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Evolution and distribution of the coregonids

Increased knowledge of the morphological and physiological plasticity of the coregonids has minimized the significance of conflicting morphological data that have retarded the development of an evolutionary theory. Of the four phyletic lines recognized through worldwide studies, Coregonus originated in the lake and stream area of northwest Eurasia, Stenodus and Prosopium evolved in the rivers of Siberia and northwest America respectively, and Leucichthys became differentiated in the lake-studded area of northeast America. Isolation and range extension were facilitated by events of the Pleistocene epoch. Inter-continental exchange most likely took place in the Bering Strait region. Representatives of each group in its area of origin are highly variable. Range extensions of each group are characterized by lesser morphological variability and at the extremes only one or two relatively stable species remain.

Journal of the Fisheries Research Board of Canada↗

The relationship between annual survival rate and migration distance in mallards: An examination of the time-allocation hypothesis for the evolution of migration

Predictions of the time-allocation hypothesis were tested with several a posteriori analyses of banding data for the mallard (Anas platyrhynchos). The time-allocation hypothesis states that the critical difference between resident and migrant birds is their allocation of time to reproduction on the breeding grounds and survival on the nonbreeding grounds. Residents have higher reproduction and migrants have higher survival. Survival and recovery rates were estimated by standard band-recovery methods for banding reference areas in the central United States and central Canada. A production-rate index was computed for each reference area with data from the U.S. Fish and Wildlife Service May Breeding Population Survey and July Production Survey. An analysis of covariance was used to test for the effects of migration distance and time period (decade) on survival, recovery, and production rates. Differences in migration chronology were tested by comparing direct-recovery distributions for different populations during the fall migration. Differences in winter locations were tested by comparing distributions of direct recoveries reported during December and January. A strong positive relationship was found between survival rate, and migration distance for 3 of the 4 age and sex classes. A weak negative relationship was found between recovery rate and migration distance. No relationship was found between production rate and migration distance. During the fall migration, birds from the northern breeding populations were located north of birds from the southern breeding populations. No pattern could be found in the relative locations of breeding and wintering areas. Although our finding that survival rate increased with migration distance was consistent with the time-allocation hypothesis, our results on migration chronology and location of wintering areas were not consistent with the mechanism underlying the time-allocation hypothesis. Neither this analysis nor other recent studies of life-history characteristics of migratory and resident birds supported the timeallocation hypothesis.

Canadian Journal of Zoology↗

Population structure and the evolution of sexual size dimorphism and sex ratios in an insular population of Florida box turtles ( Terrapene carolina bauri )

Hypotheses in the chelonian literature suggest that in species with sexual size dimorphism, the smaller sex will mature at a smaller size and a younger age than the larger sex, sex ratios should be biased in favor of the earlier maturing sex, and deviations from a 1:1 sex ratio result from maturation of the smaller sex at a younger age. I tested these hypotheses using data collected from 1991 to 1995 on an insular (Egmont Key) population of Florida box turtles, Terrapene carolina bauri. Contrary to predictions, the earlier maturing sex (males) grew to larger sizes than the late maturing sex. Males were significantly larger than females in mean carapace length but not mean body mass. Sex ratios were not balanced, favoring the earlier maturing sex (1.6 males:1 female), but the sex-ratio imbalance did not result from faster maturation of the smaller sex. The imbalance in the sex ratio in Egmont Key's box turtles is not the result of sampling biases; it may result from nest placement. Size-class structure and sex ratios can provide valuable insights into the status and trends of populations of long-lived turtles.

Canadian Journal of Zoology↗

Barriers on the brink? The complex intertwined roles of geologic framework, sediment availability and sea-level rise in island evolution

Sensitivity experiments in the North Carolina Outer Banks (OBX) have previously revealed that substrate sand proportion, followed by substrate slope, sea-level rise rate and sediment-loss rate are the most important factors in determining how barrier islands respond to sea-level rise. High sediment-loss rates and low substrate sand proportions cause barriers to be smaller and more deeply incised. Thus, as sea level rise rates increase, more deeply incised barriers do not need to migrate as far landward as larger, less-incised barriers to liberate sand from the shoreface. However, if the combination of sand losses and substrate sand proportions requires a barrier to migrate landward faster than the shoreface can erode to replenish losses, a barrier will change state and begin to disintegrate. Because the substrate of the OBXis sand-rich, these barriers are likely to persist in the near-term. In contrast, model simulations for the Chandeleur Islands, Louisiana suggest sediment loss rates are too high and/or substrate sand proportions are too low to be matched by liberation of shoreface sand. These simulations further suggest that a state change, from a landward-migrating barrier system to a subaqueous shoal complex, is either already underway or imminent.

Conference Paper↗

Chapter 8: The evolution of Santorini

There now follows a synthesis of the main results of this study. We interpret the observations in terms of the interplay between magmatic, tectonic and volcanic processes and place these processes within a global context. Santorini has displayed a wide variety of igneous and volcanic phenomena over a period of at least 650 000 years of almost continuous and focused volcanism. Perhaps the most intriguing issues are the causes of volcanic and magmatic cyclicity, the secular variations in magma composition and the way in which magmatic processes have influenced the style of volcanism.

Santorini↗

Ordovician K-bentonites in the Argentine Precordillera: Relations to Gondwana margin evolution

Ordovician K-bentonites have now been recorded from >20 localities in the vicinity of the Argentine Precordillera. Most occur in the eastern thrust belts, in the San Juan Limestone and the overlying the Gualcamayo Formation, but a few ash beds are known also from the central thrust belts. The oldest occur in the middle Arenig I. victoriae lunatus graptolite ( Oe. evae conodont) Zone, and the youngest in the middle Llanvirn P. elegans ( P. suecicus ) Zone. Mineralogical characteristics, typical of other Ordovician K-bentonites, include a matrix of illite/smectite mixed-layer clay and a typical felsic volcanic phenocryst assemblage: biotite, beta-form quartz, alkali and plagioclase feldspar, apatite, and zircon, with lesser amounts of hornblende, clinopyroxene, titanite and Fe-Ti oxides. The proportions of the mineral phases and variations in their crystal chemistry are commonly unique to individual (or small groups of) K-bentonite beds. Glass melt inclusions preserved in quartz are rhyolitic in composition. The sequence is unique in its abundance of K-bentonite beds, but a close association between the Precordillera and other Ordovician sedimentary basins cannot be established. The ash distribution is most consistent with palaeogeographical reconstructions in which early Ordovician drifting of the Precordillera occurred in proximity to one or more volcanic arcs, and with eventual collision along the Andean margin of Gondwana during the mid-Ordovician Ocloyic event of the Famatinian orogeny. The Puna-Famatina terrane northeast of the Precordillera might have served as the source of the K-bentonite ashes, possibly in concert with active arc magmatism on the Gondwana plate itself.

Geological Society Special Publication↗

Tectonic evolution of Gulf of Anadyr and formation of Anadyr and Navarin basins

New seismic reflectionand refraction data reveal that Anadyr basin is separated from Navarin basin by Anadyr ridge, a southeast-northwest-trending bedrock high that is characterized by high-amplitude, short-wavelength magnetic anomalies. Anadyr ridge may be an offshore extension of the melange belt underlying the Koryak Range. Sonobuoy refraction data indicate that the velocity profile of strata in East Anadyr trough is similar to that in Navarin basin. Structurally, the basins are different.

Siberia↗

Molecular and isotopic gas composition of the Devonian Berea Sandstone and implications for gas evolution, eastern Kentucky

Since 2011, the Devonian Berea Sandstone in northeastern Kentucky has produced oil where thermal maturity studies indicate that likely source rocks, namely, the Devonian Ohio Shale and Mississippian Sunbury Shale, are thermally immature. Downdip, where source rocks are mature for oil, the Berea Sandstone and Ohio Shale primarily produce gas. To investigate this thermal maturity discordancy, the molecular and isotopic composition of gases from the Berea Sandstone (oil associated) and Ohio Shale (nonassociated) were analyzed to understand the gas generation and migration history. Collected along a northwest-southeast transect in eastern Kentucky, samples range from 1079 to 4602 ft, respectively. All are wet gases with a thermogenic origin (δ 13 C-CH 4 = −53.5‰ to −46.1‰). This is mostly consistent with a reevaluation of thermal maturity in a companion study that expands the area mature for oil. Gas migration is required, however, in updip parts of the Berea play where vitrinite reflectance ( VR o ) values are less than 0.6%. Southeast regional dip exerts a first-order influence on thermal maturity as gases become drier and isotopically heavier downdip. Correlation of δ 13 C values for heavier hydrocarbon gases in a natural gas plot with VR o contour spacing demonstrates the influence of nearby source rock thermal maturity on gas composition. Downdip, migration of oil and the attendant increase in permeability relative to gas may account for the dominance of gas production in the Ohio Shale. Migration along with basin uplift after the Alleghany orogeny may have contributed to development of a gas phase that exsolved from oil.

Kentucky↗

Porosity evolution of upper Miocene reefs, Almeria Province, southern Spain

Sea cliffs 40 km east of Almeria, southeastern Spain, expose upper Miocene reefs and patch reefs of the Plomo formation. These reefs are formed of scleractinian corals, calcareous algae, and mollusks. The reef cores are as much as 65 m thick and several hundred meters wide. Fore-reef talus beds extend 1,300 m across and are 40 m thick. The reefs and reef breccias are composed of calcitic dolomite. They lie on volcanic rocks that have a K-Ar date of 11.5 m.y. and in turn are overlain by the upper Miocene Vicar Formation. In the reef cores and fore-reef breccia beds, porosity is both primary and postdepositional. Primary porosity is of three types: (a) boring clam holes in the scleractinian coral heads, cemented reef rocks, and breccias; (b) intraparticle porosity within the corals, Halimeda plates, and vermetid worm tubes; and (c) interparticle porosity between bioclastic fragments and in the reef breccia. Postdepositional moldic porosity was formed by the solution of aragonitic material such as molluscan and coral fragments. The Plomo reef carbonate rocks have high porosity and permeability, and retain a great amount of depositional porosity. Pores range in size from a few micrometers to 30 cm. The extensive intercrystalline porosity and high permeability resulted from dolomitization of micritic matrix. Dolomite rhombs are between 10 and 30 µ across. More moldic porosity was formed by the dissolution of the calcite bioclasts. Some porosity reduction has occurred by incomplete and partial sparry calcite infilling of interparticular, moldic, and intercrystalline voids. The high porosity and permeability of these reefs make them important targets for petroleum exploration in the western Mediterranean off southern Spain. In these offshore areas in the subsurface the volcanic ridge and the Plomo reef complex are locally onlapped or overlapped by 350 m or more of Miocene(?) and Pliocene fine-grained sedimentary rocks. The possibility exists that the buried Plomo reef deposits may form traps for oil and gas in the offshore areas southwest of the type locality. Stratigraphic traps also may occur where the Neogene sequence above the Plomo reef complex onlaps the volcanic ridge.

Almeria Province↗

Northwest margin of California continental borderland: Marine geology and tectonic evolution

The northwest margin of the California continental borderland consists mainly of two northwest-trending pre-Neogene lithologic belts blanketed by Miocene and younger strata. These belts, which are lithologically and structurally correlated with the Franciscan Complex and Great Valley sequence of northern California, are interpreted to represent facies corresponding to the subduction complex and forearc-basin deposits of a late Mesozoic and Paleogene continental-margin arc-trench system. The outer belt, which forms the acoustic basement underlying the continental slope and Patton Ridge, is characterized by: (1) moderately high compressional velocities (5.1 to 6.2 km/sec), (2) discordant and discontinuous seismic reflectors, and (3) nonfossiliferous and diverse rock types t at range in metamorphic grade from zeolite-bearing arkosic sandstone to blueschist. Landward, the inner Great Valley sequence belt underlies Santa Rosa-Cortes Ridge and Santa Cruz and San Nicolas Basins; in contrast to the outer belt, this belt is characterized by: (1) intermediate compressional velocities (4.5 to 4.6 km/sec), (2) concordant and relatively continuous reflectors, and (3) a thick turbidite sequence of relatively undeformed and unmetamorphosed fossiliferous Cretaceous and Paleogene strata. The inner belt overlies a 6.6-km/sec layer that is probably oceanic crust, a relation consistent with the hypothesis that trapped old oceanic crust underlies the forearc basins along parts of some convergent margins. Well-developed wrench-fault structures in overlying Miocene strata record a transition from Paleogene subduction to Neogene transform faulting. Related events include widespread andesitic to dacitic volcanism and concurrent uplift and erosion of Miocene and older rocks. The proximity of these Miocene silicic volcanic rocks to the paleotrench (< 20 km) is best explained as resulting from passage of the southward-migrating Rivera ridge-fault-trench triple junction along the margin. On the basis of the age of the adjacent sea floor, passage of this triple junction occurred about 17 m.y. ago, a date that corresponds closely to the inception of volcanism and uplift in the northwest margin of the borderland. A widespread unconformity that separates highly deformed lower and lower middle iocene strata from relatively undeformed younger strata is assumed to mark the subsequent onset of wrench faulting. The timing of volcanism and uplift, and of the slightly younger wrenching event, closely matches the plate-tectonic model of Atwater and Molnar if the northwest margin of the borderland is restored to an earlier tectonic setting off Baja California, approximately 260 km southeast of its present position.

California↗

Structure and evolution of Bering Sea shelf south of St. Lawrence Island

The virtually featureless Beringian shelf south of St. Lawrence Island is underlain structurally by at least 14 basins. Encompassing a total area of more than 300,000 sq km, most of the basins are either elongate structural sags, grabens, or half (asymmetric) grabens beneath the outer shelf. The regional trend of these basins is northwest, parallel with that of the continental margin. Two of the basins, St. George and Navarin, contain 7 to 10 km of Upper Cretaceous(?) and Cenozoic sedimentary strata. A major divergence in dip of beds in the upper half of the sedimentary section may reflect an abrupt shelf-wide change in the rate of sedimentation and/or subsidence, probably during the Miocene. The outer sub-shelf basement grabens and adjacent ridges (horsts) are bounded by high-angle normal faults that exhibit growth-type structure. St. Matthew basin, an elongate, southwest-trending feature of the inner shelf, lies along the offshore expression of the Kaltag fault of western Alaska. The Kaltag fault, like the Denali fault in southwestern Alaska, does not extend to the outer Bering Sea shelf but ends or turns parallel with the margin within the inner shelf. The inner shelf is underlain by a broad basement high, Nunivak arch, the seaward half of which is characterized by an arcuate belt of high-frequency and high-amplitude magnetic anomalies. This zone of intense magnetic anomalies along the shelf is probably the signature of a Mesozoic magmatic arc that extends from southwestern Alaska to eastern Siberia and consists of Jurassic to Cretaceous plutonic and volcanic rocks. We speculate that this magmatic arc resulted from oblique convergence and subduction in the Mesozoic between the Kula(?) and North American plates along the eastern Beringian margin. Folding and uplift in the area of the present outer shelf occurred contemporaneously with magmatism along the inner shelf. Plate convergence apparently ceased by the end of the Mesozoic or t e beginning of the Cenozoic. Subsequently, the foldbelt underlying the outer shelf was eroded extensively and rifted extensionally to form large, deep basins. On the average, the shelf has subsided more than 1.5 km. Subsidence and sediment burial of the eroded orogen formed the modern Beringian shelf.

AAPG Bulletin↗

Subsidence, crustal structure, and thermal evolution of Georges Bank basin

A geophysical study of Georges Bank basin defines a deep crustal structure that is interpreted in terms of the basin's tectonic and thermal history. Gravity models along three basin cross sections delineate two zones of crustal thinning at the basement hinge zone and oceanic crustal margins. These two zones bound rift-stage crust (about 25 km thick) which underlies the central portion of the basin. Subsidence analysis of the basin, using data from multichannel seismic reflection lines and two COST wells, suggests a rifting and (uniform) extensional origin. Two-dimensional finite difference modeling of the basin defines a crustal structure that concurs with the gravity and subsidence studies. The resulting isotherms show no major changes in the thermal structure since the ate Jurassic. In some areas of the basin, temperatures sufficient for oil generation are determined from maturation studies of Jurassic sediments. Hydrocarbon generation is questionable, however, because of the probable lack of proper and sufficient kerogen in the Jurassic deposits.

American Association of Petroleum Geologists Bulle↗

Thickness distributions and evolution of growth mechanisms of NH4-illite from the fossil hydrothermal system of Harghita Bai, Eastern Carpathians, Romania

The crystal growth of NH 4 -illite (NH 4 -I) from the hydrothermal system of Harghita Bãi (Eastern Carpathians) was deduced from the shapes of crystal thickness distributions (CTDs). The 4-illite-smectite (I-S) interstratified structures (R1, R2, and R3-type ordering) with a variable smectite-layer content. The NH 4 -I-S (40–5% S) structures were identified underground in a hydrothermal breccia structure, whereas the K-I/NH 4 -I mixtures were found at the deepest level sampled (−110 m). The percentage of smectite interlayers generally decreases with increasing depth in the deposit. This decrease in smectite content is related to the increase in degree of fracturing in the breccia structure and corresponds to a general increase in mean illite crystal thickness. In order to determine the thickness distributions of NH 4 -I crystals (fundamental illite particles) which make up the NH 4 -I-S interstratified structures and the NH 4, -I/K-I mixtures, 27 samples were saturated with Li + and aqueous solutions of PVP-10 to remove swelling and then were analyzed by X-ray diffraction. The profiles for the mean crystallite thickness (T mean ) and crystallite thickness distribution (CTD) of NH 4 -I crystallites were determined by the Bertaut-Warren-Averbach method using the MudMaster computer code. The T mean of NH 4 -I from NH 4 -I-S samples ranges from 3.4 to 7.8 nm. The T mean measured for the NH 4 -I/K-I mixture phase ranges from 7.8 nm to 11.7 nm (NH 4 -I) and from 12.1 to 24.7 nm (K-I). The CTD shapes of NH 4 -I fundamental particles are asymptotic and lognormal, whereas illites from NH 4 -I/K-I mixtures have bimodal shapes related to the presence of two lognormal-like CTDs corresponding to NH 4 -I and K-I. The crystal-growth mechanism for NH 4 -I samples was simulated using the Galoper code. Reaction pathways for NH 4 -I crystal nucleation and growth could be determined for each sample by plotting their CTD parameters on an α–β 2 diagram constructed using Galoper . This analysis shows that NH 4 -I crystals underwent simultaneous nucleation and growth, followed by surface-controlled growth without simultaneous nucleation.

Harghita Bai↗

Assessing the geologic and climatic forcing of biodiversity and evolution surrounding the Gulf of California

For almost a century the Baja California peninsula (Peninsula), Gulf of California (Gulf), and broader Sonoran Desert region (figure 1) have drawn geologists and biologists alike to study its unique physical and evolutionary processes (e.g., Wittich 1920; Darton 1921; Nelson 1921; Johnston 1924; Beal 1948; Durham and Allison 1960). The challenge remains to untangle the long, intricate, and at times enigmatic geological and climatological histories that have shaped the high levels of endemism and biodiversity observed in the region today (Van Devender 1990; Grismer 2000; Riddle et al. 2000).

Gulf of California↗

Landsat-7 long-term acquisition plan radiometry - evolution over time

The Landsat-7 Enhanced Thematic Mapper Plus instrument has two selectable gains for each spectral band. In the acquisition plan, the gains were initially set to maximize the entropy in each scene. One unintended consequence of this strategy was that, at times, dense vegetation saturated band 4 and deserts saturated all bands. A revised strategy, based on a land-cover classification and sun angle thresholds, reduced saturation, but resulted in gain changes occurring within the same scene on multiple overpasses. As the gain changes cause some loss of data and difficulties for some ground processing systems, a procedure was devised to shift the gain changes to the nearest predicted cloudy scenes. The results are still not totally satisfactory as gain changes still impact some scenes and saturation still occurs, particularly in ephemerally snow-covered regions. A primary conclusion of our experience with variable gain on Landsat-7 is that such an approach should not be employed on future global monitoring missions.

Photogrammetric Engineering and Remote Sensing↗

Comparative genomic analyses and a novel linkage map for cisco (Coregonus artedi) provides insight into chromosomal evolution and rediploidization across salmonids

Whole-genome duplication (WGD) is hypothesized to be an important evolutionary mechanism that can facilitate adaptation and speciation. Genomes that exist in states of both diploidy and residual tetraploidy are of particular interest, as mechanisms that maintain the ploidy mosaic after WGD may provide important insights into evolutionary processes. The Salmonidae family exhibits residual tetraploidy, and this, combined with the evolutionary diversity formed after an ancestral autotetraploidization event, makes this group a useful study system. In this study, we generate a novel linkage map for cisco ( Coregonus artedi ), an economically and culturally important fish in North America and a member of the subfamily Coregoninae, which previously lacked a high-density haploid linkage map. We also conduct comparative genomic analyses to refine our understanding of chromosomal fusion/fission history across salmonids. To facilitate this comparative approach, we use the naming strategy of protokaryotype identifiers (PKs) to associate duplicated chromosomes to their putative ancestral state. The female linkage map for cisco contains 20,292 loci, 3,225 of which are likely within residually tetraploid regions. Comparative genomic analyses revealed that patterns of residual tetrasomy are generally conserved across species, although interspecific variation persists. To determine the broad-scale retention of residual tetrasomy across the salmonids, we analyze sequence similarity of currently available genomes and find evidence of residual tetrasomy in seven of the eight chromosomes that have been previously hypothesized to show this pattern. This interspecific variation in extent of rediploidization may have important implications for understanding salmonid evolutionary histories and informing future conservation efforts.

G3 Genes, Genomes, Genetics↗

Collaborative approaches to the evolution of migration and the development of science-based conservation in shorebirds

Shorebirds are among the most highly migratory creatures on earth. Both the study of their ecology and ongoing efforts to conserve their populations must reflect this central aspect of their biology. Many species of shorebirds use migration and staging sites scattered throughout the hemisphere to complete their annual migrations between breeding areas and nonbreeding habitats (Morrison 1984). The vast distances between habitats they use pose significant challenges for studying their migration ecology. At the same time, the large number of political boundaries shorebirds cross during their epic migrations create parallel challenges for organizations working on their management and conservation. Nebel et al. (2002) represent a collaborative effort to understand the conservation implications of Western Sandpiper ( Calidris mauri ) migration ecology on a scale worthy of this highly migratory species. The data sets involved in the analysis come from four U.S. states, two Canadian provinces, and a total of five nations. Only by collaborating on this historic scale were the authors able to assemble the information necessary to understand important aspects of the migration ecology of this species, and the implications for conservation of the patterns they discovered. Collaborative approaches to shorebird migration ecology developed slowly over several decades. The same period also saw the creation of large-scale efforts to monitor and conserve shorebirds. This overview first traces the history of the study of migration ecology of shorebirds during that fertile period, and then describes the monitoring and protection efforts that have been developed in an attempt to address the enormous issues of scale posed by shorebird migration ecology and conservation.

The Auk↗

Phylogenetics, systematics, paleoecology, and evolution of the trilobite genera Paladin and Kaskia from the United States

Late Mississippian and earliest Pennsylvanian trilobite faunas of North America are dominated by the Paladin and Kaskia clades. Phylogenetic analysis of middle Carboniferous species of these clades demonstrates the close ancestral relationship between these groups. The Kaskia clade consists of eight species: K. chesterensis Weller, 1936, K. osagensis (Cisne, 1967), K. longispina (Strong, 1872), K. wilsoni (Walter, 1924), K. genevievensis (Walter, 1924), K. rosei (Cisne, 1967), K. gersnai n. sp., and K. rollinsi n. sp. Kaskia Weller, 1936 ranges from late Osagean to middle Chesterian (early Visean-early Serpukhovian) and is restricted to cyclothemic shelf and nearshore deposits. Species of Kaskia appear to have evolved in areas of shallow water and high environmental stress. The Paladin clade consists of 12 species including previously named species P. morrowensis (Mather, 1915), P. girtyianus Hahn and Hahn, 1970, P. rarus Whittington, 1954, P. helmsensis Whittington, 1954, and P. moorei (Branson, 1937). New species belonging to this clade are Paladin moorefieldensis n. sp., P. pleisiomorphus n. sp., P. imoensis n. sp., P. mangeri n. sp., and P. wapanukaensis n. sp. This group ranges from the early Chesterian to early Morrowan (late Viseanlate Bashkirian). Species of Paladin appear to be confined to outer shelf shelf-edge and off-shelf facies where presumably deeper water environments existed. This is manifested in their paleogeographic distribution, which is paleoenvironmentally controlled. Copyright ?? 2008, The Paleontological Society.

Journal of Paleontology↗