness; (2) wild and domestic herbivore use; and (3) degree of visibility obstruction by the tree layer. We then used structural equation modeling to consider the potential influences that tree density may have on herbivores and herbaceous community properties. Tree density was associated with substantial variation in herbaceous species composition and richness. Cattle exhibited a fairly uniform use of the landscape, whereas wild herbivores, with the exception of elephants, exhibited a strong preference for areas of low tree density. Model results suggest that this was not a response to variation in herbaceous‐community characteristics, but rather a response to the greater visibility associated with more open places. Elephants, in contrast, preferred areas with higher densities of trees, apparently because of greater forage availability. These results suggest that, for all but the largest species, top‐down behavioral effects of predator avoidance on herbivores are mediated by tree density. This, in turn, appears to have cascading effects on the herbaceous vegetation. These results shed light on one of the major features of the “landscape of fear” in which African savanna herbivores exist.

Laikipia County↗

Sediment sources and connectivity linked to hydrologic pathways and geomorphic processes: A conceptual model to specify sediment sources and pathways through space and time

Sediment connectivity is a framework for transfer and storage of sediment among different geomorphic compartments across upland and channel network of the catchment sediment cascade. Sediment connectivity and dysconnectivity (i.e., source delivery and storage processes) are linked to the water cycle and hydrologic systems with the associated multiscale interactions with climate, soil, topography, ecology, and landuse/landcover under natural variability and human intervention. We review the sediment connectivity concept and frameworks developed in the last few decades to examine and quantify water and sediment transfer in catchment systems. Past conceptual models of connectivity have attempted to integrate multiple processes into sediment domain, including geomorphic, hydrologic, and ecological processes (i.e., “holistic approach to connectivity”). In particular, multiple studies highlight the importance of sediment and water interaction in defining landscape connectivity. There are also efforts to quantify the topographic controls on sediment connectivity, in the advent of increasingly high-resolution digital terrain models. More recent modeling efforts have integrated structural and functional connectivity through coupling topographic information with hydrologic simulation models. Though this recent modeling development is encouraging, a comprehensive sediment connectivity framework that integrates geomorphic and hydrologic processes across spatiotemporal scales is yet to be conceived. Such an effort will require understanding the governing hydrologic and geomorphic processes that control sediment source, storage, and transport. A conceptual model is proposed to describe dominant hydrologic-sediment connectivity regimes through spatial-temporal feedbacks between hydrologic processes (rainfall, flow routing, and water residence time) and geomorphic drivers (upland soil erosion and deposition, and geomorphic channel erosion and deposition response). Recent advancements in landscape monitoring techniques using geochemical tracers, remote-sensing, increasing availability of hydrologic monitoring data, and the integration of various analytic methods (e.g., isotopic hydrograph separation, stormflow concentration-discharge, hysteretic behavior analysis) have the potential to broaden the spatial and temporal scales of geomorphic observations and understanding of landscape sediment connectivity. Using the conceptual model as a “thinking” space, we examine sediment and hydrologic interactions in real world examples of watershed studies using multiple lines of evidence and modeling techniques.

Conference Paper↗

Freshwater availability and coastal wetland foundation species: ecological transitions along a rainfall gradient

Climate gradient-focused ecological research can provide a foundation for better understanding critical ecological transition points and nonlinear climate-ecological relationships, which is information that can be used to better understand, predict, and manage ecological responses to climate change. In this study, we examined the influence of freshwater availability upon the coverage of foundation plant species in coastal wetlands along a northwestern Gulf of Mexico rainfall gradient. Our research addresses the following three questions: (1) what are the region-scale relationships between measures of freshwater availability (e.g., rainfall, aridity, freshwater inflow, salinity) and the relative abundance of foundation plant species in tidal wetlands; (2) How vulnerable are foundation plant species in tidal wetlands to future changes in freshwater availability; and (3) What is the potential future relative abundance of tidal wetland foundation plant species under alternative climate change scenarios? We developed simple freshwater availability-based models to predict the relative abundance (i.e., coverage) of tidal wetland foundation plant species using climate data (1970-2000), estuarine freshwater inflow-focused data, and coastal wetland habitat data. Our results identify regional ecological thresholds and nonlinear relationships between measures of freshwater availability and the relative abundance of foundation plant species in tidal wetlands. In drier coastal zones, relatively small changes in rainfall could produce comparatively large landscape-scale changes in foundation plant species abundance which would affect some ecosystem good and services. Whereas a drier future would result in a decrease in the coverage of foundation plant species, a wetter future would result in an increase in foundation plant species coverage. In many ways, the freshwater-dependent coastal wetland ecological transitions we observed are analogous to those present in dryland terrestrial ecosystems.

Gulf Of Mexico↗

Modeling ecological minimum requirements for distribution of greater sage-grouse leks: implications for population connectivity across their western range, U.S.A.

Greater sage-grouse Centrocercus urophasianus (Bonaparte) currently occupy approximately half of their historical distribution across western North America. Sage-grouse are a candidate for endangered species listing due to habitat and population fragmentation coupled with inadequate regulation to control development in critical areas. Conservation planning would benefit from accurate maps delineating required habitats and movement corridors. However, developing a species distribution model that incorporates the diversity of habitats used by sage-grouse across their widespread distribution has statistical and logistical challenges. We first identified the ecological minimums limiting sage-grouse, mapped similarity to the multivariate set of minimums, and delineated connectivity across a 920,000 km 2 region. We partitioned a Mahalanobis D 2 model of habitat use into k separate additive components each representing independent combinations of species–habitat relationships to identify the ecological minimums required by sage-grouse. We constructed the model from abiotic, land cover, and anthropogenic variables measured at leks (breeding) and surrounding areas within 5 km. We evaluated model partitions using a random subset of leks and historic locations and selected D 2 (k = 10) for mapping a habitat similarity index (HSI). Finally, we delineated connectivity by converting the mapped HSI to a resistance surface. Sage-grouse required sagebrush-dominated landscapes containing minimal levels of human land use. Sage-grouse used relatively arid regions characterized by shallow slopes, even terrain, and low amounts of forest, grassland, and agriculture in the surrounding landscape. Most populations were interconnected although several outlying populations were isolated because of distance or lack of habitat corridors for exchange. Land management agencies currently are revising land-use plans and designating critical habitat to conserve sage-grouse and avoid endangered species listing. Our results identifying attributes important for delineating habitats or modeling connectivity will facilitate conservation and management of landscapes important for supporting current and future sage-grouse populations.

Ecology and Evolution↗

Evaluation of trap capture in a geographically closed population of brown treesnakes on Guam

1. Open population mark-recapture analysis of unbounded populations accommodates some types of closure violations (e.g. emigration, immigration). In contrast, closed population analysis of such populations readily allows estimation of capture heterogeneity and behavioural response, but requires crucial assumptions about closure (e.g. no permanent emigration) that are suspect and rarely tested empirically. 2. In 2003, we erected a double-sided barrier to prevent movement of snakes in or out of a 5-ha semi-forested study site in northern Guam. This geographically closed population of >100 snakes was monitored using a series of transects for visual searches and a 13 ?? 13 trapping array, with the aim of marking all snakes within the site. Forty-five marked snakes were also supplemented into the resident population to quantify the efficacy of our sampling methods. We used the program mark to analyse trap captures (101 occasions), referenced to census data from visual surveys, and quantified heterogeneity, behavioural response, and size bias in trappability. Analytical inclusion of untrapped individuals greatly improved precision in the estimation of some covariate effects. 3. A novel discovery was that trap captures for individual snakes consisted of asynchronous bouts of high capture probability lasting about 7 days (ephemeral behavioural effect). There was modest behavioural response (trap happiness) and significant latent (unexplained) heterogeneity, with small influences on capture success of date, gender, residency status (translocated or not), and body condition. 4. Trapping was shown to be an effective tool for eradicating large brown treesnakes Boiga irregularis (>900 mm snout-vent length, SVL). 5. Synthesis and applications. Mark-recapture modelling is commonly used by ecological managers to estimate populations. However, existing models involve making assumptions about either closure violations or response to capture. Physical closure of our population on a landscape scale allowed us to determine the relative importance of covariates influencing capture probability (body size, trappability periods, and latent heterogeneity). This information was used to develop models in which different segments of the population could be assigned different probabilities of capture, and suggests that modelling of open populations should incorporate easily measured, but potentially overlooked, parameters such as body size or condition. ?? 2008 The Authors.

Journal of Applied Ecology↗

Protected areas as social-ecological systems: perspectives from resilience and social-ecological systems theory

Conservation biology and applied ecology increasingly recognize that natural resource management is both an outcome and a driver of social, economic, and ecological dynamics. Protected areas offer a fundamental approach to conserving ecosystems, but they are also social-ecological systems whose ecological management and sustainability are heavily influenced by people. This editorial, and the papers in the invited feature that it introduces, discuss three emerging themes in social-ecological systems approaches to understanding protected areas: (1) the resilience and sustainability of protected areas, including analyses of their internal dynamics, their effectiveness, and the resilience of the landscapes within which they occur; (2) the relevance of spatial context and scale for protected areas, including such factors as geographic connectivity, context, exchanges between protected areas and their surrounding landscapes, and scale dependency in the provision of ecosystem services; and (3) efforts to reframe what protected areas are and how they both define and are defined by the relationships of people and nature. These emerging themes have the potential to transform management and policy approaches for protected areas and have important implications for conservation, in both theory and practice.

Ecological Applications↗

Pardus in the press: Drivers of leopard (Panthera pardus fusca) attack occurrence on humans in Nepal

The negative impact of large carnivore presence in human-dominated landscapes manifests as livestock depredation and in extreme cases as attacks on humans. In the case of conflict with leopards in Nepal, attacks resulting in human fatality have become more frequent over time, thus creating an urgent socio-ecological and management issue. We estimated the occurrence of leopard attacks in Nepal from human-leopard conflict cases reported in the media. We used occupancy models to analyse data collected from online news reports on incidents of leopard attacks on humans to explore drivers of leopard attacks on a landscape scale. Our results suggest that the probability of occurrence of leopard attack is associated with human population density, terrain ruggedness and livestock density. The human population density effect may be indicative of a density-dependent relationship, where attacks are more likely in areas where an increased abundance of humans increases encounter rates with leopards. The positive effect of livestock density suggests that livestock may be drawing leopards into human settlements, and consequently increasing the likelihood of attacks on humans. Terrain ruggedness might be offering ideal conditions to facilitate attacks on humans, for example remoteness and high amounts of cover to launch ambush attacks. We provide inference and insights into key determinants of leopard attacks on humans on a landscape scale. These insights can be used to guide future research, inform mitigation measures to reduce leopard attacks and foster a better understanding of the interaction between people and leopards. This study demonstrates the applicability and novelty of using a hierarchical modelling framework applied to freely and publicly available media reports to inform the applied management of human-wildlife conflict at a national scale.

People and Nature↗

Biophysical methods and data analysis for simulating overland flow in the Everglades

The Everglades in south Florida supply fresh drinking water for more than 7 million people, host a National Park, and are classified as a Ramsar wetland of international distinction. Predicting trajectories of water flow and water storage changes in the future is important to managing the Congressionally authorized restoration of the Everglades. Here we describe the needed data sources and analysis approaches to build the inputs for biophysically based modeling that can protect water and ecological resources in the face of changing water management and climate conditions. A biophysical approach to modeling overland flow in the Everglades can help predict future outcomes for ecological habitat, water storage during droughts, and water conveyance during floods. The needed data include measurements of vegetation stem architecture, microtopography, and landscape pattern metrics. Stem architecture measurements present the opportunity to estimate flow roughness of distinct vegetation communities based on hydraulic principles. At a larger scale, the microtopography and the connectivity of the sloughs between ridges offer a way to quantify the effects of flow blockage and tortuous flow paths on overland flow. Combined with theory these data provide the capacity to simulate overland flow in both the historical, pre-drainage Everglades as well as in the present-day managed Everglades. Also provided are the hydrologic data, e.g., water slopes, water depths and overland flow velocities, that can be used to verify a biophysical model. Ultimately, the purpose is to anticipate how changing flow and water depth will interact with evolving vegetation and landscape conditions to influence future water availability for society and for the ecosystem, both in the Everglades and in other low-gradient floodplains.

Florida↗

Ecological thresholds: The key to successful enviromental management or an important concept with no practical application?

An ecological threshold is the point at which there is an abrupt change in an ecosystem quality, property or phenomenon, or where small changes in an environmental driver produce large responses in the ecosystem. Analysis of thresholds is complicated by nonlinear dynamics and by multiple factor controls that operate at diverse spatial and temporal scales. These complexities have challenged the use and utility of threshold concepts in environmental management despite great concern about preventing dramatic state changes in valued ecosystems, the need for determining critical pollutant loads and the ubiquity of other threshold-based environmental problems. In this paper we define the scope of the thresholds concept in ecological science and discuss methods for identifying and investigating thresholds using a variety of examples from terrestrial and aquatic environments, at ecosystem, landscape and regional scales. We end with a discussion of key research needs in this area.

Ecosystems↗

Dust in the Critical Zone: North American case studies

The dust cycle facilitates the exchange of particles among Earth's major systems, enabling dust to traverse ecosystems, cross geographic boundaries, and even move uphill against the natural flow of gravity. Dust in the atmosphere is composed of a complex and ever-changing mixture that reflects the evolving human footprint on the landscape. The emission, transport, and deposition of dust interacts with and connects Critical Zone processes at all spatial and temporal scales. Landscape properties, land use, and climatic factors influence the wind erosion of soil and nutrient loss, which alters the long-term ecological dynamics at erosional locations. Once in the atmosphere, dust particles influence the amount of solar radiation reaching Earth, and interact with longwave (terrestrial) radiation, with cascading effects on the climate system. Finally, the wet and dry deposition of particles influences ecosystem structure, composition, and function over both short and long-term scales. Tracking dust particles from source to sink relies on monitoring and measurement of the geochemical composition and size distribution of the particles, space-borne and ground-based remote sensing, and dust modeling. Dust is linked to human systems via land use and policies that contribute to dust emissions and the health-related consequences of particulate loads and composition. Despite the significant influence dust has in shaping coupled natural-human systems, it has not been considered a key component of the Critical Zone. Here, we demonstrate that dust particles should be included as a key component of the Critical Zone by outlining how dust interacts with and shapes Earth System processes from generation, through transport, to deposition. We synthesize current understanding from global research and identify critical data and knowledge gaps while showcasing case studies from North America.

Earth-Science Reviews↗

Mapping wildland fuels for fire management across multiple scales: integrating remote sensing, GIS, and biophysical modeling

Fuel maps are essential for computing spatial fire hazard and risk and simulating fire growth and intensity across a landscape. However, fuel mapping is an extremely difficult and complex process requiring expertise in remotely sensed image classification, fire behavior, fuels modeling, ecology, and geographical information systems (GIS). This paper first presents the challenges of mapping fuels: canopy concealment, fuelbed complexity, fuel type diversity, fuel variability, and fuel model generalization. Then, four approaches to mapping fuels are discussed with examples provided from the literature: (1) field reconnaissance; (2) direct mapping methods; (3) indirect mapping methods; and (4) gradient modeling. A fuel mapping method is proposed that uses current remote sensing and image processing technology. Future fuel mapping needs are also discussed which include better field data and fuel models, accurate GIS reference layers, improved satellite imagery, and comprehensive ecosystem models.

International Journal of Wildland Fire↗

Ecological contingency in the effects of climatic warming on forest herb communities

Downscaling from the predictions of general climate models is critical to current strategies for mitigating species loss caused by climate change. A key impediment to this downscaling is that we lack a fully developed understanding of how variation in physical, biological, or land-use characteristics mediates the effects of climate change on ecological communities within regions. We analyzed change in understory herb communities over a 60-y period (1949/1951–2007/2009) in a complex montane landscape (the Siskiyou Mountains, Oregon) where mean temperatures have increased 2 °C since 1948, similar to projections for other terrestrial communities. Our 185 sites included primary and secondary-growth lower montane forests (500–1.200 m above sea level) and primary upper montane to subalpine forests (1,500–2,100 m above sea level). In lower montane forests, regardless of land-use history, we found multiple herb-community changes consistent with an effectively drier climate, including lower mean specific leaf area, lower relative cover by species of northern biogeographic affinity, and greater compositional resemblance to communities in southerly topographic positions. At higher elevations we found qualitatively different and more modest changes, including increases in herbs of northern biogeographic affinity and in forest canopy cover. Our results provide community-level validation of predicted nonlinearities in climate change effects.

Oregon↗

Economical defence of resources structures territorial space use in a cooperative carnivore

Ecologists have long sought to understand space use and mechanisms underlying patterns observed in nature. We developed an optimality landscape and mechanistic territory model to understand mechanisms driving space use and compared model predictions to empirical reality. We demonstrate our approach using grey wolves ( Canis lupus ). In the model, simulated animals selected territories to economically acquire resources by selecting patches with greatest value, accounting for benefits, costs and trade-offs of defending and using space on the optimality landscape. Our approach successfully predicted and explained first- and second-order space use of wolves, including the population's distribution, territories of individual packs, and influences of prey density, competitor density, human-caused mortality risk and seasonality. It accomplished this using simple behavioural rules and limited data to inform the optimality landscape. Results contribute evidence that economical territory selection is a mechanistic bridge between space use and animal distribution on the landscape. This approach and resulting gains in knowledge enable predicting effects of a wide range of environmental conditions, contributing to both basic ecological understanding of natural systems and conservation. We expect this approach will demonstrate applicability across diverse habitats and species, and that its foundation can help continue to advance understanding of spatial behaviour.

Montana↗

USDA conservation program and practice effects on wetland ecosystem services in the Prairie Pothole Region

Implementation of the U.S. Department of Agriculture (USDA) Conservation Reserve Program (CRP) and Wetlands Reserve Program (WRP) has resulted in the restoration of >2 million ha of wetland and grassland habitats in the Prairie Pothole Region (PPR). Restoration of habitats through these programs provides diverse ecosystem services to society, but few investigators have evaluated the environmental benefits achieved by these programs. We describe changes in wetland processes, functions, and ecosystem services that occur when wetlands and adjacent uplands on agricultural lands are restored through Farm Bill conservation programs. At the scale of wetland catchments, projects have had positive impacts on water storage, reduction in sedimentation and nutrient loading, plant biodiversity, carbon sequestration, and wildlife habitat. However, lack of information on the geographic location of restored catchments relative to landscape‐level factors (e.g., watershed, proximity to rivers and lakes) limits interpretation of ecosystem services that operate at multiple scales such as floodwater retention, water quality improvement, and wildlife habitat suitability. Considerable opportunity exists for the USDA to incorporate important landscape factors to better target conservation practices and programs to optimize diverse ecosystem services. Restoration of hydrologic processes within wetlands (e.g., hydroperiod, water level dynamics) also requires a better understanding of the influence of conservation cover composition and structure, and management practices that occur in uplands surrounding wetlands. Although conservation programs have enhanced delivery of ecosystem services in the PPR, the use of programs to provide long‐term critical ecosystem services is uncertain because when contracts (especially CRP) expire, economic incentives may favor conversion of land to crop production, rather than reenrollment. As demands for agricultural products (food, fiber, biofuel) increase, Farm Bill conservation programs will become increasingly important to ensure provisioning of ecosystem services to society, especially in agriculturally dominated landscapes. Thus, continued development and support for conservation programs legislated through the Farm Bill will require a more comprehensive understanding of wetland ecological services to better evaluate program achievements relative to conservation goals.

Ecological Applications↗

Nine endangered taxa, one recovering ecosystem: Identifying common ground for recovery on Santa Cruz Island, California

It is not uncommon to have several rare and listed taxa occupying habitats in one landscape or management area where conservation amounts to defense against the possibility of further loss. It is uncommon and extremely exciting, however, to have several listed taxa occupying one island that is managed cooperatively for conservation and recovery. On Santa Cruz Island, the largest of the northern California island group in the Santa Barbara Channel, we have a golden opportunity to marry ecological knowledge and institutional "good will" in a field test of holistic rare plant conservation. Here, the last feral livestock have been removed, active weed control is underway, and management is focused on understanding and demonstrating system response to conservation management. Yet funding limitations still exist and we need to plan the most fiscally conservative and marketable approach to rare plant restoration. We still experience the tension between desirable quick results and the ecological pace of system recovery. Therefore, our research has focused on identifying fundamental constraints on species recovery at individual, demographic, habitat, and ecosystem levels, and then developing suites of actions that might be taken across taxa and landscapes. At the same time, we seek a performance middle ground that balances an institutional need for quick demonstration of hands-on positive results with a contrasting approach that allows ecosystem recovery to facilitate species recovery in the long term. We find that constraints vary across breeding systems, life-histories, and island locations. We take a hybrid approach in which we identify several actions that we can take now to enhance population size or habitat occupancy for some taxa by active restoration, while allowing others to recover at the pace of ecosystem change. We make our recommendations on the basis of data we have collected over the last decade, so that management is firmly grounded in ecological observation.

California↗

Effects of climate change on ecological disturbance in the Northern Rockies Region [Chapter 8]

This chapter describes the ecology of important disturbance regimes in the Forest Service, U.S. Department of Agriculture (USFS) Northern Region and the Greater Yellowstone Area, hereafter called the Northern Rockies region, and potential shifts in these regimes as a consequence of observed and projected climate change. The term disturbance regime describes the general temporal and spatial characteristics of a disturbance agent - insect, disease, fire, weather, even human activity - and the effects of that agent on the landscape (table 8.1). More specifically, a disturbance regime is the cumulative effect of multiple disturbance events over space and time (Keane 2013). Disturbances disrupt an ecosystem, community, or population structure and change elements of the biological environment, physical environment, or both (White and Pickett 1985). The resulting shifting mosaic of diverse ecological patterns and structures in turn affects future patterns of disturbance, in a reciprocal, linked relationship that shapes the fundamental character of landscapes and ecosystems. Disturbance creates and maintains biological diversity in the form of shifting, heterogeneous mosaics of diverse communities and habitats across a landscape (McKinney and Drake 1998), and biodiversity is generally highest when disturbance is neither too rare nor too frequent on the landscape (Grime 1973).

General Technical Report↗