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Survival and population size estimation in raptor studies: A comparison of two methods

ABSTRACT.--The Jolly-Seber model is a capture-recapture model that can provide less-biased survival and population size estimates than those produced from simple counting procedures. Parameter estimation by simple counts and Jolly-Seber methods are based on certain assumptions that directly determine the validity of estimates. Evuluation of assumptions for parameter estimation is a focus of this paper and used as a basis for determining which methods are more likely to produce better estimates. An example of population size and survival estimation for a peregrine falcon(Falco peregrinus) population in western Greenland is used to compare the two methods.Based on results from the Greenland peregrine population, and an assessment of the underlying assumptions of simple counts and the Jolly-Seber model,we suggest that Jolly-Seber estimation of survival and population size is less biased than simple counts in studies with marked birds. We recommend the use of a Jolly-Seber analysis of data when capture-recapture techniques are employed in raptor population studies.

Journal of Raptor Research↗

Nesting phenology and success of ring-necked ducks in east-central Maine

Selected aspects of the nesting biology of Ring-necked Ducks in east-central Maine were studied during 1983-1985 and compared to previous studies. Nesting chronology, clutch size, and hatching success were similar to results reported from previous studies, but nest success was 32-42% lower. Thirty-eight percent of the nests were successful; % of the eggs in successful nests hatched. Among unsuccessful nests, predation and flooding caused most failures.

Journal of Field Ornithology↗

Lead poisoning in captive Andean condors (Vultur gryphus)

Elevated lead in the tissues of raptors, especially those that scavenge, is a common occurrence, and lead poisoning appears to be a significant problem in the ongoing recovery effort for California condors (Gymnogyps californianus). Elevated blood lead levels have been found in released birds, and a number of birds have died of lead poisoning. In earlier work, we dosed turkey vultures (Cathartes aura) with lead shot but found them to be a poor model for lead poisoning. In this study, we dosed four Andean condors (Vultur gryphus) with lead shot and found them to be quite sensitive, as two of the birds died and the other two exhibit signs of lead poisoning within 50 days. All lead-responsive parameters were affected, and regurgitation of dosed shot occurred only once. The response of the Andean condors appeared to mimic California condors, suggesting that once exposed to lead, the possibility of survival is poor. This is consistent with observations in the wild, where otherwise healthy birds exposed to metallic lead quickly succumb. At the very least, the release program has to maintain constant surveillance and an active lead monitoring program.

Journal of Wildlife Diseases↗

Influence of tidal range on the stability of coastal marshland

Early comparisons between rates of vertical accretion and sea level rise across marshes in different tidal ranges inspired a paradigm that marshes in high tidal range environments are more resilient to sea level rise than marshes in low tidal range environments. We use field‐based observations to propose a relationship between vegetation growth and tidal range and to adapt two numerical models of marsh evolution to explicitly consider the effect of tidal range on the response of the marsh platform channel network system to accelerating rates of sea level rise. We find that the stability of both the channel network and vegetated platform increases with increasing tidal range. Our results support earlier hypotheses that suggest enhanced stability can be directly attributable to a vegetation growth range that expands with tidal range. Accretion rates equilibrate to the rate of sea level rise in all experiments regardless of tidal range, suggesting that comparisons between accretion rate and tidal range will not likely produce a significant relationship. Therefore, our model results offer an explanation to widely inconsistent field‐based attempts to quantify this relationship while still supporting the long‐held paradigm that high tidal range marshes are indeed more stable.

Journal of Geophysical Research↗

Use of isotopically labeled fertilizer to trace nitrogen fertilizer contributions to surface, soil, and ground water

The fate and transport of a single N fertilizer application through plants, soil, runoff, and the unsaturated and saturated zones was determined for four years at a field site under continuous corn (Zea mays L.) management. Claypan soils, which underlie the site, were hypothesized to restrict the movement of agrichemicals from the soil surface to ground water. However, N fertilizer moved rapidly through preferential flow paths in the soil and into the underlying glacial till aquifer. Most N transport occurred during the fall and winter when crops were not available to use excess N. Forty months after application, 33 percent of the fertilizer had been removed by grain harvests, 30 percent had been transpired to the atmosphere, and 33 percent had migrated to ground water. Although runoff volumes were 50 percent greater than infiltration, less than 2 percent of the fertilizer was lost to runoff. Small measured denitrification rates and large measured dissolved oxygen concentrations in ground water favor the long-term stability of NO3-1 in ground water. Successive fertilizer applications, in areas that lack the ability to moderate N concentrations through consumptive N reactions, risk the potential of N-saturated ecosystems.

Journal of Environmental Hydrology↗

Geology of El Chichon volcano, Chiapas, Mexico

The (pre-1982) 850-m-high andesitic stratovolcano El Chicho??n, active during Pleistocene and Holocene time, is located in rugged, densely forested terrain in northcentral Chiapas, Me??xico. The nearest neighboring Holocene volcanoes are 275 km and 200 km to the southeast and northwest, respectively. El Chicho??n is built on Tertiary siltstone and sandstone, underlain by Cretaceous dolomitic limestone; a 4-km-deep bore hole near the east base of the volcano penetrated this limestone and continued 770 m into a sequence of Jurassic or Cretaceous evaporitic anhydrite and halite. The basement rocks are folded into generally northwest-trending anticlines and synclines. El Chicho??n is built over a small dome-like structure superposed on a syncline, and this structure may reflect cumulative deformation related to growth of a crustal magma reservoir beneath the volcano. The cone of El Chicho??n consists almost entirely of pyroclastic rocks. The pre-1982 cone is marked by a 1200-m-diameter (explosion?) crater on the southwest flank and a 1600-m-diameter crater apparently of similar origin at the summit, a lava dome partly fills each crater. The timing of cone and dome growth is poorly known. Field evidence indicates that the flank dome is older than the summit dome, and K-Ar ages from samples high on the cone suggest that the flank dome is older than about 276,000 years. At least three pyroclastic eruptions have occurred during the past 1250 radiocarbon years. Nearly all of the pyroclastic and dome rocks are moderately to highly porphyritic andesite, with plagioclase, hornblende and clinopyroxene the most common phenocrysts. Geologists who mapped El Chicho??n in 1980 and 1981 warned that the volcano posed a substantial hazard to the surrounding region. This warning was proven to be prophetic by violent eruptions that occurred in March and April of 1982. These eruptions blasted away nearly all of the summit dome, blanketed the surrounding region with tephra, and sent pyroclastic flows down radial drainages on the flanks of the cone; about 0.3 km3 of material (density of all products normalized to 2.6 g cm-3) was erupted. More debris entered the stratosphere than from any other volcanic eruption within at least the past two decades. Halite and a calcium sulfate mineral (anhydrite?) recovered from the stratospheric cloud, and anhydrite as a common accessory mineral in 1982 juvenile erupted products may reflect contamination of El Chicho??n magma by the evaporite sequence revealed by drilling. ?? 1984.

Journal of Volcanology and Geothermal Research↗

Anomalous chloride flux discharges from Yellowstone National Park

The chloride concentration of some thermal springs in and adjacent to Yellowstone National Park is constant through time although their discharge varies seasonally. As a result the chloride flux from these springs increases during periods of increased discharge. We believe that this is caused by changes in the height of the local groundwater table, which affects the discharge of the springs but not their chloride concentration. The discharge from Mammoth Hot Springs varies seasonally, but its chloride concentration remains constant. We take this as evidence that this major thermal feature is derived from orifices that are tapping the local water table close to its surface. Three of the four major rivers (Yellowstone, Snake and Falls) exiting the Park also show an increased chloride flux during the spring runoff that cannot be explained solely by the contribution of snowmelt, nor by release of hot-spring-derived chloride stored in the soil during the winter and released in the spring. The increased chloride flux in these rivers is attributed to their draining shallow hot springs similar to those mentioned above. In contrast to the Yellowstone, Snake and Falls Rivers, the Firehole and Gibbon Rivers, which unite to form the Madison River and which collectively drain several major geyser basins, display a poor correlation between chloride flux and discharge. The cause, we believe, is that a large part of the thermal water input to these two rivers originated at great depths where the seasonal variation in the height of the water table had a negligible effect on hot spring discharge. Monitoring of seasonal discharge and chloride concentration of thermal features yields information on the depths at which these thermal features tap the local water table. ?? 1990.

Journal of Volcanology and Geothermal Research↗

Toggling of seismicity by the 1997 Kagoshima earthquake couplet: A demonstration of time-dependent stress transfer

Two M ??? 6 well-recorded strike-slip earthquakes struck just 4 km and 48 days apart in Kagoshima prefecture, Japan, in 1997, providing an opportunity to study earthquake interaction. Aftershocks are abundant where the Coulomb stress is calculated to have been increased by the first event, and they abruptly stop where the stress is dropped by the second event. This ability of the main shocks to toggle seismicity on and off argues that static stress changes play a major role in exciting aftershocks, whereas the dynamic Coulomb stresses, which should only promote seismicity, appear to play a secondary role. If true, the net stress changes from a sequence of earthquakes might be expected to govern the subsequent seismicity distribution. However, adding the stress changes from the two Kagoshima events does not fully capture the ensuing seismicity, such as its rate change, temporal decay, or migration away from the ends of the ruptures. We therefore implement a stress transfer model that incorporates rate/state friction, in which seismicity is treated as a sequence of independent nucleation events that are dependent on the fault slip, slip rate, and elapsed time since the last event. The model reproduces the temporal response of seismicity to successive stress changes, including toggling, decay, and aftershock migration. Nevertheless, the match of observed to predicted seismicity is quite imperfect, due perhaps to inadequate knowledge of several model parameters. However, to demonstrate the potential of this approach, we build a probabilistic forecast of larger earthquakes on the expected rate of small aftershocks, taking advantage of the large statistical sample the small shocks afford. Not surprisingly, such probabilities are highly time- and location-dependent: During the first decade after the main shocks, the seismicity rate and the chance of successive large shocks are about an order of magnitude higher than the background rate and are concentrated exclusively in the stress triggering zones. Copyright 2003 by the American Geophysical Uion.

Journal of Geophysical Research B: Solid Earth↗

Seasonal survival of radiomarked emperor geese in western Alaska

The population of emperor geese ( Chen canagica ) in western Alaska, USA, declined by >50% from the 1960s to the mid‐1980s and has increased only slightly since. Rates of population increase among arctic geese are especially sensitive to changes in adult survival. Improving adult survival in seasons or geographic areas where survival is low may be the best means of increasing the emperor goose population. We monitored fates of 133 adult female emperor geese that were radiomarked with surgically implanted very high frequency or satellite radiotransmitters from 1999 to 2004 to assess whether monthly survival varied among years, seasons, or geographic areas. Because of uncertainties in determining whether a bird had died based on the radio signal, we analyzed 2 versions of the data. One version used conservative criteria to identify which birds had died based on radio signals and the other used more liberal criteria. In the conservative version of the data we detected 12 mortalities of emperor geese, whereas in the liberal interpretation there were 18 mortalities. In both versions, the models with greatest support indicated that monthly survival varied seasonally and that compared to most seasons estimated monthly survival was lower (φ = 0.95–0.98) in May and August when emperor geese were mainly on the Yukon‐Kuskokwim Delta. From 44% to 47% of annual mortality occurred in those months. Estimated monthly survival was higher (φ = 0.98–1.0) from September through March when emperor geese were at autumn staging or wintering areas and in June and July when birds were nesting, rearing broods, or molting. Estimated annual survival was 0.85 (95% CI = 0.77–0.92) in the best‐supported model when we used conservative criteria to identify mortalities and 0.79 (95% CI = 0.74–0.85) under the best model using liberal mortality criteria. Lower survival in August and May corresponded to periods when subsistence harvest of emperor geese was likely highest. Managers may be able to most effectively influence population growth rate of emperor geese by reducing subsistence harvest on the Yukon‐Kuskokwim Delta in May and August.

Alaska↗

Regional gravity and magnetic anomalies in the eastern Snake River Plain, Idaho

Over the eastern Snake River Plain, the Bouguer gravity anomaly and the magnetic intensity are, in general, high. In detail, both the gravity and the magnetic anomalies are a complex of highs and lows, in contrast to the simpler anomalies over the western Snake River Plain. The broad gravity high associated with the eastern Snake River Plain cannot be produced by a dense mass at shallow depth under the plain, but must be produced either by a deep feature under the plain or by a broader, shallower feature extending well beyond the plain. The high could be produced by a thinning of the crust under the plain by 4.5 to 7 kilometers. The data suggest that the Cenozoic rocks of the plain are underlain by rocks that have density and magnetization similar to the density and magnetization of the pre-Cenozoic rocks north and south of the plain. The data do not suggest rifting of the upper crust such as is inferred for the western Snake River Plain. The plain is in approximate isostatic equilibrium with adjoining regions to the north and south and thus contrasts with the adjoining areas where the basin-and-range topography is not in isostatic equilibrium. The local gravity anomalies on the plain are probably produced by variations in the thickness of Cenozoic rocks, but the wavelength of the anomalies is about twice that of the basin-and-range structural features in areas adjoining the plain. The gravity expression of basin-and-range structural features does not extend far onto the plain. Magnetic anomalies on the plain are produced by Cenozoic volcanic rocks.

Idaho↗

Wood River mining district, Idaho - intrusion-related lead-silver deposits derived from country rock source

Lead-silver deposits in the Wood River mining district occur in shear zones in hornfelsed argillite of the Devonian Milligen Formation near granitic plutons and under the Wood River thrust fault. The principal ore minerals are argentiferous galena and sphalerite; siderite is the principal gangue. The δ 34 S values of the sulfide minerals range from +2.2 to +15.0 permil, indicating that the sulfur had a shallow crustal source. Δ 34 S values between sphalerite and galena range from +2.3 to +3.4 permil, corresponding to sulfur isotope temperatures between 280° and 182 °C. Hydrothermal barite has a δ 34 S of +13.2 permil. Lead isotope ratios are radiogenic, also pointing to a shallow crustal source. Quartz gangue has δ 18 O of +16.4 permil and a calculated δ 18 O H20 at 270°C of +8.4 permil. This value is reasonable for a hydrothermal fluid that had reached equilibrium with the argillite country rock. The siderite gangue has δ 18 O and δ 13 C values of +14.0 and -5.5 permil, respectively. Fluid inclusions have homogenization temperatures of 244°-307°C and average 270°C. Freezing-stage measurements ranged from -1.85 to -2.8°C, suggesting salinities of 3.2 to 4.8 weight percent. The δD values of inclusion fluid in ore and gangue minerals are -110 to -120 permil. The geology, isotope, and fluid-inclusion data are consistent with a model of hydrothermal systems of meteoric water in faulted and shattered Paleozoic rocks near plutonic masses. This environment permitted deep circulation of the hydrothermal fluids, which dissolved the metals and sulfur from the Paleozoic host rocks and deposited ore in favorable beds or structures under the regional Wood River thrust fault.

Idaho↗

The effects of the Hurricane Agnes flood on channel geometry and sediment discharge of selected streams in the Susquehanna River basin, Pennsylvania

The Hurricane Agnes flood seems to have hardly changed the channel geometry of the Pennsylvania streams studied in this report. The 10 sites studied generally showed that the width of the stream channels had been changed little by the flood and that streambed altitudes had been lowered less than a foot (0.3 m). The velocity of the streams at a given discharge had decreased. In comparison, the sediment deposited by the December 1964 flood in northwestern California commonly raised the streambed altitudes several feet. The differences in the effects of the two floods may have been produced by the availability of sediment for transport, by the relative size of sediment carried in suspension or bedload, or by the relative magnitudes of the floods.

Pennsylvania↗

A typical cross section based on magnetic data of lower and middle Keweenawan volcanic rocks, Ironwood area, Michigan

A north-trending aeromagnetic profile of a sequence of east-striking Keweenawan volcanic rocks near Ironwood, Mich., can be matched to a calculated profile over a model consisting of a series of dipping layers. (The dips were those measured by H. A. Hubbard along the north-trending valley of the Black River.) Remanent and induced magnetizations of 39 oriented cores from the Black River valley were determined by K. G. Books; felsite from Chippewa Hill and basalt from Algonquin Falls of middle Keweenawan age have normal Keweenawan magnetization and lower Keweenawan Powder Mill rocks have reverse magnetization. In the model these magetizations were assigned to 26 layers which alternated with very weakly magnetized layers. The best match of the calculated composite anomalies of remanent and induced magnetization and the aeromagnetic profile occurs if a deeper block of steeply dipping Powder Mill rocks is assumed to underlie the middle Keweenawan flows.

Michigan↗