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Carbon dioxide in the ocean surface: The homogeneous buffer factor

The amount of carbon dioxide that can be dissolved in surface seawater depends at least partially on the homogeneous buffer factor, which is a mathematical function of the chemical equilibrium conditions among the various dissolved inorganic species. Because these equilibria are well known, the homogeneous buffer factor is well known. Natural spatial variations depend very systematically on sea surface temperatures, and do not contribute significantly to uncertainties in the present or future carbon dioxide budget.

Science

A geological synthesis of the Precambrian shield in Madagascar

Available U–Pb geochronology of the Precambrian shield of Madagascar is summarized and integrated into a synthesis of the region’s geological history. The shield is described in terms of six geodynamic domains, from northeast to southwest, the Bemarivo, Antongil–Masora, Antananarivo, Ikalamavony, Androyan–Anosyan, and Vohibory domains. Each domain is defined by distinctive suites of metaigneous rocks and metasedimentary groups, and a unique history of Archean (∼2.5 Ga) and Proterozoic (∼1.0 Ga, ∼0.80 Ga, and ∼0.55 Ga) reworking. Superimposed within and across these domains are scores of Neoproterozoic granitic stocks and batholiths as well as kilometer long zones of steeply dipping, highly strained rocks that record the effects of Gondwana’s amalgamation and shortening in latest Neoproterozoic time (0.560–0.520 Ga). The present-day shield of Madagascar is best viewed as part of the Greater Dharwar Craton, of Archean age, to which three exotic terranes were added in Proterozoic time. The domains in Madagascar representing the Greater Dharwar Craton include the Antongil–Masora domain, a fragment of the Western Dharwar of India, and the Neoarchean Antananarivo domain (with its Tsaratanana Complex) which is broadly analogous to the Eastern Dharwar of India. In its reconstructed position, the Greater Dharwar Craton consists of a central nucleus of Paleo-Mesoarchean age (>3.1 Ga), the combined Western Dharwar and Antongil–Masora domain, flanked by mostly juvenile “granite–greenstone belts” of Neoarchean age (2.70–2.56 Ga). The age of the accretionary event that formed this craton is approximately 2.5–2.45 Ga. The three domains in Madagascar exotic to the Greater Dharwar Craton are the Androyan–Anosyan, Vohibory, and Bemarivo. The basement to the Androyan–Anosyan domain is a continental terrane of Paleoproterozoic age (2.0–1.78 Ga) that was accreted to the southern margin (present-day direction) of the Greater Dharwar Craton in pre-Stratherian time (>1.6 Ga), and rejuvenated at 1.03–0.93 Ga with the creation of the Ikalamavony domain. The Vohibory domain, an oceanic terrane of Neoproterozoic age was accreted to the Androyan–Anosyan domain in Cryogenian time (∼0.63–0.60 Ga). The Bemarivo domain of north Madagascar is a terrane of Cryogenian igneous rocks, with a cryptic Paleoproterozoic basement, that was accreted to the Greater Dharwar Craton in latest Ediacaran to earliest Cambrian time (0.53–0.51 Ga).

Journal of African Earth Sciences

Comments to Middle Miocene closure of the Central American Seaway

In a recent paper proposing an early (mid-Miocene) closure of the Central American Seaway (CAS), Montes et al. 2015 (1) disregard existing paleogeographic data that invalidate Panama as a source for zircons, and inappropriately ignore the evidence for trans-isthmian marine connections until 4-3 Ma. They also fail to cite previous work (2, 3), that had reconstructed the Central American arc already docked with South America by 12 Ma. Montes et al. 2015 (1) (Fig. 1) disregard the Atrato-San Juan sedimentary basin (3), a shallowing Oligocene to Pliocene, Pacific to Caribbean seaway (3, 4, 5). This deep graben (6) is filled with thousands of meters of Pre-Pliocene marine sediments (3, 5, 6) that now occupy a lowland between the Baudo uplift to the west and the Western Cordillera to the east. The Mande Batholith and numerous Eocene and younger volcanic rocks (4), the most proximal source of the zircons, are situated to the east of this seaway and would have shed zircons eastward towards the Cordillera Central. There is no evidence for any rivers crossing the seaway (3, 5), and thus no Panamanian source of zircons. Instead this seaway is evidence of a significant marine connection between the Pacific and Atlantic oceans into the Pliocene. The authors assume that the middle Miocene closure of the CAS effectively creates a continuous land bridge connecting North and South America and separating the Atlantic from the Pacific. They acknowledge, but then discount, marine connections across the Isthmus until 4-3 Ma even though these satisfactorily explain (Coates and Stallard, 2014 (6)) the oceanographic, molecular and Great American Biological Interchange events ignore unexplained by Montes et al. 2015. Only by conspicuously ignoring these events can they imply that the Isthmus was formed at 15-13 Ma. References 1. C. Montes et al., Middle Miocene closure of the Central American Seaway. Science 348, 226-229 (2015). 2. A. G. Coates, R. F. Stallard, How old is the Isthmus of Panama? B Mar Sci 89, 801-813 (2013). 3. H. Duque-Caro, The Choco Block in the northwestern corner of South America: Structural, tectonostratigraphic, and paleogeographic implications. Journal of South American Earth Sciences 3, 71-84 (1990). 4. H. Duque-Caro, Neogene stratigraphy, paleoceanography and paleobiogeography in northwest South America and the evolution of the Panama Seaway. Pal

Science

Future of gas hydrate research

Methane hydrates are ice‐like inclusion compounds, in which every volume of hydrate can contain as much as 180 volumes (STP) of gas.The amount of methane in natural gas hydrates is twice the total recoverable fossil fuel reserve. Because of their natural abundance in oceans and permafrost, hydrates have become an exciting national and international research issue. The movement of the gas and oil industry to ever deepening waters where hydrates occur, the compelling size and distribution of hydrate deposits, and strong international interest all support identification of crucial elements in a hydrate research program.

Eos, Earth and Space Science News

Climate change, coral loss, and the curious case of the parrotfish paradigm: Why don't marine protected areas improve reef resilience?

Scientists have advocated for local interventions, such as creating marine protected areas and implementing fishery restrictions, as ways to mitigate local stressors to limit the effects of climate change on reef-building corals. However, in a literature review, we find little empirical support for the notion of managed resilience. We outline some reasons for why marine protected areas and the protection of herbivorous fish (especially parrotfish) have had little effect on coral resilience. One key explanation is that the impacts of local stressors (e.g., pollution and fishing) are often swamped by the much greater effect of ocean warming on corals. Another is the sheer complexity (including numerous context dependencies) of the five cascading links assumed by the managed-resilience hypothesis. If reefs cannot be saved by local actions alone, then it is time to face reef degradation head-on, by directly addressing anthropogenic climate change—the root cause of global coral decline.

Annual Review of Marine Science

Location and age of foraminifer samples examined by Chevron Petroleum Company paleontologists from more than 2,500 oil test wells in California

Chevron Petroleum Company in 2001 donated an estimated 50,000 foraminifer slides, 5,000 well logs, geologic and surface locality maps, and paleontologic reports to the California Academy of Sciences and Stanford University for safekeeping, because they stopped or cut back exploration for petroleum deposits in California. The material was loaned to Earl Brabb temporarily so that information useful to the U.S. Geological Survey could be extracted. Among the estimated 5,000 well logs, more than 2,500 were printed on fragile Ozalid paper that had deteriorated by turning brown and hardening so that they could be easily damaged. These 2,516 well logs were scanned to provide a digital copy of the information. The 2,516 wells extend over an area from Eureka in Humboldt County south to the Imperial Valley and from the Pacific Ocean east to the eastern side of the Great Valley and the Los Angeles Basin. The wells are located in 410 7.5-minute quadrangle maps in 42 counties. The digital information herein preserves the data, makes the logs easily distributed to others interested in subsurface geology, and makes previously proprietary information widely available to the public for the first time.

California

Collection methods and descriptions of coral cores extracted from massive corals in Dry Tortugas National Park, Florida, U.S.A.

Cores from living coral colonies were collected from Dry Tortugas National Park, Florida, U.S.A., to obtain skeletal records of past coral growth and allow geochemical reconstruction of environmental variables during the corals’ centuries-long lifespans. The samples were collected as part of the U.S. Geological Survey Coral Reef Ecosystems Studies project ( http:/coastal.er.usgs.gov/crest ) that provides science to assist resource managers tasked with the stewardship of coral reef resources. Three colonies each of the coral species Orbicella faveolata and Siderastrea siderea were collected in May 2012 using the methods described herein and as approved under National Park Service scientific collecting permit number DRTO-2012-SCI-0001 and are cataloged under accession number DRTO-353. These coral samples can be used to retroactively construct environmental parameters, including sea-surface temperature, by measuring the elemental composition of the coral skeleton. The cores described here, and others (see http://olga.er.usgs.gov/coreviewer/ ), can be requested, on loan, for scientific study. Photographic images for each coral in its ocean environment, the coral cores as curated and slabbed, and the X-rays of the slabs can be found in an associated U.S. Geological Survey Data Release.

Florida

Nutrients, phytoplankton, zooplankton, and macrobenthos

Lower trophic levels support the prey fish on which most sport fish depend. Therefore, understanding the production potential of lower trophic levels is integral to the management of Lake Ontario’s fishery resources. Lower trophic-level productivity differs among offshore and nearshore waters. In the offshore, there is concern about the ability of the lake to support Alewife (Table 1) production due to a perceived decline in productivity of phytoplankton and zooplankton whereas, in the nearshore, there is a concern about excessive attached algal production (e.g., Cladophora) associated with higher nutrient concentrations—the oligotrophication of the offshore and the eutrophication of the nearshore (Mills et al. 2003; Holeck et al. 2008; Dove 2009; Koops et al. 2015; Stewart et al. 2016). Even though the collapse of the Alewife population in Lake Huron in 2003 (and the associated decline in the Chinook Salmon fishery) may have been precipitated by a cold winter (Dunlop and Riley 2013), Alewife had not returned to high abundances in Lake Huron as of 2014 (Roseman et al. 2015). Failure of the Alewife population to recover from collapse has been attributed to declines in lower trophic-level production (Barbiero et al. 2011; Bunnell et al. 2014; but see He et al. 2015). In Lake Michigan, concerns of a similar Alewife collapse led to a decrease in the number of Chinook Salmon stocked. If lower trophic-level production declines in Lake Ontario, a similar management action could be considered. On the other hand, in Lake Erie, which supplies most of the water in Lake Ontario, eutrophication is increasing and so are harmful algal blooms. Thus, there is also a concern that nutrient levels and algal blooms could increase in Lake Ontario, especially in the nearshore. Solutions to the two processes of concern—eutrophication in the nearshore and oligotrophication in the offshore—may be mutually exclusive. In either circumstance, fisheries management needs information on the productivity of lower trophic levels in Lake Ontario. In this chapter, we review the status of lower trophic levels in Lake Ontario with special attention to the current (2008-2013) and previous (2003-2007) reporting periods. During the two reporting periods, three whole-lake surveys of lower trophic levels were conducted: the Lower Trophic Level Assessment (LOLA) in 2003 and 2008 (Makarewicz and Howell 2012; Munawar et al. 2015b) and the Cooperative Science and Management Initiative (CSMI) in 2013. Analyses of the CSMI data are ongoing. In addition to the three one-year sources of information on lower trophic levels, several multi-year sources of information are available, including data from the surveillance program conducted since 1965 by Environment Canada (EC) (Dove 2009), monitoring conducted since 1980 by the U.S. Environmental Protection Agency’s (EPA) Great Lakes National Program Office (GLNPO) (Barbiero et al. 2014; Reavie et al. 2014), sampling for a Bioindex Program at two stations, one offshore and one in the Eastern Basin, assessments of Mysis diluviana (formerly Mysis relicta) conducted since 1980 by Fisheries and Oceans Canada (Johannsson et al. 1998, 2011) and the Ontario Ministry of Natural Resources and Forestry (OMNRF), and monitoring conducted since 1995 by the Biomonitoring Program (BMP) on the New York side of the lake (Holeck et al. 2015b). The BMP is a collaboration of the New York State Department of Environmental Conservation (DEC), U.S. Fish and Wildlife Service, U.S. Geological Survey (USGS), and Cornell University.

Special Publication

Sources of land-derived runoff to a coral reef-fringed embayment identified using geochemical tracers in nearshore sediment traps

Geochemical tracers, including Ba, Co, Th, 7 Be, 137 Cs and 210 Pb, and magnetic properties were used to characterize terrestrial runoff collected in nearshore time-series sediment traps in Hanalei Bay, Kauai, during flood and dry conditions in summer 2006, and to fingerprint possible runoff sources in the lower watershed. In combination, the tracers indicate that runoff during a flood in August could have come from cultivated taro fields bordering the lower reach of the river. Land-based runoff associated with summer floods may have a greater impact on coral reef communities in Hanalei Bay than in winter because sediment persists for several months. During dry periods, sediment carried by the Hanalei River appears to have been mobilized primarily by undercutting of low 7 Be, low 137 Cs riverbanks composed of soil weathered from tholeiitic basalt with low Ba and Co concentrations. Following a moderate rainfall event in September, high 7 Be sediment carried by the Hanalei River was probably mobilized by overland flow in the upper watershed. Ba-desorption in low-salinity coastal water limited its use to a qualitative runoff tracer in nearshore sediment. 210 Pb had limited usefulness as a terrestrial tracer in the nearshore due to a large dissolved oceanic source and scavenging onto resuspended bottom sediment. 210 Pb-scavenging does, however, illustrate the role resuspension could play in the accumulation of particle-reactive contaminants in nearshore sediment. Co and 137 Cs were not affected by desorption or geochemical scavenging and showed the greatest potential as quantitative sediment provenance indicators in material collected in nearshore sediment traps.

Estuarine, Coastal and Shelf Science

Tracking carbon from subduction to outgassing along the Aleutian-Alaska Volcanic Arc

Subduction transports volatiles between Earth’s mantle, crust, and atmosphere, ultimately creating a habitable Earth. We use isotopes to track carbon from subduction to outgassing along the Aleutian-Alaska Arc. We find substantial along-strike variations in the isotopic composition of volcanic gases, explained by different recycling efficiencies of subducting carbon to the atmosphere via arc volcanism and modulated by subduction character. Fast and cool subduction facilitates recycling of ~43 to 61% sediment-derived organic carbon to the atmosphere through degassing of central Aleutian volcanoes, while slow and warm subduction favors forearc sediment removal, leading to recycling of ~6 to 9% altered oceanic crust carbon to the atmosphere through degassing of western Aleutian volcanoes. These results indicate that less carbon is returned to the deep mantle than previously thought and that subducting organic carbon is not a reliable atmospheric carbon sink over subduction time scales.

Alaska

Development of the Circum-Antarctic Current

Deep-sea drilling in the Southern Ocean south of Australia and New Zealand shows that the Circum-Antarctic Current developed about 30 million years ago in the middle to late Oligocene when final separation occurred between Antarctica and the continental South Tasman Rise. Australia had commenced drifting northward from Antarctica 20 million years before this.

Science

Long-term perspectives on giant earthquakes and tsunamis at subduction zones

Histories of earthquakes and tsunamis, inferred from geological evidence, aid in anticipating future catastrophes. This natural warning system now influences building codes and tsunami planning in the United States, Canada, and Japan, particularly where geology demonstrates the past occurrence of earthquakes and tsunamis larger than those known from written and instrumental records. Under favorable circumstances, paleoseismology can thus provide long-term advisories of unusually large tsunamis. The extraordinary Indian Ocean tsunami of 2004 resulted from a fault rupture more than 1000 km in length that included and dwarfed fault patches that had broken historically during lesser shocks. Such variation in rupture mode, known from written history at a few subduction zones, is also characteristic of earthquake histories inferred from geology on the Pacific Rim. Copyright ?? 2007 by Annual Reviews. All rights reserved.

Annual Review of Earth and Planetary Sciences

Sources, distributions and dynamics of dissolved organic matter in the Canada and Makarov Basins

A comprehensive survey of dissolved organic carbon (DOC) and chromophoric dissolved organic matter (CDOM) was conducted in the Canada and Makarov Basins and adjacent seas during 2010–2012 to investigate the dynamics of dissolved organic matter (DOM) in the Arctic Ocean. Sources and distributions of DOM in polar surface waters were very heterogeneous and closely linked to hydrological conditions. Canada Basin surface waters had relatively low DOC concentrations (69 ± 6 μmol L −1 ), CDOM absorption ( a 325 : 0.32 ± 0.07 m −1 ) and CDOM-derived lignin phenols (3 ± 0.4 nmol L −1 ), and high spectral slope values ( S 275–295 : 31.7 ± 2.3 μm −1 ), indicating minor terrigenous inputs and evidence of photochemical alteration in the Beaufort Gyre. By contrast, surface waters of the Makarov Basin had elevated DOC (108 ± 9 μmol L −1 ) and lignin phenol concentrations (15 ± 3 nmol L −1 ), high a 325 values (1.36 ± 0.18 m −1 ), and low S 275–295 values (22.8 ± 0.8 μm −1 ), indicating pronounced Siberian river inputs associated with the Transpolar Drift and minor photochemical alteration. Observations near the Mendeleev Plain suggested limited interactions of the Transpolar Drift with Canada Basin waters, a scenario favoring export of Arctic DOM to the North Atlantic. The influence of sea-ice melt on DOM was region-dependent, resulting in an increase (Beaufort Sea), a decrease (Bering-Chukchi Seas), and negligible change (deep basins) in surface DOC concentrations and a 325 values. Halocline structures differed between basins, but the Canada Basin upper halocline and Makarov Basin halocline were comparable in their average DOC (65–70 μmol L −1 ) and lignin phenol concentrations (3–4 nmol L −1 ) and S 275–295 values (22.9–23.7 μm −1 ). Deep-water DOC concentrations decreased by 6–8 μmol L −1 with increasing depth, water mass age, nutrient concentrations, and apparent oxygen utilization. Maximal estimates of DOC degradation rates (0.036–0.039 μmol L −1 yr −1 ) in the deep Arctic were lower than those in other ocean basins, possibly due to low water temperatures. DOC concentrations in bottom waters (>2500 m; 46 ± 2 μmol L −1 ) of the Canada and Makarov Basins were slightly lower than those reported for deep waters of the Eurasian Basin and Nordic Seas. Elevated a 325 values (by 10–20%) were observed near the seafloor, indicating biological activity in Arctic basin sediments.

Frontiers in Marine Science

Mercury in Pacific bluefin tuna (Thunnus orientalis):bioaccumulation and trans-Pacific Ocean migration

Pacific bluefin tuna (Thunnus orientalis) have the largest home range of any tuna species and are well known for the capacity to make transoceanic migrations. We report the measurement of mercury (Hg) concentrations in wild Pacific bluefin tuna (PBFT), the first reported with known size-of-fish and capture location. The results indicate juvenile PBFT that are recently arrived in the California Current from the western Pacific Ocean have significantly higher Hg concentrations in white muscle (0.51 ug/g wet mass, wm) than PBFT of longer California Current residency (0.41 ug/g wm). These new arrivals are also higher in Hg concentration than PBFT in farm pens (0.43 ug/g wm) that were captured on arrival in the California Current and raised in pens on locally derived feed. Analysis by direct Hg analyzer and attention to Hg by tissue type and location on the fish allowed precise comparisons of mercury among wild and captive fish populations. Analysis of migration and nearshore residency, determined through extensive archival tagging, bioaccumulation models, trophic investigations, and potential coastal sources of methylmercury, indicates Hg bioaccumulation is likely greater for PBFT juvenile habitats in the western Pacific Ocean (East China Sea, Yellow Sea) than in the eastern Pacific Ocean (California Current). Differential bioaccumulation may be a trophic effect or reflect methylmercury availability, with potential sources for coastal China (large hypoxic continental shelf receiving discharge of three large rivers, and island-arc volcanism) different from those for coastal Baja California (small continental shelf, no large rivers, spreading-center volcanism).

Canadian Journal of Fisheries and Aquatic Sciences

Coastal landforms and processes at the Cape Cod National Seashore, Massachusetts—A primer

Anyone who spends more than a few days on Cape Cod (the Cape) quickly becomes a coastal geologist, quickly learning the rhythms of daily tides and the seasonal cycles of beaches growing and being swept away by storms; swimmers and surfers track how the breakers appear, and dog-walkers notice the hard-packed sand blanketed overnight by an airy layer that leaves deep labored tracks. Careful observers whose paths wander to the ocean’s edge will observe many of the landforms and coastal processes described in this book and if we have done our job well, the stories told here will seem familiar. Watchful experience brings insights; indeed, this is how scientists and perhaps how artists work, describing patterns that explain and predict. When is the next high tide? What will the winter bring? Where do we build, fish, swim? How do wind and waves offshore in the North Atlantic help arrange the plants and dunes and hollows on the beach? And most of all, as human animals drawn to live and play on the edge of the ocean, how do we get the benefits of this complex natural system of geology and biology? How do we affect coastal processes; how is the coast changing now and how is the coast likely to change in years ahead with climate warming and climate change? This book is about the highly dynamic coastal landforms of Cape Cod—the beaches, bluffs, spits, dunes, barrier beaches, estuaries, and salt marshes. What they are, why they are where they are, how they behave with respect to the greater Cape Cod coastal system—how the landforms respond to day-to-day and long-term geologic processes, such as waves and currents, change in sediment transport, relative sea-level rise, and meteorological processes such as hurricanes, nor’easters, and cold front passages. It is also about how the landforms got to be where they are and the way they are and where they are headed in the near future with the predicted effects of global climate warming and change. Our objective is to provide a single source of understandable and readable scientific information for those who live, play, and work on outer Cape Cod and at the Cape Cod National Seashore, as well as to provide an introduction to Cape Cod’s coastal landforms for anyone with an interest in Earth science and nature who wants a better understanding of coastal systems and processes. Basic to an understanding of coastal landforms is the fact that they work together—they interact—as elements of many systems, and therefore our ultimate concern is not the individual landform itself but rather the geologic systems that make up Cape Cod and the Cape Cod National Seashore. Much of this discussion can be applied as well to Nantucket, Martha’s Vineyard, and other coastal regions. The coast of outer Cape Cod, about 15,000 years old and about 30 miles (mi; 50 kilometers [km]) long, is but a tiny piece of the global Earth system that operates within a much larger realm of space and time. Cape Cod’s coastal landforms are temporary holding patterns within a continual interplay of land, sea, atmosphere, climate, ice, and life, including a variety of human activities that both affect and are affected by these processes. These interactions produce the landforms, and the landforms alter the interactions. The resulting landforms provide habitats for a wide variety of coastal plants and animals. The habitats along with their inhabitants and the interacting environmental factors controlling them constitute the Cape’s complex and varied ecosystems. But for now, we are here to enjoy it. We welcome you to delight and wonder at the perpetually changing handshake between the ocean and shore at New England’s Great Beach.

Massachusetts

A 107-year-old coral from Florida Bay: barometer of natural and man- induced catastrophes?

The 107-yr growth history of a massive coral Solenastrea bournoni from Florida Bay was reconstructed with X-ray imagery from a single 4 in. diameter (10 cm) core that penetrated the exact epicenter of the 95.3 cm high colony. Growth increments totalled 952.9 mm, averaging 8.9 mm/yr over the life of the coral. Growth rate trends in the Florida Bay coral were compared to those in a Montastraea annularis of similar age from a nearby patch reef on the Atlantic Ocean side of the Florida Keys. It was concluded that growth rate, at least in these specimens, is a questionable indicator of past hurricanes and freezes. There does appear to be, however, a possible cause-and-effect relationship between major man-induced environmental perturbations and a prolonged reduction in growth rate in each coral's growth record. -from Authors

Bulletin of Marine Science

Scanning electron microscope studies of some early Miocene diatoms from the equatorial Pacific Ocean with descriptions of two new species, Actinocyclus jouseae Barron and Actinocyclus nigriniae Barron

Scanning electron microscope (SEM) and light microscope (LM) studies are used to propose and describe two new species, Actinocyclus jouseae Barron , sp. nov. and Actinocyclus nigriniae Barron , sp. nov. from lower Miocene sediments from equatorial Pacific ODP Site 1219. Parallel SEM and LM studies reveal that Thalassiosira bukryi Barron should be transferred to Azpeitia and suggest that Actinocyclus barronii Radionova is likely to be a variety of A. radionovae Barron

Proceedings of the California Academy of Sciences

Influence of hydrologic processes on reproduction of the introduced bivalve Potamocorbula amurensis in northern San Francisco Bay, California

Monthly censusing of reproductive condition of the Asian clam Potamocorbula amurensis at four sites in northern San Francisco Bay over a 9-yr period revealed year-to-year differences in local reproductive activity that are associated with patterns of hydrologic variability. Between 1989 and 1992, Northern California experienced a drought, whereas the period between 1993 and 1998 was marked by a mix of wet and dry years. We took advantage of the extreme year-to-year differences to examine reproductive responses to river inflow patterns. Populations of P. amurensis at the upstream sites in Suisun Bay and Carquinez Strait were more reproductively active during wet years than dry years. Conversely, at the downstream site in San Pablo Bay, the population was more reproductively active during dry years than wet years. We suggest that the different reproductive patterns observed reflect the clam's response to different sources of food. During wet years, organic matter from the rivers augments food supplies in Suisun Bay. During dry years, when inflow into the San Francisco Bay Estuary from the rivers is reduced, water transported from the adjacent ocean into the estuary as far as San Pablo Bay provides a supplemental food supply for the local production. The populations take advantage of these spatially distinct food supplies by initiating and maintaining local reproductive activity. We conclude that the ability of P. amurensis to consume and use various types of food to regulate its reproductive activity is part of the reason for its success as an invasive species.

California