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At least 433 records · Page 24Linked to original sources

Conceptual data modeling of wildlife response indicators to ecosystem change in the Arctic

Large research studies are often challenged to effectively expose and document the types of information being collected and the reasons for data collection across what are often a diverse cadre of investigators of differing disciplines. We applied concepts from the field of information or data modeling to the U.S. Geological Survey (USGS) Changing Arctic Ecosystems (CAE) initiative to prototype an application of information modeling. The USGS CAE initiative is collecting information from marine and terrestrial environments in Alaska to identify and understand the links between rapid physical changes in the Arctic and response of wildlife populations to these ecosystem changes. An associated need is to understand how data collection strategies are informing the overall science initiative and facilitating communication of those strategies to a wide audience. We explored the use of conceptual data modeling to provide a method by which to document, describe, and visually communicate both enterprise and study level data; provide a simple means to analyze commonalities and differences in data acquisition strategies between studies; and provide a tool for discussing those strategies among researchers and managers.

Alaska

Annotated list of plants of Arctic Slope of Alaska

The accompanying 5-page list of 439 higher plants was compiled as part of a manuscript to be included in a series of professional papers on Naval Petroleum Reserve No. 4 and adjoining areas. This check list describing the distribution, abundance, flowering period, plant growth on various rook types, and other habitat relationships, is based primarily on extensive collections of arctic plants by the author and other members of the Survey. In 1948 and 1949 the Arctic Institute of North America sponsored these studies, which were done primarily out of Survey field camps. Analysis of these collections which are probably the most extensive and complete of any from northern Alaska, was done at the University of Minnesota.

Alaska

Morphology, sedimentation, and seismic characteristics of an arctic beach, Nome, Alaska - with economic significances

Arctic beaches exhibit characteristics that do not occur on beaches in more temperate zones. In the summer of 1967 morphological, sedimentological, and seismic refraction studies were made on an arctic beach near Nome, Alaska, in order to better define these distinguishing characteristics. Several distinct microrelief structures are developed during ice break-up and the melting of ice and snow on the beaches including ice-pushed ridges, kaimoo ridge and deposits, sea-ice kettles, sea- ice sand and sea-ice gravel cones, micro-outwash deposits, and microdeltaic deposits. Sedimentological studies showed that sediment distribution along the beach is random with no increasing or decreasing median diameters, or sorting coefficients, or sand-gravel percentages, either along the strandline or along the beach profile. This random grain size distribution is attributed to the relatively low energy along the coast of the Seward Peninsula. Further, the beaches are insulated from the elements by ice for about 6 months out of the year and the beach sediments are disturbed annually by ice-push. Longshore drift appeared to vary from east to west during the summer of 1967 with a net transport to the east. The seismic refraction study on the Nome beach indicated that internal stratigraphy of the overburden could be interpreted and correlated with known onshore and offshore stratigraphic units. Bedrock was well defined in the seismograms and exhibited a gently undulating topography with several buried channels that may be sites of gold placer deposits.

Alaska

Stamukhi shoals of the Arctic - some observations from the Beaufort Sea

A number of linear shoals, representing pronounced topographic anomalies on the surface of the Arctic shelf, have been studied in the Prudhoe Bay area. These shoals have been referred to in several previous studies. Based on seismic reflection records, Reimnitz et al., (1972a), stated that the shoals are constructional features younger than the post-Wisconsin transgression. Large chunks of grounded ice have frequently been seen on the linear shoals of the inner shelf. These ice chunks form barriers parallel to the shore (Reimnitz, et al. 1972b). Reimnitz and Barnes (1974) considered the lack of gravel concentrations on the shoals to be evidence that the stream of pack ice drifting past northern Alaska carries very little gravel today. Lewellen (1977) referred to the linear shoals as submerged barrier islands, while Reimnitz, et al., (1977b) pointed out that, although similar in shape to barrier islands, the linear shoals are very different in composition and do not appear to represent drowned harrier islands. They show that the shoals localize the formation of major shear and pressure events in the ice, which in turn cause the formation of linear belts of deformed and grounded ice. Today the shoals appear to be migrating under the influence of ice-bottom interaction, and indeed may have formed in response to ice-bottom interaction within the "stamukhi zone". The shoals migrate rather slowly and retain their shapes over periods of 25 years, yet control the location and stabilize the outer edge of the floating fast ice zone, and provide shelter for the inner shelf and coast. Reimnitz et al. (1977b) surmised that similar artificial structures might be used to modify the ice environment on the arctic shelf.

Alaska

Oil and gas resources of the Arctic Alaska Petroleum Province

The Arctic Alaska Petroleum Province, encompassing all the lands and adjacent Continental Shelf areas north of the Brooks Range-Herald arch, is one of the most petroleum-productive areas in the United States, having produced about 15 billion bbl of oil. Seven unitized oil fields currently contribute to production, and three additional oil fields have been unitized but are not yet producing. Most known petroleum accumulations involve structural or combination structural-stratigraphic traps related to closure along the Barrow arch, a regional basement high, which has focused regional hydrocarbon migration since Early Cretaceous time. Several oil accumulations in stratigraphic traps have been developed in recent years. In addition to three small gas fields producing for local consumption, more than 20 additional oil and gas discoveries remain undeveloped. This geologically complex region includes prospective strata within passive-margin, rift, and foreland-basin sequences. Oil and gas were generated from multiple source rocks throughout the region. Although some reservoired oils appear to be derived from a single source rock, evidence for significant mixing of hydrocarbons from multiple source rocks indicates a composite petroleum system. Both extensional and contractional tectonic structures provide ample exploration targets, and recent emphasis on stratigraphic traps has demonstrated a significant resource potential in shelf and turbidite sequences of Jurassic through Tertiary age. Recent estimates of the total mean volume of undiscovered resources in the Arctic Alaska Petroleum Province by the U.S. Geological Survey and U.S. Minerals Management Service are more than 50 billion bbl of oil and natural-gas liquids and 227 trillion ft3 of gas, distributed approximately equally between Federal offshore and combined onshore and State offshore areas.

Alaska

Arctic Alaska’s Lower Cretaceous (Hauterivian and Barremian) mudstone succession—Linking lithofacies, texture, and geochemistry to marine processes

We present new images and descriptions of the lithofacies and organic facies of the pebble shale unit and lower part of the Hue Shale (Lower Cretaceous) of Arctic Alaska at a high magnification that illustrates their textural characteristics. Our aims were to describe and determine the distribution of facies in these petroleum source rocks and to identify the processes that formed them. We sampled at high-resolution and applied new petrographic techniques combined with scanning electron microscopy and geochemical analyses to samples collected from three widely spaced sections—located in exposures along the Canning River and continuous core from the Mikkelsen Bay State 1 and Orion 1 wells. Results from these three locations indicate that this succession consists primarily of clay-rich mudstones that are variously silt- or sand-bearing and clay-dominated mudstones that exhibit mainly relict, 2–5 millimeter thick bedding and common but variable microbioturbation, rare macrobioturbation, and common fabrics of pelleted clay and silt. These mudstones contain rare, poorly sorted, silt-rich basal laminae that are often discontinuous and have wavy, sharp bases and crude upward fining. In addition, mud-supported, outsized clasts (dropstones) of fine sand to pebble size are present throughout the succession as isolated clasts or in clusters. We interpret these textures and much of this succession to result from intermittent deposition by suspension settling from melting seasonal sea ice—sometimes sediment-laden—and associated primary productivity. Overall, this mudstone succession fines and deepens upward from the pebble shale unit into the Hue Shale. In the Hue Shale of the Orion well, however, different processes intermittently deposited thin, discrete intervals of coarser sediment that probably represent deposition from density currents. Also in the Hue Shale of the Orion well, several thicker sandstone and silt-dominated mudstone units with discordant, scoured bases and cut and fill structures represent erosion during higher energy events such as major storms. Other lithofacies within the succession are graded tuffs/bentonites and tuffaceous/bentonitic mudstones from episodic volcanic ash falls; these are abundant in the Hue Shale, and very rare in the pebble shale unit of the two wells. Organic-carbon rich strata associated with volcanic ash intervals of the pebble shale unit and Hue Shale in the Mikkelsen 1 well have some of the best petroleum source rock potential determined for this succession. Authigenic pyrite and carbonate-cement-dominated mudstone are also present in both units of all three sections. The carbonate-cemented units indicate breaks in sedimentation and are common in the Hue Shale and in sections of the pebble shale unit interpreted to be more distal, such as along the Canning River. Our results document the variation in facies and textures of the Hauterivian and Barremian Lower Cretaceous mudstone succession of Arctic Alaska. Comparison of these characteristics to the products of modern processes on the North Slope of Alaska, in the Beaufort Sea, and elsewhere suggest that this succession formed primarily from depositional processes related to seasonal sea ice with intermittent fluvial-sourced sediment deposited by density currents and episodic erosion and reworking by storms and other currents.

Alaska

Upper Cretaceous and Lower Jurassic strata in shallow cores on the Chukchi Shelf, Arctic Alaska

Shallow cores collected in the 1980s on the Chukchi Shelf of western Arctic Alaska sampled pre-Cenozoic strata whose presence, age, and character are poorly known across the region. Five cores from the Herald Arch foreland contain Cenomanian to Coniacian strata, as documented by biostratigraphy, geochronology, and thermochronology. Shallow seismic reflection data collected during the 1970s and 1980s show that these Upper Cretaceous strata are truncated near the seafloor by subtle angular unconformities, including the Paleogene mid-Brookian unconformity in one core and the Pliocene-Pleistocene unconformity in four cores. Sedimentary structures and lithofacies suggest that Upper Cretaceous strata were deposited in a low accommodation setting that ranged from low-lying coastal plain (nonmarine) to muddy, shallow-marine environments near shore. These observations, together with sparse evidence from the adjacent western North Slope, suggest that Upper Cretaceous strata likely were deposited across all of Arctic Alaska. A sixth core from the Herald Arch contains lower Toarcian marine strata, indicated by biostratigraphy, truncated by a Neogene or younger unconformity. These Lower Jurassic strata evidently were deposited south of the arch, buried structurally to high levels of thermal maturity during the Early Cretaceous, and uplifted on the Herald thrust-fault system during the mid to Late Cretaceous. These interpretations are based on regional stratigraphy and apatite fission-track data reported in a complementary report and are corroborated by the presence of recycled palynomorphs of Early Jurassic age and high thermal maturity found in Upper Cretaceous strata in two of the foreland cores. This dataset provides evidence that uplift and exhumation of the Herald thrust belt provided sediment to the foreland during the Late Cretaceous.

Professional Paper

Methodology for assessment of undiscovered oil and gas resources for the 2008 Circum-Arctic Resource Appraisal

The methodological procedures used in the geologic assessments of the 2008 Circum-Arctic Resource Appraisal (CARA) were based largely on the methodology developed for the 2000 U.S. Geological Survey World Petroleum Assessment. The main variables were probability distributions for numbers and sizes of undiscovered accumulations with an associated risk of occurrence. The CARA methodology expanded on the previous methodology in providing additional tools and procedures more applicable to the many Arctic basins that have little or no exploration history. Most importantly, geologic analogs from a database constructed for this study were used in many of the assessments to constrain numbers and sizes of undiscovered oil and gas accumulations.

Professional Paper

Surficial geologic map of the Gates of the Arctic National Park and Preserve, Alaska

The Gates of the Arctic National Park and Preserve (GAAR) is centered over the central Brooks Range of northern Alaska. To the west, it abuts the Noatak National Preserve; its eastern boundary is the transportation corridor occupied by the Dalton Highway and the Alyeska Pipeline. The GAAR extends northward beyond the northern flank of the Brooks Range into the southern Arctic Foothills. Its southern boundary lies beyond the south flank of the Brooks Range within foothills and depositional basins of interior Alaska. The accompanying surficial geologic map covers all of the GAAR with the addition of a 10-km (6.2-mi) belt or "buffer zone" beyond its boundaries. A narrower (5-km) buffer zone is employed where the GAAR extends farthest north and south of the Brooks Range, in the north-central and southwestern parts of the map area, respectively. The surfical geologic map incorporates parts of ten surficial geologic maps previously published at 1:250,000 scale. In addition, a small part of the buffer zone mapped in the southwest corner of the map area was compiled from unpublished surficial geologic mapping of the Shungnak 1:250,000-scale quadrangle. Each of those individual maps was developed from (1) aerial and surface observations of morphology and composition of unconsolidated deposits, (2) tracing the distribution and interrelation of terraces, abandoned meltwater channels, moraines, abandoned lake beds, and other landforms, (3) stratigraphic study of exposures along lake shores and river bluffs, (4) examination of sediments and soil profiles in auger borings and test pits, and exposed in roadcuts and placer workings, and (5) analysis of previously published geologic maps and reports. The map units used for those maps and employed in the present compilation are defined on the basis of their physical character, genesis, and age. Relative and absolute ages of the map units were determined from their geographic locations and from their stratigraphic positions and radiocarbon ages.

Alaska

Aufeis accumulations in stream bottoms in arctic and subarctic environments as a possible indicator of geologic structure: Chapter F in Recent U.S. Geological Survey studies in the Tintina Gold Province, Alaska, United States, and Yukon, Canada--results of a 5-year project

Thick accumulations of ice, called “aufeis,” form during winter along stream and river valleys in arctic and subarctic regions. In high-gradient alpine streams, aufeis forms mostly as a result of ground-water discharge into the stream channel. The ice occludes this discharge, perturbing the steady-state condition, and causing an incremental rise in the local water table until discharge occurs higher on the stream bank above the previously formed ice. Successive freezing of onlapping ice layers can lead to aufeis accumulations several meters thick. The location and extent of aufeis in high-gradient streams may be useful to relate local hydrology to geologic structure. In the Goodpaster River basin study area, mineral deposits are known to occur, the location of which may be structurally controlled. Therefore, a more thorough understanding of regional geologic structures may facilitate a more detailed understanding of the genesis of the mineral deposits. Extensive aufeis was observed during visits to the Goodpaster River basin in east-central Alaska during 1999, 2001, and 2002. Seeps from the sides of the valleys caused ice to build up, giving the ice surface a concave-upward shape perpendicular to the stream direction. This concavity is evidence for ground-water discharge along the length of the aufeis, as opposed to discharge from a single upstream point. During thaw, streamflow is commonly observed out of the normal channel, evidence that occlusion of the channel (and shallow sediments) by ice is a viable mechanism for causing the water table to rise. The thickest (>3 meters) and most extensive aufeis (100’s of meters to kilometers along valleys) coincided with locations of laterally extensive (>5 kilometers) mapped high-angle brittle fault zones, suggesting that the fault zones are hydraulically conductive. Additional evidence of water flow is provided by observed changes in stream-water chemistry in reaches in which aufeis forms, despite a lack of surface tributaries. Minor or no aufeis was observed in many other drainage valleys where no laterally extensive structures have been mapped, implying that aufeis formation results from more than a topographic effect or discharge from bank storage. Thus, the presence of thick, laterally extensive aufeis in highgradient streams may be a useful aid to geologic structural mapping in arctic and subarctic climates.

Alaska;Yukon

Distribution and biophysical processes of beaded streams in Arctic permafrost landscapes

Beaded streams are widespread in permafrost regions and are considered a common thermokarst landform. However, little is known about their distribution, how and under what conditions they form, and how their intriguing morphology translates to ecosystem functions and habitat. Here we report on a Circum-Arctic survey of beaded streams and a watershed-scale analysis in northern Alaska using remote sensing and field studies. We mapped over 400 channel networks with beaded morphology throughout the continuous permafrost zone of northern Alaska, Canada, and Russia and found the highest abundance associated with medium- to high- ground ice content permafrost in moderately sloping terrain. In the Fish Creek watershed, beaded streams accounted for half of the drainage density, occurring primarily as low-order channels initiating from lakes and drained lake basins. Beaded streams predictably transition to alluvial channels with increasing drainage area and decreasing channel slope, although this transition is modified by local controls on water and sediment delivery. Comparison of one beaded channel using repeat photography between 1948 and 2013 indicate a relatively stable landform and 14C dating of basal sediments suggest channel formation may be as early as the Pleistocene-Holocene transition. Contemporary processes, such as deep snow accumulation in riparian zones effectively insulates channel ice and allows for perennial liquid water below most beaded stream pools. Because of this, mean annual temperatures in pool beds are greater than 2°C, leading to the development of perennial thaw bulbs or taliks underlying these thermokarst features. In the summer, some pools thermally stratify, which reduces permafrost thaw and maintains coldwater habitats. Snowmelt generated peak-flows decrease rapidly by two or more orders of magnitude to summer low flows with slow reach-scale velocity distributions ranging from 0.1 to 0.01 m/s, yet channel runs still move water rapidly between pools. The repeating spatial pattern associated with beaded stream morphology and hydrological dynamics may provide abundant and optimal foraging habitat for fish. Thus, beaded streams may create important ecosystem functions and habitat in many permafrost landscapes and their distribution and dynamics are only beginning to be recognized in Arctic research.

Alaska

Reviews and syntheses: Effects of permafrost thaw on Arctic aquatic ecosystems

The Arctic is a water-rich region, with freshwater systems covering about 16 % of the northern permafrost landscape. Permafrost thaw creates new freshwater ecosystems, while at the same time modifying the existing lakes, streams, and rivers that are impacted by thaw. Here, we describe the current state of knowledge regarding how permafrost thaw affects lentic (still) and lotic (moving) systems, exploring the effects of both thermokarst (thawing and collapse of ice-rich permafrost) and deepening of the active layer (the surface soil layer that thaws and refreezes each year). Within thermokarst, we further differentiate between the effects of thermokarst in lowland areas vs. that on hillslopes. For almost all of the processes that we explore, the effects of thaw vary regionally, and between lake and stream systems. Much of this regional variation is caused by differences in ground ice content, topography, soil type, and permafrost coverage. Together, these modifying factors determine (i) the degree to which permafrost thaw manifests as thermokarst, (ii) whether thermokarst leads to slumping or the formation of thermokarst lakes, and (iii) the manner in which constituent delivery to freshwater systems is altered by thaw. Differences in thaw-enabled constituent delivery can be considerable, with these modifying factors determining, for example, the balance between delivery of particulate vs. dissolved constituents, and inorganic vs. organic materials. Changes in the composition of thaw-impacted waters, coupled with changes in lake morphology, can strongly affect the physical and optical properties of thermokarst lakes. The ecology of thaw-impacted lakes and streams is also likely to change; these systems have unique microbiological communities, and show differences in respiration, primary production, and food web structure that are largely driven by differences in sediment, dissolved organic matter, and nutrient delivery. The degree to which thaw enables the delivery of dissolved vs. particulate organic matter, coupled with the composition of that organic matter and the morphology and stratification characteristics of recipient systems will play an important role in determining the balance between the release of organic matter as greenhouse gases (CO 2 and CH 4 ), its burial in sediments, and its loss downstream. The magnitude of thaw impacts on northern aquatic ecosystems is increasing, as is the prevalence of thaw-impacted lakes and streams. There is therefore an urgent need to quantify how permafrost thaw is affecting aquatic ecosystems across diverse Arctic landscapes, and the implications of this change for further climate warming.

Biogeosciences

Multi-year, spatially extensive, watershed-scale synoptic stream chemistry and water quality conditions for six permafrost-underlain Arctic watersheds

Repeated sampling of spatially distributed river chemistry can be used to assess the location, scale, and persistence of carbon and nutrient contributions to watershed exports. Here, we provide a comprehensive set of water chemistry measurements and ecohydrological metrics describing the biogeochemical conditions of permafrost-affected Arctic watersheds. These data were collected in watershed-wide synoptic campaigns in six stream networks across northern Alaska. Three watersheds are associated with the Arctic Long-Term Ecological Research site at Toolik Field Station (TFS), which were sampled seasonally each June and August from 2016 to 2018. Three watersheds were associated with the National Park Service (NPS) of Alaska and the U.S. Geological Survey (USGS) and were sampled annually from 2015 to 2019. Extensive water chemistry characterization included carbon species, dissolved nutrients, and major ions. The objective of the sampling designs and data acquisition was to characterize terrestrial–aquatic linkages and processing of material in stream networks. The data allow estimation of novel ecohydrological metrics that describe the dominant location, scale, and overall persistence of ecosystem processes in continuous permafrost. These metrics are (1) subcatchment leverage, (2) variance collapse, and (3) spatial persistence. Raw data are available at the National Park Service Integrated Resource Management Applications portal (O'Donnell et al., 2021, https://doi.org/10.5066/P9SBK2DZ ) and within the Environmental Data Initiative (Abbott, 2021, https://doi.org/10.6073/pasta/258a44fb9055163dd4dd4371b9dce945 ).

Alaska

Seasonal thaw settlement at drained thermokarst lake basins, Arctic Alaska

Drained thermokarst lake basins (DTLBs) are ubiquitous landforms on Arctic tundra lowland. Their dynamic states are seldom investigated, despite their importance for landscape stability, hydrology, nutrient fluxes, and carbon cycling. Here we report results based on high-resolution Interferometric Synthetic Aperture Radar (InSAR) measurements using space-borne data for a study area located on the North Slope of Alaska near Prudhoe Bay, where we focus on the seasonal thaw settlement within DTLBs, averaged between 2006 and 2010. The majority (14) of the 18 DTLBs in the study area exhibited seasonal thaw settlement of 3–4 cm. However, four of the DTLBs examined exceeded 4 cm of thaw settlement, with one basin experiencing up to 12 cm. Combining the InSAR observations with the in situ active layer thickness measured using ground penetrating radar and mechanical probing, we calculated thaw strain, an index of thaw settlement strength along a transect across the basin that underwent large thaw settlement. We found thaw strains of 10–35% at the basin center, suggesting the seasonal melting of ground ice as a possible mechanism for the large settlement. These findings emphasize the dynamic nature of permafrost landforms, demonstrate the capability of the InSAR technique to remotely monitor surface deformation of individual DTLBs, and illustrate the combination of ground-based and remote sensing observations to estimate thaw strain. Our study highlights the need for better description of the spatial heterogeneity of landscape-scale processes for regional assessment of surface dynamics on Arctic coastal lowlands.

Alaska

InSAR detects possible thaw settlement in the Alaskan Arctic Coastal Plain

Satellite interferometric synthetic aperture radar (InSAR) has proven to be an effective tool for monitoring surface deformation from volcanoes, earthquakes, landslides, and groundwater withdrawal. This paper seeks to expand the list of applications of InSAR data to include monitoring subsidence possibly associated with thaw settlement over the Alaskan Arctic Coastal Plain. To test our hypothesis that InSAR data are sufficiently sensitive to detect subsidence associated with thaw settlement, we acquired all Japanese Earth Resources Satellite-1 (JERS-1) L-band data available for the summers of 1996, 1997, and 1998 over two sites on the Alaska North Slope. The least amount of subsidence for both study sites was detected in the interferograms covering the summer of 1996 (2-3 cm), interferograms from 1997 and 1998 revealed that about 3 cm of subsidence occurred at the northern Cache One Lake site, and about 5 cm of subsidence was detected at the southern Kaparuk River site. These preliminary results illustrate the capacity of the L-band (24 cm) wavelength JERS-1 radar data to penetrate the short Arctic vegetation to monitor subsidence possibly associated with thaw settlement of the active layer and (or) other hydrologic changes over relatively large areas.

Canadian Journal of Remote Sensing

Morphotypical and geochemical variations of planktic foraminiferal species in Siberian and Central Arctic Ocean core tops

In this work, we utilize a transect of core top, mid- to late Holocene, sediments from the Eastern Siberian Sea to the central Arctic Ocean, spanning gradients in upper-ocean water column properties, to examine regional planktic foraminiferal species abundances and geochemistry. We present species- and morphotype-specific foraminiferal assemblages at these sites and stable isotope analyses of neogloboquadrinids. We find little variation in planktic species populations, and only small variations in N. pachyderma morphotype distributions, between sites. Spatial averages of N. pachyderma morphotype and N. incompta δ 18 O values show no significant differences, suggesting a similar calcification depth for all morphotypes of N. pachyderma and N. incompta across our sites, which we estimate to be between ∼ 50–150 m. Values of δ 18 O of a group of unencrusted specimens delineate a shallower calcification habitat. Neogloboquadrina pachyderma-2 Mg/Ca values yield temperatures outside the range of observations using available calibration equations, pointing toward the need for more Arctic-specific Mg/Ca-temperature calibrations.

Journal of Foraminiferal Research

Malocclusion in the jaws of captive bred Arctic wolves

Similar abnormalities in the skulls of captive Arctic Wolves (Canis lupus arctos) and a wild Arctic wolf found dead on Ellesmere Island, Canada, in 1986 are described. The malocclusion is likely to be recessively inherited and would be expressed more frequently in association with increased levels of inbreeding. A re-shaping of the skulls may have occurred due to the effects of the malocclusive trait. The Ellesmere skull was short and wide in comparison to the captive skulls which were long and narrow. The focus of effect was in a foreshortening of the rostrum and the resulting shortened toothrow.

Canadian Field-Naturalist

Summer movements and behavior of an Arctic wolf, Canis lupus, pack without pups

A pupless arctic Wolf pack (two adults, three yearlings) studied 5-30 July 1993 on Ellesmere Island, Northwest Territories, traveled nomadically around an area >381 km2 but the alpha pair sometimes left the yearlings at a rendezvous site. All pack members hunted Arctic hares. The alpha pair sometimes fed the yearlings, the alpha male doing so more than the alpha female.

Canadian Field-Naturalist