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Response of white-footed mice (Peromyscus leucopus) to fire and fire surrogate fuel reduction treatments in a southern Appalachian hardwood forest

An experiment conducted as part of the multidisciplinary National Fire and Fire Surrogate Study was designed to determine effects of three fuel reduction techniques on small mammals and habitat structure in the southern Appalachian mountains. Four experimental units, each >14-ha were contained within each of three replicate blocks at the Green River Game Land, Polk County, NC. Treatments were (1) prescribed burning (B); (2) mechanical felling of shrubs and small trees (M); (3) mechanical felling + burning (MB); (4) controls (C). Mechanical understory felling treatments were conducted in winter 2001-2002, and prescribed burning was conducted in March 2003. After treatment, there were fewer live trees, more snags, and greater canopy openness in MB than in other treatments. Leaf litter depth was reduced by burning in both B and MB treatments, and tall shrub cover was reduced in all fuel reduction treatments compared to C. Coarse woody debris pieces and percent cover were similar among treatments and controls. We captured 990 individuals of eight rodent species a total of 2823 times. Because white-footed mice composed >79% of all captures, we focused on this species. Populations in experimental units increased 228% on average between 2001 and 2002, but there was no evidence of an effect of the mechanical treatment. From 2002 to 2003, all units again showed an average increase in relative population size, but increases were greater in MB than in the other treatments. Age structure and male to female ratio were not affected by the fuel reduction treatment. Average adult body weight declined from 2001 to 2002, but less so in M than in units that remained C in both years. The proportion of mice captured near coarse woody debris was similar to the proportion captured in open areas for all treatments, indicating that white-footed mice did not use coarse woody debris preferentially or change their use patterns in response to fuel reduction treatments. Land managers should understand possible effects of different fuel reduction treatments on white-footed mouse populations, as they are an important component of the fauna and food chain of deciduous southern Appalachian forests.

Forest Ecology and Management↗

Holocene paleoenvironments of Northeast Iowa

This paper presents the biotic, sedimentary, geomorphic, and climatic history of the upper part of the Roberts Creek Basin, northeastern Iowa for the late—glacial and Holocene, and compares these records with a C—O isotopic sequence from Coldwater Cave, 60 km northwest of Roberts Creek. The biotic record (pollen, vascular plant and bryophyte macrofossils, and insects) is preserved in floodplain alluvium that underlies three constructional surfaces separated by low scarps. Each surface is underlain by a lithologically and temporally distinct alluvial fill. The highest surface is underlain by the Gunder Member of the Deforest Formation, dating from 11 000 to 4000 yr BP; beneath the intermediate level is the Roberts Creek Member, dating from 4000 to 400 yr BP; and the lowest level is underlain by the Camp Creek Member, deposited during the last 380 yr. Pollen and plant macrofossils in the alluvial fill show that a typical late—glacial spruce forest was replaced by Quercus and Ulmus in the early Holocene. This early—to—middle Holocene forest became dominated by mesic elements such as Acer saccharum, Tilia americana, Ostrya virginiana, and Carpinus caroliniana as late as 5500 yr BP; in contrast, the closest sites to the west and north were at their warmest and driest and were covered by prairie vegetation between 6500 and 5500 yr BP. After 5500 yr BP, the forest in the Roberts Creek area was replaced by prairie, as indicated by a rich assemblage of plant macrofossils, although only Ambrosia and Poaceae became abundant in the pollen record. The return of Quercus ≈ 3000 BP (while nonarboreal pollen percentages remained relatively high) indicates that oak savanna prevailed with little change until settlement time. The bryophyte assemblages strongly support the vascular plant record. Rich fen species characteristic of boreal habitats occur only in the late—glacial. They are replaced by a number of deciduous—forest elements when early—to—middle Holocene forests were present, but mosses of forest habitats completely disappear when prairie became dominant. A few deciduous—forest taxa return during the late—Holocene, when oak savanna prevailed. The C—O isotopic record from stalagmite s in Coldwater Cave indicates a relatively stable environment from ≈ 8000 to 5100 yr BP, when the δ 13 C values indicate a change in vegetation dominated by C 3 (predominantly forest) to C 4 (predominantly prairie) plants. About 4900 yr BP, the rise in 18 O values indicates a temperature increase of ≈ 1.5 ° C. The fact that the vegetational change suggested by the δ 13 C values preceded the temperature increase suggests that fire may have been an important factor in converting forest to prairie. Abundant charred seeds and other plant material at Roberts Creek 4830 yr BP support this hypothesis. The 18 O values remain constant from ≈ 5100 to ≈ 3000 yr BP, but the δ 13 C values gradually rise, indicating that soil formed under forest takes at least 2000 yr for its carbon to reach equilibrium after replacement by prairie vegetation. The return of oak to form savanna is reflected in the gradual decline of δ 13 C values in the last 3000 yr BP; O isotopic values drop sharply by ≈ 1 ° C ≈ 2800 yr BP and then were relatively stable. In contrast to the vegetational and isotopic records, the insect assemblages suggest little change in the local environments throughout most of the Holocene. All of the beetle taxa presently occur in eastern Iowa. The relative stability through the Holocene indicates that both open grassland and riparian woodland elements were present throughout. Settlement, land clearing, and land cultivation by EuroAmericans in the region caused rapid erosion of the upland landscape, the deposition of 1—2 m of sediment across the floodplain, a replacement of the native vegetation with ruderal species, a decimation of the native insect fauna, and a degradation of water quality in the stream. These changes in the landscape, vegetation, and insect faunas are as striking as those associated with glacial—interglacial transitions. The timing and direction of changes in the vegetation at Roberts Creek generally correlate well with the carbon and oxygen isotopic record in speleothems at nearby Coldwater Cave and indicate that climate was the main forcing function. However, the contrast between the vegetational change and the stability of the beetle population suggests that climatic changes were subtle. We hypothesize that the factors involved in the Holocene changes were seasonal changes in temperature and precipitation that may not have resulted in much mean annual change. Such changes may have affected the vegetation more than the insect fauna.

Iowa↗

Establishing native grasses in a big sagebrush-dominated site: An intermediate restoration step

Many semiarid rangelands in the Great Basin, U.S.A., are shifting dominance to woody species as a consequence of land degradation including intense livestock grazing and fire suppression. Whereas past rehabilitation efforts in Big sagebrush ( Artemisia tridentata ) steppes removed the shrub and added introduced forage grasses to successfully shift communities from shrublands to grasslands, current consensus is that native species should be included in restoration projects and that retention of some woody plants is desirable. We examined the potential for interseeding grasses into dense shrub communities as a precursor to thinning shrubs and releasing grasses from shrub interference. We compared seedling establishment of the native grass, Bluebunch wheatgrass ( Pseudoroegneria spicata ), with that of the Eurasia grass, Crested wheatgrass ( Agropyron desertorum ), in dense Ar. tridentata stands. Shrubs may play an important role as nurse plants for seedling establishment (reduced solar radiation, “island of fertility” effect) but result in highly contrasting light environments and root interference for seedlings. In experimental plots, we examined effects of Ar. tridentata shade levels (0, 40, 70, and 90% reduction of solar radiation) and initial root exclusion (present/absent) on the establishment and growth of P. spicata and Ag. desertorum seedlings. With this design we evaluated the interference effects of Ar. tridentata on the two grasses and identified the most beneficial microsites for grass restoration in Ar. tridentata –dominated communities. We predicted seedling survival and growth to be greater under moderate shade (40% reduction) and limited root competition than under no or strong shade conditions (0 and 90%) and unrestricted root interactions. Fifty to 85% of the P. spicata and Ag. desertorum seedlings survived the dry summer months of 1995 and 1996 and the intervening winter. Neither shading nor root exclusion from Ar. tridentata affected final seedling survival of either species. Seedling biomass of both grass species was negatively affected by initial root interactions with Ar. tridentata. However, the analysis of seedling biomass variability (coefficient of variation) indicated that in all shade and root-exclusion treatments, some seedlings of both species developed to large individuals to survive in Ar. tridentata –dominated rangelands. Thus, the use of interseeding techniques shows promise for restoring herbaceous species in dense Ar. tridentata stands and should be given further consideration when shrub retention is an important consideration.

Utah↗

Scale-dependent approaches to modeling spatial epidemiology of chronic wasting disease.

This e-book is the product of a second workshop that was funded and promoted by the United States Geological Survey to enhance cooperation between states for the management of chronic wasting disease (CWD). The first workshop addressed issues surrounding the statistical design and collection of surveillance data for CWD. The second workshop, from which this document arose, followed logically from the first workshop and focused on appropriate methods for analysis, interpretation, and use of CWD surveillance and related epidemiology data. Consequently, the emphasis of this e-book is on modeling approaches to describe and gain insight of the spatial epidemiology of CWD. We designed this e-book for wildlife managers and biologists who are responsible for the surveillance of CWD in their state or agency. We chose spatial methods that are popular or common in the spatial epidemiology literature and evaluated them for their relevance to modeling CWD. Our opinion of the usefulness and relevance of each method was based on the type of field data commonly collected as part of CWD surveillance programs and what we know about CWD biology, ecology, and epidemiology. Specifically, we expected the field data to consist primarily of the infection status of a harvested or culled sample along with its date of collection (not date of infection), location, and demographic status. We evaluated methods in light of the fact that CWD does not appear to spread rapidly through wild populations, relative to more highly contagious viruses, and can be spread directly from animal to animal or indirectly through environmental contamination. We discovered that many of the wellpublished methods were developed for fast-spreading human diseases, such as influenza and measles. While these methods are applicable to fast spreading wildlife diseases, such as foot-and-mouth disease or West Nile virus, many are not likely to work well for CWD. Only limited data exist to evaluate geographic and spatial spread because many locations where we find CWD tend to be locations where samples have just been taken or sample sizes have just become large enough to have a high probability of detecting a low prevalence. Consequently, methods that work well to describe or predict the spread of foot-and-mouth disease throughout England, which occurred within a year, do not work well for describing or predicting CWD spread. We did not exclude methods that we regarded as inappropriate; rather, we included methods that are commonly used for disease epidemiology and then discussed their applicability for modeling the spatial epidemiology of CWD. We hope including inappropriate methods with an explanation of why they are ill-suited for CWD will make it easier to drop them from consideration and explain to others why they were not recommended for spatial modeling of CWD. We organized the three chapters by scale and extent for which each method was developed or best suited. The first chapter covers methods appropriate to multi-jurisdictional or multi-state modeling, which we call “regional” scale. The second chapter covers methods appropriate for within state areas such as wildlife management units or metapopulations, which we call “landscape” scale. The third chapter covers methods appropriate for population or individual-based modeling, which we call “fine” scale. We know this rubric is somewhat artificial because many methods work at multiple scales. We hope, however, that this structure addresses some of the challenges faced by managers that work at local, regional, state, and national scales. Further, the resolution of empirical data often changes with spatial scale, which affects the utility of different modeling approaches. For example, individual-based models work best at modeling spread within populations, while risk analysis is most useful for summarizing data over larger scales such as a region. Because some methods are applicable at several scales, however, we included a graphic at the beginning of each method that indicates the range of scales for which it applies. For example, the graphic to the right indicates that the method is most applicable for regional-scale modeling. There is also a question of resolution as well as scale and extent for each method. CWD surveillance data have been collected over large areas, such as a wildlife management unit or state, but the resolution of the data may be fine scale with GPS locations for many samples. For each method, we described the required resolution of the data and describe the type of data required, as well as what questions the method could answer and how useful the method is, given typical CWD data. For each scale, we presented a focal approach that would be useful for understanding the spatial pattern and epidemiology of CWD, as well as being a useful tool for CWD management. The focal approaches include risk analysis and micromaps for the regional scale, cluster analysis for the landscape scale, and individual based modeling for the fine scale of within population. For each of these methods, we used simulated data and walked through the method step by step to fully illustrate the “how to”, with specifics about what is input and output, as well as what questions the method addresses. We also provided a summary table to, at a glance, describe the scale, questions that can be addressed, and general data required for each method described in this e-book. We hope that this review will be helpful to biologists and managers by increasing the utility of their surveillance data, and ultimately be useful for increasing our understanding of CWD and allowing wildlife biologists and managers to move beyond retroactive fire-fighting to proactive preventative action.

Book↗

Preliminary assessment of channel stability and bed-material transport in the Coquille River basin, southwestern Oregon

This report summarizes a preliminary study of bed-material transport, vertical and lateral channel changes, and existing datasets for the Coquille River basin, which encompasses 2,745 km 2 (square kilometers) of the southwestern Oregon coast. This study, conducted to inform permitting decisions regarding instream gravel mining, revealed that: The 115.4-km-long study area on the South Fork and mainstem Coquille River can be divided into four reaches on the basis of topography and hydrology. In the fluvial (nontidal, or dominated by riverine processes) reaches on the South Fork Coquille River, the channel consists of bedrock and alluvium in the Powers Reach and mostly alluvium in the Broadbent Reach. In both fluvial reaches, the channel alternates between confined and unconfined segments and contains gravel bars. In the tidally affected Myrtle Point and Bandon Reaches, the channel consists of alluvial deposits and contains sparse gravel and sand bars as well as expansive mud flats and tidal marshes near the Coquille River mouth. The 15.4- and 14.6-km-long study areas on the Middle and North Forks of the Coquille River, respectively, were treated as distinct reaches. The channel beds consist of mixed bedrock and alluvium in the Bridge Reach on the Middle Fork Coquille River and alluvium in the Gravelford Reach on the North Fork Coquille River. Both of these reaches contain fewer bars than the Powers and Broadbent Reaches on the South Fork Coquille River and are predominately fluvial. Channel condition, bed-material transport, and the distribution and area of bars have likely been influenced by logging and splash damming, dredging and wood removal for navigation, historical and ongoing instream gravel mining, gold mining, fires, and mass movements. These anthropogenic and natural disturbances likely have varying effects on channel condition and sediment flux throughout the study area and over time. Available data include at least eight sets of aerial and orthophotographs that were taken of the study area from 1939 to 2011 that are available for assessing long-term changes in channel condition, bar area, and vegetation establishment patterns. Additionally, a high-resolution Light Detection And Ranging (LiDAR) survey conducted in 2008 for nearly the entire study area would be useful in future quantitative analyses of channel morphology and bed-material transport. Previous studies found (1) substantial bank erosion in the Broadbent Reach, resulting in banks with near vertical profiles and heights exceeding 7.6 m, (2) erosion of over 40,000 square meters of riparian land from 1939 to 1992, (3) incision along the South Fork Coquille River, and (4) potential for lateral channel migration at several locations along the mainstem and South Fork Coquille River. A review of deposited and mined bed-material estimates derived largely from repeat surveys at instream mining sites on the South Fork Coquille River indicates that bed material transported by the river tends to rebuild mined bar surfaces in most years. Reported annual deposition volumes for 1996–2009 indicate average transport of over 34,700 cubic meters per year (m 3 /yr) of bed material into the South Fork Coquille River study area. The spatial variation in the number and area of gravel bars is controlled by factors such as valley confinement, channel slope, basin geology, and tidal extent. The Powers and Broadbent Reaches of the South Fork Coquille River have the greatest abundance of gravel bars, likely owing to a substantial area of the South Fork Coquille River basin draining the gravel-producing Klamath Mountains geologic province. From 1939 to 2009, the fluvial reaches all had a net loss in bar area, ranging from 24 percent in the Powers Reach to 56 percent in the Bridge Reach. In the Powers and Broadbent Reaches, the declines in active bar area were associated primarily with vegetation establishment on bar surfaces and lateral bar erosion. The reductions in active bar area were attributed to vegetation establishment in the Bridge and Gravelford Reaches as well as some lateral bar erosion in the Bridge Reach. In contrast, the tidal Myrtle Point and Bandon Reaches had a net increase in bar area (28 and 29 percent, respectively) from 1939 to 2009. In the Myrtle Point Reach, these increases in bar area were primarily attributed to lateral channel migration that led to the deposition of bed material at newly formed bars. In the Bandon Reach, bar area increased primarily in the lower 5.4 km of the reach owing possibly to factors such as tide differences between the photographs and sediment deposition. Analyses of multiple channel cross sections along the South Fork Coquille River as well as historical stage-discharge data collected by the U.S. Geological Survey (USGS) at Powers, Oregon, indicate that the bed of the South Fork Coquille River has locally lowered, as much as 1.9 m from 1994 to 2008 for one site in the Broadbent Reach. Stage-discharge data indicate persistent incision at the Powers site since 1939 (with a net incision of about 0.3 m) that has been interrupted by episodic aggradation apparently corresponding with large floods. For the Bridge and Gravelford Reaches on the Middle and North Forks of the Coquille River, channel cross sections indicate a mix of aggradation and incision as well as bank erosion and deposition from 1992 to 2010 and 2000 to 2009, respectively. Cross sections in the tidal reaches indicate local incision of 0.4 m in at one site in the Myrtle Point Reach from 2004 to 2008 and 0.5 m at one site in in the Bandon Reach from 2000 to 2010. On the South Fork Coquille River, the median diameter of surface particles varied from 78.0 mm (millimeters) at China Flat Bar slightly upstream of the study area to 48.8 mm at Seals Bar in the Broadbent Reach. The armoring ratio (or ratio of the median grain sizes of the surface and subsurface layers) for Seals Bar was 3.5, indicating that the river’s transport capacity likely exceeds sediment supply at this site. Most fluvial reaches in the Coquille River study area are likely supply-limited, meaning that the river’s transport capacity exceeds the supply of bed-material, as indicated by the intermittent bedrock outcrops in the Powers and Bridge Reaches and the paucity of bars in the Bridge and Gravelford Reaches. The Broadbent Reach of the South Fork Coquille River may be presently and probably was historically transport-limited, meaning that bed-material transport is primarily a function of local transport capacity. However, the locally coarse bed texture, high armoring ratio measured at Seals Bar, and recent channel incision indicate that sediment supply has likely diminished relative to transport capacity in recent decades. Because of exceedingly low gradients, the tidal Myrtle Point and Bandon Reaches are transport limited. Bed material in these reaches is primarily sand and finer grain-size material, much of which is probably transported as suspended load from upstream reaches. The tidal reaches will be most susceptible to watershed conditions affecting the supply and transport of fine sediment. Compared to the nearby Chetco and Rogue Rivers and Hunter Creek on the southwestern Oregon coast, the Coquille River likely has lower overall transport of gravel bed material. While the conclusion of lower bed-material transport in the Coquille River is tentative in the absence of actual transport measurements or transport capacity calculations, empirical evidence including the much lower area and frequency of bars for most of the Coquille River study area and the head of tide reaching to RKM (river kilometer) 63.2 on the South Fork Coquille River supports this conclusion. More detailed investigations of bed-material transport rates and channel morphology would support assessments of lateral and vertical channel condition and longitudinal trends in bed material. Such assessments would be most practical for the Powers and Broadbent Reaches and relevant to several ongoing management and ecological issues pertaining to sand and gravel transport. The tidal Bandon and Myrtle Point Reaches may also be logical subjects for in-depth analyses of fine sediment deposition and transport (and associated channel and riparian conditions and processes) rather than coarse bed material.

Oregon↗

Connectivity of Mojave Desert tortoise populations—Management implications for maintaining a viable recovery network

Executive Summary The historic distribution of Mojave desert tortoises ( Gopherus agassizii ) was relatively continuous across the range, and the importance of tortoise habitat outside of designated tortoise conservation areas (TCAs) to recovery has long been recognized for its contributions to supporting gene flow between TCAs and to minimizing impacts and edge effects within TCAs. However, connectivity of Mojave desert tortoise populations has become a concern because of recent and proposed development of large tracts of desert tortoise habitat that cross, fragment, and surround designated conservation areas. This paper summarizes the underlying concepts and importance of connectivity for Mojave desert tortoise populations by reviewing current information on connectivity and providing information to managers for maintaining or enhancing desert tortoise population connectivity as they consider future proposals for development and management actions. Maintaining an ecological network for the Mojave desert tortoise, with a system of core habitats (TCAs) connected by linkages, is necessary to support demographically viable populations and long-term gene flow within and between TCAs. There are four points for wildlife and land-management agencies to consider when making decisions that could affect connectivity of Mojave desert tortoise populations (for example, in updating actions in resource management plans or amendments that could help maintain or restore functional connectivity in light of the latest information): Management of all desert tortoise habitat for persistence and connectivity . Desert tortoise populations continue to decline within most TCAs, and it is unlikely that trends are better in populations outside protected areas. Fragmentation exacerbates negative population trends by breaking large continuous populations into smaller isolated populations. Connectivity within large populations can enhance resilience to localized disturbances due to rescue by neighboring individuals. In contrast, smaller fragmented populations are resistant to rescue by their isolation and thus could suffer irreversible declines to extirpation from a variety of threats and stochastic events. Enhanced threat reduction to reverse declines within TCAs and to maintain occupied habitat in the surrounding matrix would help reduce the variability in population growth rates and improve the resilience of protected populations even while implementing efforts to improve connectivity. Each TCA has unique strengths and weaknesses regarding its ability to support minimum sustainable populations based on areal extent and its ability to support population increases based on landscape connection with adjacent populations. Considering how proposed projects (inside or outside of TCAs) affect connectivity and the ability of TCAs to support at least 5,000 adult tortoises (the numerical goal for each TCA) could help managers to maintain the resilience of TCAs to population declines. The same project, in an alternative location, could have very different impacts on local and regional populations. For example, within the habitat matrix surrounding TCAs, narrowly delineated corridors may not allow for natural population dynamics if they do not accommodate overlapping home ranges along most of their widths so that tortoises reside, grow, find mates, and produce offspring that can replace older tortoises. In addition, most habitat outside TCAs may receive more surface disturbance than habitat within TCAs. Therefore, managing the entire remaining matrix of desert tortoise habitat for permeability may be better than delineating fixed corridors. These concepts apply, especially given uncertainty about long-term condition of habitat, within and outside of TCAs under a changing climate. Ultimately, questions such as “ What are the critical linkages that need to be protected ?” could be better framed as “ How can we manage the remaining habitat matrix in ways that sustain ecological processes and habitat suitability for special status species ?” Land-management decisions made in the context of the latter question may be more conducive to maintenance of a functional ecological network. Limitations on landscape-level disturbance across habitat managed for the desert tortoise Clearly delineating habitat linkages and differentiating them from non-delineated areas by the uses that are permitted or prohibited within them by specific management guidelines can help achieve functional connectivity. Such guidelines would be most effective if they considered and accounted for all surface disturbances (for example, temporary disturbances such as fiberoptic lines or off-highway vehicle routes, right-of-ways, utility-scale solar development, urbanization) to the extent possible. A weighted framework that varies with the permanence or severity of the disturbance, and can be additive to quantify cumulative effects, could be useful (Xiong, 2020). For example, minor roads can alter tortoise movements independently of other features (Peaden and others, 2017; Hromada and others, 2020), but if the isolated dirt road is accompanied by a powerline that encourages raven predation (Xiong, 2020), then the two features together may be additive. Ignoring minor or temporary disturbance on the landscape could result in a cumulatively large impact that is not explicitly acknowledged (Goble, 2009); therefore, understanding and quantifying all surface disturbance on a given landscape is prudent. In California, the Bureau of Land Management established 0.1–1.0 percent caps on new surface-disturbance for TCAs and mapped linkages that address the issues described in number 1 of this list. Nevada, Utah, and Arizona currently do not have surface-disturbance limits. Limits comparable to those in the Desert Renewable Energy Conservation Plan (DRECP) would be 0.5 percent within TCAs and 1 percent within the linkages modeled by Averill-Murray and others (2013). Limits in some areas of California within the Desert Renewable Energy Conservation Plan, such as Ivanpah Valley, are more restrictive, at 0.1 percent. Continuity across the state line in Nevada could be achieved with comparable limits in the adjacent portion of Ivanpah Valley, as well as the Greater Trout Canyon Translocation Area and the Stump Springs Regional Augmentation Site. These more restrictive limits would help protect remaining habitat in the major interstate connectivity pathway through Ivanpah Valley and focal areas of population augmentation that provide additional population connectivity along the western flank of the Spring Mountains. In a recent study that analyzed 13 years of desert tortoise monitoring data, nearly all desert tortoise observations were at sites in which 5 percent or less of the surrounding landscape within 1 kilometer was disturbed (Carter and others, 2020a). To help maintain tortoise habitability and permeability across all other non-conservation-designated tortoise habitat, all surface disturbance could be limited to less than 5-percent development per square kilometer because the 5-percent threshold for development is the point at which tortoise occupation drops precipitously (Carter and others, 2020a). However, although individual desert tortoises were observed at development levels up to 5 percent, we do not know the fitness or reproductive characteristics of these individuals. This level of development also may not allow for long-term persistence of healthy populations that are of adequate size needed for demographic or functional connectivity; therefore, a conservative interpretation suggests that, ideally, development could be lower. Lower development levels would be particularly useful in areas within the upper 5th percentile of connectivity values modeled by Gray and others (2019). Reducing ancillary threats in places where connectivity is restricted to narrow strips of habitat, for example, narrow mountain passes or vegetated strips between solar development, could enhance the functionality of these vulnerable linkages. In such areas, maintaining multiple, redundant linkages could further enhance overall connectivity. Minimization of mortality from roads and maximization of passage under roads . Roads pose a significant threat to the long-term persistence of local tortoise populations, and roads of high traffic volume lead to severe population declines, which ultimately fragments populations farther away from the roads. Three points (a.–c.) pertain to reducing direct mortality of tortoises on the many paved roads that cross desert tortoise habitat and to maintaining a minimal level of permeability across these roads: Tortoise-exclusion fencing tied into culverts, underpasses, overpasses, or other passages below roads in desert tortoise habitat, would limit vehicular mortality of tortoises and provide opportunities for movement across the roads. Installation of shade structures on the habitat side of fences installed in areas with narrow population-depletion zones would limit overheating of tortoises that may pace the fence. Passages below highways could be maintained or retrofitted to ensure safe tortoise access, for example, by filling eroded drop-offs or modifying erosion-control features such as rip-rap to make them safer and more passable for tortoises. Wildlife management agencies could work with transportation departments to develop construction standards that are consistent with hydrologic/erosion management goals, while also incorporating a design and materials consistent with tortoise survival and passage and make the standards widely available. The process would be most effective if the status of passages was regularly monitored and built into management plans. Healthy tortoise populations along fenced highways could be supported by ensuring that land inside tortoise-exclusion fences is not so degraded that it leads to degradation of tortoise habitat outside the exclusion areas. For example, severe invasive plant infestations inside a highway exclusion could cause an increase of invasive plants outside the exclusion area and degrade habitat; therefore, invasive plants inside road rights of way could be mown or treated with herbicide to limit their spread into adjacent tortoise habitat and minimize the risk of these plants carrying wildfires into adjacent habitat. Adaptation of management based on new information . Future research will continue to build upon and refine models related to desert tortoise population connectivity and develop new ones. New models could consider landscape levels of development and be constructed such that they share common foundations to support future synthesis efforts. If model development was undertaken in partnership with entities that are responsible for management of desert tortoise habitat, it would facilitate incorporation of current and future modeling results into their land management decisions. There are specific topics that may be clarified with further evaluation: The effects of climate change on desert tortoise habitat, distribution, and population connectivity; The effects of large-scale fires, especially within repeatedly burned habitat, on desert tortoise distribution and population connectivity; The ability of solar energy facilities or similar developments to support tortoise movement and presence by leaving washes intact; leaving native vegetation intact whenever possible, or if not possible, mowing the site, allowing vegetation to re-sprout, and managing weeds; and allowing tortoises to occupy the sites; and The design and frequency of underpasses necessary to maintain functional demographic and genetic connectivity across linear features, like highways.

Arizona, California, Nevada↗

Preliminary assessment of channel stability and bed-material transport in the Rogue River basin, southwestern Oregon

This report summarizes a preliminary assessment of bed-material transport, vertical and lateral channel changes, and existing datasets for the Rogue River basin, which encompasses 13,390 square kilometers (km 2 ) along the southwestern Oregon coast. This study, conducted to inform permitting decisions regarding instream gravel mining, revealed that: The Rogue River in its lowermost 178.5 kilometers (km) alternates between confined and unconfined segments, and is predominately alluvial along its lowermost 44 km. The study area on the mainstem Rogue River can be divided into five reaches based on topography, hydrology, and tidal influence. The largely confined, active channel flows over bedrock and coarse bed material composed chiefly of boulders and cobbles in the Grants Pass (river kilometers [RKM] 178.5–152.8), Merlin (RKM 152.8–132.7), and Galice Reaches (RKM 132.7–43.9). Within these confined reaches, the channel contains few bars and has stable planforms except for locally wider segments such as the Brushy Chutes area in the Merlin Reach. Conversely, the active channel flows over predominately alluvial material and contains nearly continuous gravel bars in the Lobster Creek Reach (RKM 43.9–6.7). The channel in the Tidal Reach (RKM 6.7–0) is also alluvial, but tidally affected and unconfined until RKM 2. The Lobster Creek and Tidal Reaches contain some of the most extensive bar deposits within the Rogue River study area. For the 56.6-km-long segment of the Applegate River included in this study, the river was divided into two reaches based on topography. In the Upper Applegate River Reach (RKM 56.6–41.6), the confined, active channel flows over alluvium and bedrock and has few bars. In the Lower Applegate River Reach (RKM 41.6–0), the active channel alternates between confined and unconfined segments, flows predominantly over alluvium, shifts laterally in unconfined sections, and contains more numerous and larger bars. The 6.5-km segment of the lower Illinois River included in this study was treated as one reach. This stretch of the Illinois River is fully alluvial, with nearly continuous gravel bars flanking the channel. The width of the active channel is confined by the narrow topography of the valley. The primary human activities that have likely influenced channel condition, bed-material transport, and the extent and area of bars are (1) historical gold mining throughout the basin, (2) historical and ongoing gravel mining from instream sites in the Tidal Reach and floodplain sites such as those in the Lower Applegate River Reach, (3) hydropower and flow control structures, (4) forest management and fires throughout the basin, and (5) dredging. These anthropogenic activities likely have varying effects on channel condition and the transport and deposition of sediment throughout the study area and over time. Several vertical (aspect) aerial photographs (including the complete coverages of the study area taken in 1995, 2000, 2005, and 2009 and the partial coverages taken in 1967, 1968, 1969, and 1990) are available for assessing long-term changes in attributes such as channel condition, bar area, and vegetation cover. A Light Detection And Ranging (LiDAR) survey performed in 2007–2008 provides 1-m resolution topographic data for sections of the Grants Pass (RKM 178.5–167.6) and Lobster Creek (RKM 17.8–12 and 10–6.7) Reaches and the entire Tidal Reach. Previous studies provide information for specific locations, including (1) an estimated average annual bed-material load of 76,000 m 3 at the former Savage Rapids Dam site (RKM 173.1, Grants Pass Reach), (2) over 490 m of channel shifting from 1965 to 1991 in the Brushy Chutes area (RKM 142–141, Merlin Reach), (3) active sediment transport and channel processes in the Lobster Creek Reach, (4) lateral channel migration in the Tidal Reach, and (5) up to 1.8 m of bar aggradation from the town of Agness (RKM 45.1) to the Rogue River mouth following the flood in water year 1997. Review of the repeat surveys conducted at the instream gravel-mining sites on Elephant and Wedderburn Bars tentatively indicated that these bars (1) experience some bed-material deposition in most years and more substantial deposition following high flows such as those in water years 1997 and 2006, and (2) are dynamic and subject to local scour and deposition. Results from the specific gage analyses completed for five long-term USGS streamflow-gaging stations showed that only the Grants Pass station on the Rogue River (RKM 164.4, Grants Pass Reach) experienced substantial changes in the stage–discharge relationship across a range of flows from 1938 to 2009. Observed changes indicate channel incision at this site. The Rogue and Applegate Rivers are dynamic and subject to channel shifting, aggradation, and incision, as indicated by channel cross sections surveyed during 2000–2010 on the Rogue River and 1933–2010 on the Applegate River. The elevation of the riverbed changed substantially (defined here as more than a net 0.5 m of incision or aggradation) at three locations on the Rogue River (near RKM 164.5, 139.2, and 1.3) and two on the Applegate River (near RKM 42 and 13.5). Systematic delineation of bar features from vertical photographs taken in 1967–69, 2005, and 2009 indicated that most of the repeat mapping sites had a net loss in bar area over the analysis period, ranging from 22 percent at the Oak Flat site (Illinois River Reach) to 69 percent at the Thompson Creek site (Upper Applegate River Reach). Bar area remained stable at the Williams Creek site (Lower Applegate River Reach), but increased 11 percent at the Elephant Rock site (Tidal Reach). The declines in bar area were associated primarily with the establishment of vegetation on upper bar surfaces lacking obvious vegetation in the 1960s. Some of the apparent changes in bar area may also owe to some differences in streamflow and tide levels between the vertical photographs. On the mainstem Rogue River, the median diameter of surface particles varied from 21 mm at the Wedderburn Bar in the Tidal Reach to more than 100 millimeters (mm) at some of the coarsest bars in the Galice Reach. Low armoring ratios tentatively indicated that sediment supply likely exceeds transport capacity at Orchard (Lobster Creek Reach) and Wedderburn (Tidal Reach) Bars. Conversely, relatively higher armoring ratios indicated that transport capacity likely is in balance with sediment supply at Roberston Bridge Bar (Merlin Reach) and exceeds sediment supply at Rogue River City (Grants Pass Reach), Solitude Riffle (Galice Reach), and Hooks Gulch (Galice Reach) Bars. Limited particle data were collected in the study areas on the Applegate and Illinois Rivers. Particle size measurements and armoring ratios tentatively show that sediment supply likely exceeds transport capacity at Bakery Bar in the Lower Applegate Reach. Also, the bed material exiting the Applegate River is likely finer than the bed material in the Rogue River, whereas bed material exiting the Illinois River is likely coarser than the bed material in the Rogue River. Together, these observations and findings indicate that (1) the size, area, and overall position of bars in the Rogue River study area are determined largely by valley physiography, such that unconfined alluvial sections have large channel-flanking bars, whereas confined sections have fewer and smaller bars, (2) segments within the Grants Pass, Merlin, Tidal, Upper Applegate River, and Lower Applegate River Reaches are prone to vertical and/or lateral channel adjustments, and (3) the balance between transport capacity and sediment supply varies throughout the study area. High winter flows and the steep, confined character of much of the Rogue River within the study area result in a river corridor with a high capacity to transport bed material. In the Grants Pass and Galice Reaches, the extensive in-channel bedrock as well as the sparse number and coarse texture of bars indicate that these reaches are likely supply-limited, meaning that the river’s transport capacity exceeds the supply of bed material. In contrast, the Lobster Creek and Tidal Reaches and perhaps portions of the Merlin Reach receive bed-material inputs that more closely balance or even exceed the river’s transport capacity. The lowermost reaches on the Illinois and Applegate Rivers are fully alluvial segments that are likely transport limited, meaning sediment supply likely exceeds the river's transport capacity. However, the steeper Upper Applegate River Reach is likely supply-limited as indicated by the sparse number and area of bars mapped in this reach and the intermittent bedrock outcrops in the channel. The sediment loads derived from these large tributaries draining the Klamath Mountains are probably important contributions to the overall transport of bed material in the Rogue River basin. Compared to the slightly smaller Umpqua River basin (drainage area 12,103 km 2 ) to the north, the Rogue River (13,390 km 2 ) likely transports more bed material. Although this conclusion of greater bed-material transport in the Rogue River is tentative in the absence of either actual transport measurements or transport capacity calculations, empirical evidence, including the much greater area and frequency of bars along most of the Rogue River as well as the much shorter tidal reach on the Rogue River (6.7 km) compared to the Umpqua River (40 km) supports this inference. More detailed investigations of bed-material transport rates and channel morphology would support assessments of channel condition, longitudinal trends in particle size, the relation between sediment supply and transport capacity, and the potential causes of bar area loss (such as vegetation establishment and potential changes in peak flow patterns). The reaches most practical for such assessments and relevant to several management and ecological issues are (1) the lower Rogue River basin, including the Lobster Creek and Tidal Reaches of the Rogue River as well as the Illinois River Reach and (2) the Lower Applegate River Reach.

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