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At least 631 records · Page 35Linked to original sources
USGS2018

Riparian defoliation by the invasive green alder sawfly influences terrestrial prey subsidies to salmon streams

Invasive species in riparian forests are unique as their effects can transcend ecosystem boundaries via stream‐riparian linkages. The green alder sawfly ( Monsoma pulveratum ) is an invasive wasp whose larvae are defoliating riparian thin‐leaf alder ( Alnus tenuifolia ) stands across southcentral Alaska. To test the hypothesis that riparian defoliation by this invasive sawfly negatively affects the flow of terrestrial prey resources to stream fishes, we sampled terrestrial invertebrates on riparian alder foliage, their subsidies to streams and their consumption by juvenile coho salmon ( Oncorhynchus kisutch ). Invasive sawflies altered the composition of terrestrial invertebrates on riparian alder foliage and as terrestrial prey subsidies to streams. Community analyses supported these findings revealing that invasive sawflies shifted the community structure of terrestrial invertebrates between seasons and levels of energy flow (riparian foliage, streams and fish). Invasive sawfly biomass peaked mid‐summer, altering the timing and magnitude of terrestrial prey subsidies to streams. Contrary to our hypothesis, invasive sawflies had no effect on the biomass of native taxa on riparian alder foliage, as terrestrial prey subsidies, or in juvenile coho salmon diets. Juvenile coho salmon consumed invasive sawflies when most abundant, but relied more on other prey types selecting against sawflies relative to their availability. Although we did not find effects of invasive sawflies extending to juvenile coho salmon in this study, these results could change as the distribution of invasive sawflies expands or as defoliation intensifies. Nevertheless, riparian defoliation by these invasive sawflies is likely having other ecological effects that merits further investigation.

Alaska↗
USGS2019

Managing effects of drought and other water resource challenges in Alaska and the Pacific Northwest

This is a Cooperator Report. As such, there is no specific abstract. The physical, ecological, and social environments of Alaska and the Pacific Northwest (PNW) region of the United States are extremely diverse. Alaska ranges from the Arctic Ocean and the very cold, dry environments of the North Slope to the cool and very rainy coastal North Pacific region of Southeast Alaska. Most precipitation falls as snow at higher elevations. In Arctic Alaska, average annual temperature is 14.6 F, and average annual precipitation is 11 inches. By contrast, in Southeast Alaska, average annual temperature is 35.8 F, and annual average precipitation is 143 inches. The PNW, defined here as Idaho, Oregon, and Washington, ranges from the Pacific Coast (annual precipitation of 200 inches) to interior semi-arid regions (annual precipitation of 8 inches). Precipitation patterns in the PNW are strongly governed by orographic phenomena, with high, persistent snowpack in the higher mountains (e.g., record annual snowfall of 1,130 in at Mount Baker, Washington in 1999-2000). Ecosystems in the PNW include productive temperate coniferous forests near the Pacific coast and along the (wet) west slope of the Cascade Range, less productive mixed-conifer forest along the (dry) east slope of the Cascades and in interior mountain ranges, and sagebrush-steppe and shrublands at lower elevations in much of the interior and mountain valleys. Large rivers and thousands of smaller tributaries form an extensive network of riparian, wetland, and estuarine systems that provide both critical hydrologic function and biological diversity at broad and fine spatial scales. Although Alaska and the Pacific Northwest differ in important physical, ecological, and social features, the importance of natural resources is evident in both regions. Water is important for wildlife and people. Water provides critical habitat for salmon, which are culturally and economically valuable species. Timber production has declined in recent decades. Recreation has emerged as a major revenue source.

Alaska, Oregon, Washington↗
USGS2003

Planning for population viability on Northern Great Plains national grasslands

Broad-scale information in concert with conservation of individual species must be used to develop conservation priorities and a more integrated ecosystem protection strategy. In 1999 the United States Forest Service initiated an approach for the 1.2× 10 6 ha of national grasslands in the Northern Great Plains to fulfill the requirement to maintain viable populations of all native and desirable introduced vertebrate and plant species. The challenge was threefold: 1) develop basic building blocks in the conservation planning approach, 2) apply the approach to national grasslands, and 3) overcome differences that may exist in agency-specific legal and policy requirements. Key assessment components in the approach included a bioregional assessment, coarse-filter analysis, and fine-filter analysis aimed at species considered at-risk. A science team of agency, conservation organization, and university personnel was established to develop the guidelines and standards and other formal procedures for implementation of conservation strategies. Conservation strategies included coarse-filter recommendations to restore the tallgrass, mixed, and shortgrass prairies to conditions that approximate historical ecological processes and landscape patterns, and fine-filter recommendations to address viability needs of individual and multiple species of native animals and plants. Results include a cost-effective approach to conservation planning and recommendations for addressing population viability and biodiversity concerns on national grasslands in the Northern Great Plains.

Wildlife Society Bulletin↗
USGS2013

Natural ecosystems

Natural Ecosystems analyzes the association of observed changes in climate with changes in the geographic distributions and phenology (the timing of blossoms or migrations of birds) for Southwestern ecosystems and their species, portraying ecosystem disturbances—such as wildfires and outbreaks of forest pathogens—and carbon storage and release, in relation to climate change.

Book chapter↗
USGS2019

Fire, forests and city water supply

Forest landscapes generate 57 percent of runoff worldwide and supply water to more than 4 billion people (Millennium Ecosystem Assessment, 2005). As the world population continues to increase, there is a strong need to understand how forest processes link together in a cascade to provide people with water services like hydropower, aquaculture, drinking water and flood protection (Carvalho-Santos, Honrado and Hein, 2014). Wildfire is a major disturbance affecting forested watersheds and the water they provide (Box 1) (Paton et al., 2015). Several regions have experienced shifts in wildfires from natural ignition sources (primarily lightning) to ignitions dominated by human activities, especially in areas where populations are increasing (Moritz et al., 2014; Balch et al., 2017). Occasional wildfire is essential for the health and functioning of fire-adapted ecosystems through its effects on nutrient cycling, plant diversity and succession, and pest regulation (Pausas and Keeley, 2019). It also reduces the risk of subsequent wildfires until a forest has accumulated sufficient fuels and conditions are conducive for another fire. Extreme and hazardous wildfires, on the other hand, can cause erosion, gullying, soil loss and flooding – and, in severe cases, even debris flows and flash floods – by removing the protective functions of forests on hillsides (Ebel and Moody, 2017). Extreme wildfires have become more common after decades of fire suppression, allowing forests to become much denser with vegetation and causing more fuels to build up over time. Combined with increasing summer drought, this can have impacts on water yield and the ability of upstream forests to deliver high-quality water because forest vegetation uses less water immediately after fire and, in environments influenced by snow, more snow can accumulate in forest clearings (Kinoshita and Hogue, 2015; Hallema et al., 2019). Therefore, accounting for wildfire impacts on forests in water planning has become a priority for the nexus of fire, water and society or, in other words, the connection between fire risk and water security (Figure 1) (Martin, 2016). In this article, we discuss managed forest landscapes as nature-based solutions for water and explore how fire affects the provision of water-related services.

Unasylva↗
USGS2009

Identifying baldcypress-water tupelo regeneration classes in forested wetlands of the Atchafalaya Basin, Louisiana

Baldcypress-water tupelo (cypress-tupelo) swamps are critically important coastal forested wetlands found throughout the southeastern U.S. The long-term survival and sustainability of these swamp forests is unknown due to large-scale changes in hydrologic regimes that prevent natural regeneration following logging or mortality. We used NWI wetland maps and remotely sensed hydrologic data to map cypress-tupelo communities, surface water, and the extent and location of proposed regeneration condition classes for cypress-tupelo swamps in the Atchafalaya Basin, LA. Only 6,175 ha (5.8%) of the 106,227 ha of cypress-tupelo forest in the Lower Atchafalaya Basin Floodway was classified as capable of naturally regenerating. Over 23% (24,525 ha) of the forest area was mapped as unable to regenerate either naturally or artificially. The loss and conversion of nearly 25,000 ha of cypress-tupelo forest would have significant and long-lasting impacts on ecosystem services such as wildlife habitat for birds and Louisiana black bears. Given the landscape-scale changes in surface elevations and flooding depths and durations throughout southern Louisiana, similar conditions and impacts are likely applicable to all coastal cypress-tupelo forests in Louisiana. Better data on flooding during the growing season are needed to more accurately identify and refine the location and spatial extent of the regeneration condition classes.

Louisiana↗
USGS2004

Urbanization impacts on the structure and function of forested wetlands

The exponential increase in population has fueled a significant demographic shift: 60% of the Earth's population will live in urban areas by 2030. While this population growth is significant in its magnitude, the ecological footprint of natural resource consumption and use required to sustain urban populations is even greater. The land use and cover changes accompanying urbanization (increasing human habitation coupled with resource consumption and extensive landscape modification) impacts natural ecosystems at multiple spatial scales. Because they generally occupy lower landscape positions and are linked to other ecosystems through hydrologic connections, the cascading effects of habitat alteration on watershed hydrology and nutrient cycling are particularly detrimental to wetland ecosystems. I reviewed literature relevant to these effects of urbanization on the structure and function of forested wetlands. Hydrologic changes caused by habitat fragmentation generally reduce species richness and abundance of plants, macroinvertebrates, amphibians, and birds with greater numbers of invasives and exotics. Reduction in soil saturation and lowered water tables result in greater nitrogen mineralization and nitrification in urban wetlands with higher probability of NO − 3 export from the watershed. Depressional forested wetlands in urban areas can function as important sinks for sediments, nutrients, and metals. As urban ecosystems become the predominant human condition, there is a critical need for data specific to urban forested wetlands in order to better understand the role of these ecosystems on the landscape.

Urban Ecosystems↗
USGS2009

Delayed conifer mortality after fuel reduction treatments: Interactive effects of fuel, fire intensity, and bark beetles

Many low‐elevation dry forests of the western United States contain more small trees and fewer large trees, more down woody debris, and less diverse and vigorous understory plant communities compared to conditions under historical fire regimes. These altered structural conditions may contribute to increased probability of unnaturally severe wildfires, susceptibility to uncharacteristic insect outbreaks, and drought‐related mortality. Broad‐scale fuel reduction and restoration treatments are proposed to promote stand development on trajectories toward more sustainable structures. Little research to date, however, has quantified the effects of these treatments on the ecosystem, especially delayed and latent tree mortality resulting directly or indirectly from treatments. In this paper, we explore complex hypotheses relating to the cascade of effects that influence ponderosa pine (Pinus ponderosa) and Douglas‐fir (Pseudotsuga menziesii) mortality using structural equation modeling (SEM). We used annual census and plot data through six growing seasons after thinning and four growing seasons after burning from a replicated, operational‐scale, completely randomized experiment conducted in northeastern Oregon, USA, as part of the national Fire and Fire Surrogate study. Treatments included thin, burn, thin followed by burn (thin + burn), and control. Burn and thin + burn treatments increased the proportion of dead trees while the proportion of dead trees declined or remained constant in thin and control units, although the density of dead trees was essentially unchanged with treatment. Most of the new mortality (96%) occurred within two years of treatment and was attributed to bark beetles. Bark beetle‐caused tree mortality, while low overall, was greatest in thin + burn treatments. SEM results indicate that the probability of mortality of large‐diameter ponderosa pine from bark beetles and wood borers was directly related to surface fire severity and bole charring, which in turn depended on fire intensity, which was greater in units where thinning increased large woody fuels. These results have implications when deciding among management options for restoring ecosystem health in similar ponderosa pine and Douglas‐fir forests.

Ecological Applications↗
USGS2014

The effects of atmospheric nitrogen deposition on terrestrial and freshwater biodiversity

This chapter reports the findings of a Working Group on how atmospheric nitrogen (N) deposition affects both terrestrial and freshwater biodiversity. Regional and global scale impacts on biodiversity are addressed, together with potential indicators. Key conclusions are that: the rates of loss in biodiversity are greatest at the lowest and initial stages of N deposition increase; changes in species compositions are related to the relative amounts of N, carbon (C) and phosphorus (P) in the plant soil system; enhanced N inputs have implications for C cycling; N deposition is known to be having adverse effects on European and North American vegetation composition; very little is known about tropical ecosystem responses, while tropical ecosystems are major biodiversity hotspots and are increasingly recipients of very high N deposition rates; N deposition alters forest fungi and mycorrhyzal relations with plants; the rapid response of forest fungi and arthropods makes them good indicators of change; predictive tools (models) that address ecosystem scale processes are necessary to address complex drivers and responses, including the integration of N deposition, climate change and land use effects; criteria can be identified for projecting sensitivity of terrestrial and aquatic ecosystems to N deposition. Future research and policy-relevant recommendations are identified.

Book chapter↗
USGS2017

Soil data for a thermokarst bog and the surrounding permafrost plateau forest, located at Bonanza Creek Long Term Ecological Research Site, Interior Alaska

Peatlands play an important role in boreal ecosystems, storing a large amount of soil organic carbon. In northern ecosystems, collapse-scar bogs (also known as thermokarst bogs) often form as the result of ground subsidence following permafrost thaw. To examine how ecosystem carbon balance changes with the loss of permafrost, we measured carbon and nitrogen storage within a thermokarst bog and the surrounding forest, which continues to have permafrost. These sites are a part of the Bonanza Creek Long Term Ecological Research (LTER) site and are located within Interior Alaska. Here, we report on methods used for core collection analysis as well as the cores’ physical, chemical, and descriptive properties.

Alaska↗
USGS1997

Potential effects of climate change on freshwater ecosystems of the New England/Mid-Atlantic Region

Numerous freshwater ecosystems, dense concentrations of humans along the eastern seaboard, extensive forests and a history of intensive land use distinguish the New England/Mid-Atlantic Region. Human population densities are forecast to increase in portions of the region at the same time that climate is expected to be changing. Consequently, the effects of humans and climatic change are likely to affect freshwater ecosystems within the region interactively. The general climate, at present, is humid continental, and the region receives abundant precipitation. Climatic projections for a 2 × CO 2 atmosphere, however, suggest warmer and drier conditions for much of this region. Annual temperature increases ranging from 3–5°C are projected, with the greatest increases occurring in autumn or winter. According to a water balance model, the projected increase in temperature will result in greater rates of evaporation and evapotranspiration. This could cause a 21 and 31% reduction in annual stream flow in the southern and northern sections of the region, respectively, with greatest reductions occurring in autumn and winter. The amount and duration of snow cover is also projected to decrease across the region, and summer convective thunderstorms are likely to decrease in frequency but increase in intensity.

Hydrological Processes↗
USGS2021

A new remote sensing-based Carbon Sequestration Potential Index (CSPI): A tool to support land carbon management

Integrating remote sensing into assessments of carbon stocks and fluxes has advanced our understanding of how global change affects landscapes and our capacity to support decision making about forest management. However, there remains a lack of detailed and actionable analyses conducted across widely ranging environmental conditions that are appropriate for tactical planning. We used airborne laser scanning data and multi-source satellite imagery to estimate forest aboveground carbon density and gross primary production, and to map forest cover across the main Hawaiian Islands. We used these measures to develop the Carbon Sequestration Potential Index (CSPI), which identifies where the potential for carbon sequestration following afforestation would be highest within a complex landscape of 304 management units. Variation in CSPI was high across islands and between ecosystems, with low values for cool, dry and largely intact forest systems and high values for warm, wet and largely non-forested systems. The CSPI provided a rapid, spatially-explicit and actionable assessment of Hawaiian forest reserves, which can help stewardship agencies contribute to state carbon neutrality goals through climate-smart and science-driven prescriptions that encompass conservation to restoration.

Hawaii↗
USGS2014

Temperate and boreal forest mega-fires: characteristics and challenges

Mega-fires are often defined according to their size and intensity but are more accurately described by their socioeconomic impacts. Three factors – climate change, fire exclusion, and antecedent disturbance, collectively referred to as the “mega-fire triangle” – likely contribute to today's mega-fires. Some characteristics of mega-fires may emulate historical fire regimes and can therefore sustain healthy fire-prone ecosystems, but other attributes decrease ecosystem resiliency. A good example of a program that seeks to mitigate mega-fires is located in Western Australia, where prescribed burning reduces wildfire intensity while conserving ecosystems. Crown-fire-adapted ecosystems are likely at higher risk of frequent mega-fires as a result of climate change, as compared with other ecosystems once subject to frequent less severe fires. Fire and forest managers should recognize that mega-fires will be a part of future wildland fire regimes and should develop strategies to reduce their undesired impacts.

Frontiers in Ecology and the Environment↗
USGS2011

Mapping and monitoring Louisiana's mangroves in the aftermath of the 2010 Gulf of Mexico Oil spill

Information regarding the present condition, historical status, and dynamics of mangrove forests is needed to study the impacts of the Gulf of Mexico oil spill and other stressors affecting mangrove ecosystems. Such information is unavailable for Louisiana at sufficient spatial and thematic detail. We prepared mangrove forest distribution maps of Louisiana (prior to the oil spill) at 1 m and 30 m spatial resolution using aerial photographs and Landsat satellite data, respectively. Image classification was performed using a decision-tree classification approach. We also prepared land-cover change pairs for 1983, 1984, and every 2 y from 1984 to 2010 depicting “ecosystem shifts” (e.g., expansion, retraction, and disappearance). This new spatiotemporal information could be used to assess short-term and long-term impacts of the oil spill on mangroves. Finally, we propose an operational methodology based on remote sensing (Landsat, Advanced Spaceborne Thermal Emission and Reflection Radiometer [ASTER], hyperspectral, light detection and ranging [LIDAR], aerial photographs, and field inventory data) to monitor the existing and emerging mangrove areas and their disturbance and regrowth patterns. Several parameters such as spatial distribution, ecosystem shifts, species composition, and tree height/biomass could be measured to assess the impact of the oil spill and mangrove recovery and restoration. Future research priorities will be to quantify the impacts and recovery of mangroves considering multiple stressors and perturbations, including oil spill, winter freeze, sea-level rise, land subsidence, and land-use/land-cover change for the entire Gulf Coast.

Journal of Coastal Research↗
USGS2016

Soil mercury distribution in adjacent coniferous and deciduous stands highly impacted by acid rain in the Ore Mountains, Czech Republic

Forests play a primary role in the cycling and storage of mercury (Hg) in terrestrial ecosystems. This study aimed to assess differences in Hg cycling and storage resulting from different vegetation at two adjacent forest stands - beech and spruce. The study site Načetín in the Czech Republic's Black Triangle received high atmospheric loadings of Hg from coal combustion in the second half of the 20th century as documented by peat accumulation rates reaching 100 μg m −2 y −1 . In 2004, the annual litterfall Hg flux was 22.5 μg m −2 y −1 in the beech stand and 14.5 μg m −2 y −1 in the spruce stand. Soil concentrations and pools of Hg had a strong positive relation to soil organic matter and concentrations of soil sulfur (S) and nitrogen (N). O-horizon Hg concentrations ranged from 245 to 495 μg kg −1 and were greater in the spruce stand soil, probably as a result of greater dry Hg deposition. Mineral soil Hg concentrations ranged from 51 to 163 μg kg −1 and were greater in the beech stand soil due to its greater capacity to store organic carbon (C). The Hg/C ratio increased with depth from 0.3 in the O-horizon to 3.8 μg g −1 in the C horizon of spruce soil and from 0.7 to 2.7 μg g −1 in beech soil. The Hg/C ratio was greater at all mineral soil depths in the spruce stand. The organic soil Hg pools in beech and spruce stands (6.4 and 5.7 mg m −2 , respectively) were considerably lower than corresponding mineral soil Hg pools (39.1 and 25.8 mg m −2 ). Despite the important role of S in Hg cycling, differences in soil Hg distribution at both stands could not be attributed to differences in soil sulfur speciation.

Ore Mountains↗
USGS2021

Introduced mangroves along the coast of Moloka‘i, Hawai‘i may represent novel habitats for megafaunal communities

Mangrove forests are prevalent along tropical/subtropical coastlines and provide valuable ecosystem services including coastline stabilization, storm impact reduction, and enhanced coastal productivity. However, mangroves were absent from the Hawaiian Islands and their introduction to Moloka‘i in 1902 has provided an opportunity to examine their unique influence on coastal landscapes. Previous studies indicate an inability of native detritivores to utilize tannin-rich substrates, yielding poor cycling of mangrove-derived detritus in Hawaiian tidal zones. We hypothesize that in addition to altering detrital inputs, introduced mangroves facilitate the persistence of introduced species in the Hawaiian coastal zone by providing novel habitat for juvenile megafauna. To determine whether mangrove-dominated tidal zones harbor megafaunal assemblages distinct from open sandflats, we sampled in two mangrove (M1 and M2) and two adjacent sandflat (S1 and S2) sites along the southern coast of Moloka‘i, where the most mature mangrove forests occur in Hawai‘i. There were no statistical differences in total abundances between M1 and M2 or S1 and S2; therefore, results from individual deployments were pooled across the sites in order to conduct between-habitat (mangrove vs. sandflat) comparisons. Our mangrove study site had significantly higher abundances of megafauna, including several shrimp and crab species, compared to the sandflat site. The community composition within the mangrove site differed from the sandflat site, including higher abundances of non-native mangrove crabs ( Scylla serrata ), as well as native fish Bathygobius cocosensis and crustaceans ( Thalamita crenata , Palaemon pacificus , P. debilis ) than in the sandflat site, indicating that the mangrove site may provide niches for both invasive and native species. In addition, mean body length for several similar species was smaller in the mangrove site than in the sandflat site, suggesting that these mangroves may be providing a habitat for juvenile species. While our study was spatially limited to two mangrove and two adjacent sandflat sites, our results suggest that introduced mangroves in Moloka‘i may support small-bodied, native, and non-native megafauna, influencing coastal Hawaiian trophic dynamics. Our case study provides a baseline for megafaunal fish and invertebrate communities present prior to non-native mangrove removal as well as for monitoring potential community changes following expansion of mangrove habitats due to climate change.

Hawaii↗
USGS2026

Initial condition uncertainty exerts a large and persistent influence on model simulations of ecosystem carbon dynamics in California

Uncertainties in terrestrial ecosystem models limit their predictive power. Efforts to reduce projection error have rarely focused on constraining uncertainty in the initial state of the ecosystem, however, despite evidence that matching model initial conditions to real-world observations reduces overall model bias. Here we use an integrated land change and carbon gain-loss model to evaluate the influence of initial condition uncertainty on simulations of California wildland ecosystems during the years 1985–2020. We generated 36 initial conditions scenarios by varying the source data used to initialize state variables and then ran simulations based on each of these scenarios under a constant set of historical conditions. We found that discrepancies in initial forest extent and initial forest age among scenarios generated wide uncertainty ranges in model estimates of terrestrial ecosystem carbon stocks and flux rates at the outset of the simulation period, but differences in initial forest composition had no impact. Over time, forest age became more homogeneous across model scenarios leading to exponential rates of decline in the uncertainty ranges of live biomass and dead wood carbon but little to no impact on uncertainties in litter and soil organic carbon. Uncertainties in individual carbon flux rates were consistent with uncertainties in their source pools. In contrast, model estimates of ecosystem carbon balance demonstrated a shift in system behavior not apparent in trends for individual carbon stocks and fluxes. Specifically, estimates of ecosystem carbon balance converged across scenarios for the first 20 years of the simulation period but then began to diverge at an accelerating rate, possibly due to weakened resilience to the increased frequency and severity of climate-driven disturbances. Our results demonstrate that uncertainty in the initial state of the system can have large and persistent impacts on the predictability of ecosystem carbon dynamics, and that ongoing shifts in external forcing by climate and climate-driven disturbances can exacerbate these impacts.

California↗
USGS2009

Forest Vegetation Monitoring Protocol for National Parks in the North Coast and Cascades Network

Plant communities are the foundation for terrestrial trophic webs and animal habitat, and their structure and species composition are an integrated result of biological and physical drivers (Gates, 1993). Additionally, they have a major role in geologic, geomorphologic and soil development processes (Jenny, 1941; Stevens and Walker, 1970). Throughout most of the Pacific Northwest, environmental conditions support coniferous forests as the dominant vegetation type. In the face of anthropogenic climate change, forests have a global role as potential sinks for atmospheric carbon (Goodale and others, 2002). Consequently, knowledge of the status of forests in the three large parks of the NCCN [that is, Mount Rainier (MORA), North Cascades (NOCA), and Olympic (OLYM) National Parks] is fundamental to understanding the condition of Pacific Northwest ecosystems. Diverse climate and soil properties across the Pacific Northwest result in a variety of forest types (Franklin and Dyrness, 1973; Franklin and others, 1988; Henderson and others, 1989, 1992). The mountainous terrain of Mount Rainier, North Cascades, and Olympic National Parks create steep elevational and precipitation gradients within and among the parks: collectively, these parks span from sea level to more than 4,200 m; and include areas with precipitation from 90 to more than 500 cm. The resulting forests range from coastal rainforests with dense understories and massive trees draped with epiphytes; to areas with drought-adapted Ponderosa pines; to high-elevation subalpine fir forests interspersed with meadows just below treeline (table 1). These forests, in turn, are the foundation for other biotic communities constituting Pacific Northwest ecosystems.

Techniques and Methods↗