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At least 523 records · Page 29Linked to original sources
USGS2022

Evidence of a dietary shift by the Florida manatee (Trichechus manatus latirostris) in the Indian River Lagoon inferred from stomach content analyses

Investigating the long-term fluctuations of the feeding ecology of megaherbivores such as sirenians is important, as any changes could be indicative of shifts in resource availability. The Indian River Lagoon (IRL), eastern Florida, USA, is a critical habitat for the Florida manatee ( Trichechus manatus latirostris ). However, the IRL has experienced a substantial decline in seagrass due to the persistence of several harmful algal blooms. Using microhistological analysis, we examined the diet of manatees over a discontinuous sampling period spanning over 38 years using stomach contents collected from carcasses recovered in the IRL. Samples collected between 2013–2015 (post-seagrass die-off, n = 90) were compared to archived stomach samples collected between 1977–1989 (pre-seagrass die-off, n = 103). Samples analyzed from 1977–1989 contained primarily seagrasses (61.7%), followed by algae (28.4%) and vascular plants (1.7%). In contrast, stomach samples from the post-seagrass die-off primarily contained algae (49.5%), followed by seagrasses (34%) and vascular plants (2.7%). Between 1977–1989 and 2013–2015, manatees in the IRL experienced a 44.9% decline in seagrass consumption, and a 74.3% increase in algal consumption. This dietary shift was not influenced by body length, a proxy of age, or sex. Our results indicate that the dietary shift experienced by manatees is due to the decline of available seagrass forage in the IRL, and highlight the dietary plasticity of manatees in the face of changes in resource availability. However, the individual health and population-level consequences of this dietary shift are unknown. An increase in mortality due to undetermined causes in this region since 2012 can be associated with deteriorating body conditions of manatees in the IRL, possibly resulting from a lack of seagrass diet. Future research should further investigate behavioral changes affecting manatees in relation to seagrass decline in the IRL, including the energetic costs of this dietary change.

Florida↗
USGS2026

Detection of Naegleria fowleri in thermally impacted recreational waters of western United States national parks

Naegleria fowleri is a thermophilic free-living amoeba (FLA) and the causative agent of primary amoebic meningoencephalitis, posing public health risks in warm freshwater environments. This multiyear, multiagency study surveyed 40 thermally impacted recreational waters across five western United States national parks and recreation areas–Yellowstone National Park, Grand Teton National Park, Olympic National Park, Newberry National Volcanic Monument, and Lake Mead National Recreation Area–to assess N. fowleri presence, concentration, and associated environmental conditions. A total of 185 water samples were analyzed by qPCR and Sanger sequencing, revealing widespread detection of N. fowleri in 34% of samples with positive detections from Lake Mead, Yellowstone, and Grand Teton hot springs and thermally impacted waters, with concentrations ranging from 4.9 to 115.7 cells/L. Multiple codetections of N. fowleri with nonpathogenic species including Naegleria australiensis were identified, suggesting they may inhabit similar ecological niches in the natural systems in contrast to engineered systems. These findings indicate that N. fowleri is present in thermally impacted areas across the western United States and underscore the use of enhanced monitoring, public awareness, and risk management strategies in thermally influenced recreational waters.

Arizona, Nevada, Oregon, Washington, Wyoming↗
USGS2009

Science-Based Strategies for Sustaining Coral Ecosystems

Coral ecosystems and their natural capital are at risk. Greenhouse gas emissions, overfishing, and harmful land-use practices are damaging our coral reefs. Overwhelming scientific evidence indicates that the threats are serious, and if they are left unchecked, the ecological and social consequences will be significant and widespread. Although the primary stressors to coral ecosystems are known, science-based strategies are needed to more accurately explain natural processes and forecast human-induced change. Collaborations among managers and scientists and enhanced mapping, monitoring, research, and modeling can lead to effective mitigation plans. U.S. Geological Survey scientists and their partners assess coral ecosystem history, ecology, vulnerability, and resiliency and provide study results to decisionmakers who may devise policies to sustain coral resources and the essential goods and services they provide.

Fact Sheet↗
USGS2011

Assessment of potential effects of water produced from coalbed natural gas development on macroinvertebrate and algal communities in the Powder River and Tongue River, Wyoming and Montana, 2010

Ongoing development of coalbed natural gas in the Powder River structural basin in Wyoming and Montana led to formation of an interagency aquatic task group to address concerns about the effects of the resulting production water on biological communities in streams of the area. Ecological assessments, made from 2005–08 under the direction of the task group, indicated biological condition of the macroinvertebrate and algal communities in the middle reaches of the Powder was lower than in the upper or lower reaches. On the basis of the 2005–08 results, sampling of the macroinvertebrate and algae communities was conducted at 18 sites on the mainstem Powder River and 6 sites on the mainstem Tongue River in 2010. Sampling-site locations were selected on a paired approach, with sites located upstream and downstream of discharge points and tributaries associated with coalbed natural gas development. Differences in biological condition among site pairs were evaluated graphically and statistically using multiple lines of evidence that included macroinvertebrate and algal community metrics (such as taxa richness, relative abundance, functional feeding groups, and tolerance) and output from observed/expected (O/E) macroinvertebrate models from Wyoming and Montana. Multiple lines of evidence indicated a decline in biological condition in the middle reaches of the Powder River, potentially indicating cumulative effects from coalbed natural gas discharges within one or more reaches between Flying E Creek and Wild Horse Creek in Wyoming. The maximum concentrations of alkalinity in the Powder River also occurred in the middle reaches. Biological condition in the upper and lower reaches of the Powder River was variable, with declines between some site pairs, such as upstream and downstream of Dry Fork and Willow Creek, and increases at others, such as upstream and downstream of Beaver Creek. Biological condition at site pairs on the Tongue River showed an increase in one case, near the Wyoming-Montana border, and a decrease in another case, upstream of Tongue River Reservoir. Few significant differences were noted from upstream to downstream of Prairie Dog Creek, a major tributary to the Tongue River. Further study would be needed to confirm the observed patterns and choose areas to examine in greater detail.

Wyoming;Montana↗
USGS2019

Let’s agree to disagree: Comparing auto-acoustic identification programs for northeastern bats

With the declines in abundance and changing distribution of white-nose syndrome–affected bat species, increased reliance on acoustic monitoring is now the new “normal.” As such, the ability to accurately identify individual bat species with acoustic identification programs has become increasingly important. We assessed rates of disagreement between the three U.S. Fish and Wildlife Service–approved acoustic identification software programs (Kaleidoscope Pro 4.2.0, Echoclass 3.1, and Bat Call Identification 2.7d) and manual visual identification using acoustic data collected during summers from 2003 to 2017 at Fort Drum, New York. We assessed the percentage of agreement between programs through pairwise comparisons on a total nightly count level, individual file level (e.g., individual echolocation pass call file), and grouped maximum likelihood estimate level (e.g., probability values that a species is misclassified as present when in fact it is absent) using preplanned contrasts, Akaike Information Criterion, and annual confusion matrices. Interprogram agreement on an individual file level was low, as measured by Cohen's Kappa (0.2–0.6). However, site-night level pairwise comparative analysis indicated that program agreement was higher (40–90%) using single season occupancy metrics. In comparing analytical outcomes of our different datasets (i.e., how comparable programs and visual identification are regarding the relationship between environmental conditions and bat activity), we determined high levels of congruency in the relative rankings of the model as well as the relative level of support for each individual model. This indicated that among individual software packages, when analyzing bat calls, there was consistent ecological inference beyond the file-by-file level at the scales used by managers. Depending on objectives, we believe our results can help users choose automated software and maximum likelihood estimate thresholds more appropriate for their needs and allow for better cross-comparison of studies using different automated acoustic software.

Journal of Fish and Wildlife Management↗
USGS2019

Phenology patterns indicate recovery trajectories of ponderosa pine forests after high-severity fires

Post-fire recovery trajectories in ponderosa pine (Pinus ponderosa Laws.) forests of the US Southwest are increasingly shifting away from pre-burn vegetation communities. This study investigated whether phenological metrics derived from a multi-decade remotely sensed imagery time-series could differentiate among grass, evergreen shrub, deciduous, or conifer-dominated replacement pathways. We focused on 10 fires that burned ponderosa pine forests in Arizona and New Mexico, USA before the year 2000. A total of 29 sites with discernable post-fire recovery signals were selected within high-severity burn areas. At each site, we used Google Earth Engine to derive time-series of normalized difference vegetation index (NDVI) signals from Landsat Thematic Mapper, Enhanced Thematic Mapper+, and Operational Line Imager data from 1984 to 2017. We aggregated values to 8- and 16-day intervals, fit Savitzy-Golay filters to each sequence, and extracted annual phenology metrics of amplitude, base value, peak value, and timing of peak value in the Timesat analysis package. Results show that relative to post-fire conditions, pre-burn ponderosa pine forests exhibit significantly lower mean NDVI amplitude (0.14 vs. 0.21), higher mean base NDVI (0.47 vs. 0.22), higher mean peak NDVI (0.60 vs. 0.43), and later mean peak NDVI (day of year 277 vs. 237). Vegetation succession exhibits distinct phenometric characteristics as early as year five (amplitude) and as late as year 20 (timing of peak NDVI). This study confirms the feasibility of leveraging phenology metrics derived from long-term imagery time series to identify and monitor ecological outcomes. This information may be of benefit to land resource managers who seek indicators of future landscape composition to inform management strategies.

Arizona, New Mexico↗
USGS2005

Are all prey created equal? A review and synthesis of differential predation on prey in substandard condition

Our understanding of predator-prey interactions in fishes has been influenced largely by research assuming that the condition of the participants is normal. However, fish populations today often reside in anthropogenically altered environments and are subjected to many kinds of stressors, which may reduce their ecological performance by adversely affecting their morphology, physiology, or behaviour. One consequence is that either the predator or prey, or both, may be in a substandard condition at the time of an interaction. We reviewed the literature on predator-prey interactions in fishes where substandard prey were used as experimental groups. Although most of this research indicates that such prey are significantly more vulnerable to predation, prey condition has rarely been considered in ecological theory regarding predator-prey interactions. The causal mechanisms for increased vulnerability of substandard prey to predation include a failure to detect predators, lapses in decision-making, poor fast-start performance, inability to shoal effectively, and increased prey conspicuousness. Despite some problems associated with empirical predator-prey studies using substandard prey, their results can have theoretical and applied uses, such as in ecological modelling or justification of corrective measures to be implemented in the wild. There is a need for more corroborative field experimentation, a better understanding of the causal mechanisms behind differential predation, and increased incorporation of prey condition into the research of predator-prey modellers and theoreticians. If the concept of prey condition is considered in predator-prey interactions, our understanding of how such interactions influence the structure and dynamics of fish communities is likely to change, which should prove beneficial to aquatic ecosystems.

Journal of Fish Biology↗
USGS2017

Divergent migration within lake sturgeon (Acipenser fulvescens) populations: Multiple distinct patterns exist across an unrestricted migration corridor

Population structure, distribution, abundance, and dispersal arguably underpin the entire field of animal ecology, with consequences for regional species persistence, and provision of ecosystem services. Divergent migration behaviours among individuals or among populations is an important aspect of the ecology of highly-mobile animals, allowing populations to exploit spatially- or temporally-distributed food and space resources. This study investigated the spatial ecology of lake sturgeon ( Acipenser fulvescens ) within the barrier free Huron-Erie Corridor (HEC), which connects Lake Huron and Lake Erie of the North American Laurentian Great Lakes. Over six years (2011 – 2016), movements of 268 lake sturgeon in the HEC were continuously monitored across the Great Lakes using acoustic telemetry (10 yr battery life acoustic transmitters). Five distinct migration behaviours were identified with hierarchical cluster analysis, based on the phenology and duration of river and lake use. Lake sturgeon in the HEC were found to contain a high level of intraspecific divergent migration, including partial migration with the existence of residents. Specific behaviours included year-round river residency and multiple lake-migrant behaviours that involved movements between lakes and rivers. Over 85% of individuals were assign to migration behaviours as movements were consistently repeated over the study, which suggested migration behaviours were consistent and persistent in lake sturgeon. Differential use of specific rivers or lakes by acoustic-tagged lake sturgeon further subdivided individuals into 14 “contingents” (spatiotemporally segregated subgroups). Contingents associated with one river (Detroit or St. Clair) were rarely detected in the other river, which confirmed that lake sturgeon in the Detroit and St. Clair represent two semi-independent populations that could require separate management consideration for their conservation. The distribution of migration behaviours did not vary between populations, sexes, body size, or among release locations, which indicated that intrapopulation variability in migratory behaviour is a general feature of the spatial ecology of lake sturgeon in un-fragmented landscapes.

Michigan, Ontario↗
USGS2016

A method for characterizing late-season low-flow regime in the upper Grand Ronde River Basin, Oregon

This report describes a method for estimating ecologically relevant low-flow metrics that quantify late‑season streamflow regime for ungaged sites in the upper Grande Ronde River Basin, Oregon. The analysis presented here focuses on sites sampled by the Columbia River Inter‑Tribal Fish Commission as part of their efforts to monitor habitat restoration to benefit spring Chinook salmon recovery in the basin. Streamflow data were provided by the U.S. Geological Survey and the Oregon Water Resources Department. Specific guidance was provided for selection of streamgages, development of probabilistic frequency distributions for annual 7-day low-flow events, and regionalization of the frequency curves based on multivariate analysis of watershed characteristics. Evaluation of the uncertainty associated with the various components of this protocol indicates that the results are reliable for the intended purpose of hydrologic classification to support ecological analysis of factors contributing to juvenile salmon success. They should not be considered suitable for more standard water-resource evaluations that require greater precision, especially those focused on management and forecasting of extreme low-flow conditions.

Oregon↗
USGS2000

Restoring ecological integrity of great rivers: Historical hydrographs aid in defining reference conditions for the Missouri River

Restoring the ecological integrity of regulated large rivers necessitates characterizing the natural flow regime. We applied 'Indicators of Hydrologic Alteration' to assess the natural range of variation of the Missouri River's flow regime at 11 locations before (1929-1948) and after (1967-1996) mainstem impoundment. The 3768 km long Missouri River was divided into three sections: upper basin least-altered from flow regulation, including the lower Yellowstone River; middle basin inter-reservoir, and lower basin channelized. Flow regulation was associated with a reduction in magnitude and duration of the annual flood pulse, an increase in magnitude and duration of annual discharge minima, a reduction in frequency of annual low-flow pulses, earlier timing of March-October low-flow pulses, and a general increase in frequency of flow reversals with a reduction in the rate of change in river flows. Hydrologic alterations were smallest at two least-altered upper-basin sites and most frequent and severe in inter-reservoir and upper-channelized river sections. The influence of reservoir operations on depressing the annual flood pulse was partially offset by tributary inflow in the lower 600 km of river. Reservoir operations could be modified to more closely approximate the 1929-1948 flow regime to establish a simulated natural riverine ecosystem. For inter-reservoir and upper channelized-river sections, we recommend periodic controlled flooding through managed reservoir releases during June and July; increased magnitude, frequency and duration of annual high-flow pulses; and increased annual rates of hydrograph rises and falls. All of the regulated Missouri River would benefit from reduced reservoir discharges during August-February, modified timing of reservoir releases and a reduced number of annual hydrograph reversals. Assessment of ecological responses to a reregulation of Missouri River flows that more closely approximates the natural flow regime should then be used in an adaptive fashion to further adjust reservoir operations.

Conference Paper↗
USGS2020

Climate dipoles as continental drivers of plant and animal populations

Ecological processes, such as migration and phenology, are strongly influenced by climate variability. Studying these processes often relies on associating observations of animals and plants with climate variability indices, such as the El Niño–Southern Oscillation. A characteristic of climate indices is the simultaneous emergence of opposite extremes of temperature and precipitation across continental scales, known as climate dipoles. The role of climate dipoles in shaping ecological and evolutionary processes has been largely overlooked. We review emerging evidence that climate dipoles can entrain species dynamics, and offer a framework for identifying ecological dipoles using broad-scale biological data. Given future changes in climatic and atmospheric processes, climate and ecological dipoles will likely shift in their intensity, distribution, and timing.

Trends in Ecology and Evolution↗
USGS2010

Stage structure alters how complexity affects stability of ecological networks

Resolving how complexity affects stability of natural communities is of key importance for predicting the consequences of biodiversity loss. Central to previous stability analysis has been the assumption that the resources of a consumer are substitutable. However, during their development, most species change diets; for instance, adults often use different resources than larvae or juveniles. Here, we show that such ontogenetic niche shifts are common in real ecological networks and that consideration of these shifts can alter which species are predicted to be at risk of extinction. Furthermore, niche shifts reduce and can even reverse the otherwise stabilizing effect of complexity. This pattern arises because species with several specialized life stages appear to be generalists at the species level but act as sequential specialists that are hypersensitive to resource loss. These results suggest that natural communities are more vulnerable to biodiversity loss than indicated by previous analyses.

Ecology Letters↗
USGS2006

A regional assessment of salt marsh restoration and monitoring in the Gulf of Maine

We compiled salt marsh monitoring datasets from 36 complete or imminent restoration projects in the Gulf of Maine to assess regional monitoring and restoration practices. Data were organized by functional indicators and restoration project types (culvert replacement, excavation works, or ditch plugging) then pooled to generate mean values for indicators before restoration, after restoration, and at reference sites. Monitoring data were checked against the regional standards of a voluntary protocol for the Gulf of Maine. Data inventories showed that vegetation and salinity indicators were most frequently collected (89 and 78% of sites, respectively), whereas nekton, bird, and hydrologic measures were collected at only about half of the sites. Reference conditions were monitored at 72% of sites. Indicators were analyzed to see if project sites were degraded relative to reference areas and to detect ecological responses to restoration activities. Results showed that compared to reference areas, prerestoration sites had smaller tidal ranges, reduced salinity levels, greater cover of brackish plants species, and lower cover of halophyte plants. Following restoration, physical factors rebounded rapidly with increased flood and salinity levels after about one year, especially for culvert projects. Biological responses were less definitive and occurred over longer time frames. Plant communities trended toward recovered halophytes and reduced brackish species at 3+ years following restoration. Nekton and avian indicators were indistinguishable among reference, impacted, and restored areas. The protocol was successful in demonstrating restoration response for the region, but results were limited by regional inconsistencies in field practices and relatively few multiyear datasets. To improve future assessment capabilities, we encourage greater adherence to the standard protocol throughout the Gulf of Maine salt marsh restoration community.

Restoration Ecology↗
USGS2024

Declining ecological resilience and invasion resistance under climate change in the sagebrush region, United States

In water-limited dryland ecosystems of the Western United States, climate change is intensifying the impacts of heat, drought, and wildfire. Disturbances often lead to increased abundance of invasive species, in part, because dryland restoration and rehabilitation are inhibited by limited moisture and infrequent plant recruitment events. Information on ecological resilience to disturbance (recovery potential) and resistance to invasive species can aid in addressing these challenges by informing long-term restoration and conservation planning. Here, we quantified the impacts of projected future climate on ecological resilience and invasion resistance (R&R) in the sagebrush region using novel algorithms based on ecologically relevant and climate-sensitive predictors of climate and ecological drought. We used a process-based ecohydrological model to project these predictor variables and resulting R&R indicators for two future climate scenarios and 20 climate models. Results suggested widespread future R&R decreases (24%–34% of the 1.16 million km 2 study area) that are generally consistent among climate models. Variables related to rising temperatures were most strongly linked to decreases in R&R indicators. New continuous R&R indices quantified responses to climate change; particularly useful for areas without projected change in the R&R category but where R&R still may decrease, for example, some of the areas with a historically low R&R category. Additionally, we found that areas currently characterized as having high sagebrush ecological integrity had the largest areal percentage with expected declines in R&R in the future, suggesting continuing declines in sagebrush ecosystems. One limitation of these R&R projections was relatively novel future climatic conditions in particularly hot and dry areas that were underrepresented in the training data. Including more data from these areas in future updates could further improve the reliability of the projections. Overall, these projected future declines in R&R highlight a growing challenge for natural resource managers in the region, and the resulting spatially explicit datasets provide information that can improve long-term risk assessments, prioritizations, and climate adaptation efforts.

western United States↗
USGS2023

Landscape-scale drivers of tayra abundance in the Ecuadorian Andes

Habitat conversion to agriculture and overexploitation of wildlife are the two largest drivers of biodiversity loss globally. Biodiversity loss is especially prevalent in areas undergoing rapid economic development at the expense of natural land cover as is the case across much of South America. Despite expected declines in wildlife populations associated with ongoing large-scale land conversions, for many species we lack sufficient data on key threats and drivers of abundance in order to inform appropriate management and conservation. Collecting data to estimate critical state variables such as abundance can be expensive, logistically challenging, and even implausible on spatial scales relevant to species management, especially for carnivores which are elusive and difficult to monitor. Here, we use detection-non-detection data collected using a structured camera trap survey repeated over two years in the Ecuadorian Andes to produce insights into the habitat associations and potential threats faced by the tayra, a medium-sized carnivore that despite being perceived to be relatively common in South America, remains largely understudied. We use hierarchical modelling to estimate an index of abundance for the tayra while accounting for imperfect detection. We demonstrate the tayra to be a lowland habitat generalist, with conversion of land to agriculture potentially benefitting this species in the short term, with increasing proportion of core agricultural land being associated with higher indices of abundance. However, we highlight that this state could potentially serve as an ecological trap in the long term. We provide evidence for negative impacts of human population density on tayra abundance. We hypothesize this relationship could be underpinned by conflict and subsequent persecution by humans, which is likely to be exacerbated in agricultural landscapes. These findings suggest that like many carnivores, the tayra may be able to adapt and even thrive in human modified landscapes given societal acceptance, thus, conservation strategies for the tayra that focus on fostering co-existence between humans and this medium-sized carnivore may contribute to its future.

Biodiversity and Conservation↗
USGS2010

A generalized watershed disturbance-invertebrate relation applicable in a range of environmental settings across the continental United States

It is widely recognized that urbanization can affect ecological conditions in aquatic systems; numerous studies have identified impervious surface cover as an indicator of urban intensity and as an index of development at the watershed, regional, and national scale. Watershed percent imperviousness, a commonly understood urban metric was used as the basis for a generalized watershed disturbance metric that, when applied in conjunction with weighted percent agriculture and percent grassland, predicted stream biotic conditions based on Ephemeroptera, Plecoptera, and Trichoptera (EPT) richness across a wide range of environmental settings. Data were collected in streams that encompassed a wide range of watershed area (4.4-1,714 km), precipitation (38-204 cm/yr), and elevation (31-2,024 m) conditions. Nevertheless the simple 3-landcover disturbance metric accounted for 58% of the variability in EPT richness based on the 261 nationwide sites. On the metropolitan area scale, relationship r ranged from 0.04 to 0.74. At disturbance values 15. Future work may incorporate watershed management practices within the disturbance metric, further increasing the management applicability of the relation. Such relations developed on a regional or metropolitan area scale are likely to be stronger than geographically generalized models; as found in these EPT richness relations. However, broad spatial models are able to provide much needed understanding in unmonitored areas and provide initial guidance for stream potential.

Portland; Salt Lake City; Denver; Dallas-Fort Wort↗
USGS1978

Aleuts, sea otters, and alternate stable state communities

Reexamination of stratified faunal components of a prehistoric Aleut midden excavated on Amchitka Island, Alaska, indicates that Aleut prey items changed dramatically during 2500 years of aboriginal occupation. Recent ecological studies in the Aleutian Islands have shown the concurrent existence of two alternate stable nearshore communities, one dominated by macroalgae, the other by epibenthic herbivores, which are respectively maintained by the presence or absence of dense sea otter populations. Thus, rather than cultural shifts in food preference, the changes in Aleut prey were probably the result of local overexploitation of sea otters by aboriginal Aleuts.

Alaska↗
USGS2014

Physiological condition of juvenile wading birds in relation to multiple landscape stressors in the Florida Everglades: effects of hydrology, prey availability, and mercury bioaccumulation

The physiological condition of juvenile birds can be influenced by multiple ecological stressors, and few studies have concurrently considered the effects of environmental contaminants in combination with ecological attributes that can influence foraging conditions and prey availability. Using three temporally distinct indices of physiological condition, we compared the physiological response of nestling great egrets ( Ardea alba ) and white ibises ( Eudocimus albus ) to changing prey availability, hydrology (water depth, recession rate), and mercury exposure in the Florida Everglades. We found that the physiological response of chicks varied between species and among environmental variables. Chick body condition (short-term index) and fecal corticosterone levels (medium-term) were influenced by wetland water depth, prey availability, region, and age, but not by mercury contamination. However, mercury exposure did influence heat shock protein 70 (HSP70) in egret chicks, indicating a longer-term physiological response to contamination. Our results indicate that the physiological condition of egret and ibis chicks were influenced by several environmental stressors, and the time frame of the effect may depend on the specialized foraging behavior of the adults provisioning the chicks.

Florida↗