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At least 37 records · Page 2Linked to original sources

Response of Pacific walruses to disturbances from capture and handling activities at a haul-out in Bristol Bay, Alaska

Observations were made on hems of the Pacific walrus (Odobenus rosmarus divergens) to study their response during the capturing and handling of adult males in summer 1995 at a haul-out at Cape Peirce in southwestern Alaska. Three behaviors (alertness, displacement, and dispersal) were quantified from 16 capture sessions. Herd sizes ranged from 622 to 5,289 walruses. Handling of an immobilized walrus consisted of attempts to attach telemetry devices to the tusks and collect various biological samples. Handling activities resulted in an average of about 10-fold or greater levels of behavior in alertness, displacement, and dispersal than during precapture and darting periods. High levels of behavior usually occurred within the first 45 min of handling. In 8 of 10 capture sessions, walruses returned to predisturbance levels of behavior within 40 min of cessation of the handling disturbance. Alertness and displacement were moderately and negatively correlated with herd size during the handling period, which may reflect an effect of a threshold distance from the point of disturbance to responding individuals. Observations of walruses tagged with VHF radio transmitters indicated that the activities from a given capture session did not preclude tagged walruses from using the haul-out over a subsequent 11-wk monitoring period. Moreover, non-tagged walruses continued to extensively use the haul-out during and after the period in which capture sessions were conducted.

Marine Mammal Science

Spirit rover localization and topographic mapping at the landing site of Gusev crater, Mars

By sol 440, the Spirit rover has traversed a distance of 3.76 km (actual distance traveled instead of odometry). Localization of the lander and the rover along the traverse has been successfully performed at the Gusev crater landing site. We localized the lander in the Gusev crater using two-way Doppler radio positioning and cartographic triangulations through landmarks visible in both orbital and ground images. Additional high-resolution orbital images were used to verify the determined lander position. Visual odometry and bundle adjustment technologies were applied to compensate for wheel slippage, azimuthal angle drift, and other navigation errors (which were as large as 10.5% in the Husband Hill area). We generated topographic products, including 72 ortho maps and three-dimensional (3-D) digital terrain models, 11 horizontal and vertical traverse profiles, and one 3-D crater model (up to sol 440). Also discussed in this paper are uses of the data for science operations planning, geological traverse surveys, surveys of wind-related features, and other science applications.

Journal of Geophysical Research E: Planets

Satellite telemetry and wildlife studies in India: Advantages, options and challenges

Greater spatial coverage, accuracy and non-invasiveness of satellite technology make it one of the best tools to track long-distance migrants, which is otherwise difficult using conventional radio telemetry. In this article, we review the evolution of satellite telemetry and its application. We provide examples of three recent studies in India that have demonstrated and created a widespread appreciation of the use and benefits of satellite telemetry among biologists and managers. We also discuss the future prospects of this technology vis-a-vis benefits and challenges in the Indian subcontinent.

Current Science

Estimation of walrus populations on sea ice with infrared imagery and aerial photography

Population sizes of ice-associated pinnipeds have often been estimated with visual or photographic aerial surveys, but these methods require relatively slow speeds and low altitudes, limiting the area they can cover. Recent developments in infrared imagery and its integration with digital photography could allow substantially larger areas to be surveyed and more accurate enumeration of individuals, thereby solving major problems with previous survey methods. We conducted a trial survey in April 2003 to estimate the number of Pacific walruses (Odobenus rosmarus divergens) hauled out on sea ice around St. Lawrence Island, Alaska. The survey used high altitude infrared imagery to detect groups of walruses on strip transects. Low altitude digital photography was used to determine the number of walruses in a sample of detected groups and calibrate the infrared imagery for estimating the total number of walruses. We propose a survey design incorporating this approach with satellite radio telemetry to estimate the proportion of the population in the water and additional low-level flights to estimate the proportion of the hauled-out population in groups too small to be detected in the infrared imagery. We believe that this approach offers the potential for obtaining reliable population estimates for walruses and other ice-associated pinnipeds. ?? 2007 by the Society for Marine Mammalogy.

Marine Mammal Science

Global positioning system and associated technologies in animal behaviour and ecological research

Biologists can equip animals with global positioning system (GPS) technology to obtain accurate (less than or equal to 30 m) locations that can be combined with sensor data to study animal behaviour and ecology. We provide the background of GPS techniques that have been used to gather data for wildlife studies. We review how GPS has been integrated into functional systems with data storage, data transfer, power supplies, packaging and sensor technologies to collect temperature, activity, proximity and mortality data from terrestrial species and birds. GPS 'rapid fixing' technologies combined with sensors provide location, dive frequency and duration profiles, and underwater acoustic information for the study of marine species. We examine how these rapid fixing technologies may be applied to terrestrial and avian applications. We discuss positional data quality and the capability for high-frequency sampling associated with GPS locations. We present alternatives for storing and retrieving data by using dataloggers (biologging), radio-frequency download systems (e.g. very high frequency, spread spectrum), integration of GPS with other satellite systems (e.g. Argos, Globalstar) and potential new data recovery technologies (e.g. network nodes). GPS is one component among many rapidly evolving technologies. Therefore, we recommend that users and suppliers interact to ensure the availability of appropriate equipment to meet animal research objectives.

Philosophical Transactions of the Royal Society B:

To the editor: Reply confuses newsprint with media

“Is Science in Danger of Sanctifying the Wolf” (Mech, 2012), contends that some scientists have claimed that wolf (Canis lupus) restoration has led to many ecological benefits lower in the food chain but that other scientists have questioned the evidence for those claims. Furthermore, my article claims that “the media become complicit [in promoting positive studies about wolves] by immediately publicizing such studies (Table 1)….” Dr. Bruskotter’s letter (Bruskotter, 2012) contends that Houston et al. (2010) refute that statement. However, Houston et al. (2010) covered only newsprint, including many local newspapers dating back to 1999 before most of the scientific articles I mentioned were published. My article said nothing about newsprint or news media. It stated “media” which of course includes radio, TV, blogs, websites, magazines, social media, etc. These types of media reach a wider and often different audience than newsprint. The Houston et al. (2010)analysis included none of these latter types of media so does not refute my statements.

Biological Conservation

USGS Cascades Volcano Observatory news media management guide — General protocols and templates

This guide describes general protocols and provides templates for news media management at the U.S. Geological Survey (USGS) Cascades Volcano Observatory (CVO) and is intended for use by the CVO scientist-in-charge, communications staff, scientists, and guest communications colleagues. This public version, with CVO names and contact information removed, may be useful to other agencies developing their own protocols and templates. This guide evolved from a smaller document hastily assembled out of necessity during the complex and overwhelming news media interest in the 2004–2008 Mount St. Helens eruption. News media interest exceeded the need for life-saving crisis communication and foretold of the need for future multi-faceted and well-coordinated news media and social media responses during future volcanic events. This guide accompanies the USGS Volcano Science Center’s (VSC’s) general guidelines and protocols for how communications staff at all VSC observatories will work together to respond to news media requests. The protocols and templates are applicable to (1) normal conditions when CVO has an opportunity to be proactive with its messages and to raise general awareness, (2) general responses to news media and TV documentary inquiries, (3) intense news media interest where the responsibility to communicate information and hazards rests primarily with staff at CVO, and (4) intense and overwhelming news media interest that requires a multiagency response. This guide reflects general protocols in effect at the time of publication. The information will be modified as conditions change. Although “news media” generally refers to traditional outlets such as TV, radio, and newspapers, the protocols used to engage these traditional outlets apply also when responding to bloggers, online news services, and social media comments.

Circular

The geomagnetic blitz of September 1941

Seventy-five years ago, on 18–19 September 1941, the Earth experienced a great magnetic storm, one of the most intense ever recorded. It arrived at a poignant moment in history, when radio and electrical technology was emerging as a central part of daily life and when much of the world was embroiled in World War II, which the United States had not yet officially entered. Auroras danced across the night sky as voltage surged in power grid lines. A radio blackout interrupted fan enjoyment of a baseball game, while another radio program was interrupted by private phone conversations. Citizens, already on edge, wondered if neon lights were some sort of antiaircraft signal. And far away in the North Atlantic, the illuminated night sky exposed an Allied convoy to German attack. These effects raised awareness within the scientific community and among the public of the societal significance of the effects that the Sun and outer space can have on the Earth—what we now call space weather .

Eos, Earth and Space Science News

Movement patterns of rural and suburban white-tailed deer in Massachusetts

We used satellite land cover data and the program FRAGSTATS toquantify land cover types and calculate the amount of forest edge available in suburban and rural regions of northeastern and northwestern Massachusetts. Cover categories included forest cover, open canopy vegetation, and non-deer habitat. We calculated all edge segments where forest cover abutted open canopy cover. Our open canopy vegetation category was calculated both with and without low intensity suburban development. We then compared these findings to movement data from 53 (13 males, 40 females) adult radio-marked white-tailed deerOdocoileus virginianusmonitored biweekly and diurnally from January 2001 to January 2003. The range of movements of suburban deer in eastern Massachusetts showed no difference to that of suburban deer in western Massachusetts (P = 0.7). However, the ranges for suburban deer in both eastern and western Massachusetts were 10 times less than those of deer in rural western Massachusetts (P = 0.001).Our findings suggest that landscape configuration, as described by the amount and distribution of edge due to suburban development, which is related to the amount and distribution of resources such as food and cover, affects migratory behavior of white-tailed deer, allowsdeer to have smaller ranges, and contributes to high deer densities.Inclusion of suburban edge in habitat models will increase our understanding of deer-habitat relationships for management of deer in urbanizing environments. ?? 2005 Springer Science + Business Media, Inc.

Urban Ecosystems

Initial results of rover localization and topographic mapping for the 2003 mars exploration rover mission

This paper presents the initial results of lander and rover localization and topographic mapping of the MER 2003 mission (by Sol 225 for Spirit and Sol 206 for Opportunity). The Spirit rover has traversed a distance of 3.2 km (actual distance traveled instead of odometry) and Opportunity at 1.2 km. We localized the landers in the Gusev Crater and on the Meridiani Planum using two-way Doppler radio positioning technology and cartographic triangulations through landmarks visible in both orbital and ground images. Additional high-resolution orbital images were taken to verify the determined lander positions. Visual odometry and bundle-adjustment technologies were applied to overcome wheel slippages, azimuthal angle drift and other navigation errors (as large as 21 percent). We generated timely topographic products including 68 orthophoto maps and 3D Digital Terrain Models, eight horizontal rover traverse maps, vertical traverse profiles up to Sol 214 for Spirit and Sol 62 for Opportunity, and five 3D crater models. A web-based landing-site Geographic Information System (GIS) has been set up at The Ohio State University to update and disseminate the daily localization and topographic information to support tactical and strategic operations of the mission. Also described in this paper are applications of the data for science operations planning, geological traverse survey, survey of wind-related features, and other science applications. The majority of the instruments onboard both rovers are healthy at this moment, and they will continue to explore the two landing sites on the Martian surface. We expect to report further localization and topographic mapping results to be achieved in the rest of the mission period and in post-mission data processing.

Photogrammetric Engineering and Remote Sensing

10. The surface and interior of venus

Present ideas about the surface and interior of Venus are based on data obtained from (1) Earth-based radio and radar: temperature, rotation, shape, and topography; (2) fly-by and orbiting spacecraft: gravity and magnetic fields; and (3) landers: winds, local structure, gamma radiation. Surface features, including large basins, crater-like depressions, and a linear valley, have been recognized from recent ground-based radar images. Pictures of the surface acquired by the USSR's Venera 9 and 10 show abundant boulders and apparent wind erosion. On the Pioneer Venus 1978 Orbiter mission, the radar mapper experiment will determine surface heights, dielectric constant values and small-scale slope values along the sub-orbital track between 50??S and 75??N. This experiment will also estimate the global shape and provide coarse radar images (40-80 km identification resolution) of part of the surface. Gravity data will be obtained by radio tracking. Maps combining radar altimetry with spacecraft and ground-based images will be made. A fluxgate magnetometer will measure the magnetic fields around Venus. The radar and gravity data will provide clues to the level of crustal differentiation and tectonic activity. The magnetometer will determine the field variations accurately. Data from the combined experiments may constrain the dynamo mechanism; if so, a deeper understanding of both Venus and Earth will be gained. ?? 1977 D. Reidel Publishing Company.

Space Science Reviews

Mercury concentrations and space use of pre-breeding American avocets and black-necked stilts in San Francisco Bay

We examined factors influencing mercury concentrations in pre-breeding American avocets ( Recurvirostra americana ) and black-necked stilts ( Himantopus mexicanus ), the two most abundant breeding shorebirds in San Francisco Bay, California. We tested the effects of species, site, sex, year, and date on total mercury concentrations in blood of pre-breeding adult birds and used radio telemetry to determine space use and sites of dietary mercury exposure. We collected blood from 373 avocets and 157 stilts from February to April in 2005 and 2006, radio-marked and tracked 115 avocets and 94 stilts, and obtained 2393 avocet and 1928 stilt telemetry locations. Capture site was the most important factor influencing mercury concentrations in birds, followed by species and sex. Mercury concentrations were higher in stilts (geometric mean: 1.09 μg g − 1 wet weight [ww]) than in avocets (0.25 μg g − 1 ww) and males (stilts: 1.32 μg g − 1 ww; avocets: 0.32 μg g − 1 ww) had higher levels than females (stilts: 1.15 μg g − 1 ww; avocets: 0.21 μg g − 1 ww). Mercury concentrations were highest for both species at the southern end of San Francisco Bay, especially in salt pond A8 (stilts: 3.31 μg g − 1 ww; avocets: 0.58 μg g − 1 ww). Radio telemetry data showed that birds had strong fidelity to their capture site. Avocets primarily used salt ponds, tidal marshes, tidal flats, and managed marshes, whereas stilts mainly used salt ponds, managed marshes, and tidal marshes. Our results suggest that variation in blood mercury concentrations among sites was attributed to differences in foraging areas, and species differences in habitat use and foraging strategies may increase mercury exposure in stilts more than avocets.

Science of the Total Environment

Looking forward, looking back: Building resilience today community report: St. Michael, AK

The Alaska Climate Adaptation Science Center (AK CASC), in partnership with the Aleutian Pribilof Islands Association (APIA), designed the Looking Forward, Looking Back: Building Resilience Today (hereafter ‘BRT’) project as a series of trainings and workshops with tribal community leadership and members. The overarching goal of the project was to collaboratively develop the Indigenous knowledge and western science knowledge for adaptation planning. We worked with five community teams consisting of up to four leaders from communities that chose to participate in the project: Iliamna, Kotlik, Kwigillingok, Quinhagak, and St. Michael. Community teams were developed through the application process and the project duration. Community teams were encouraged to have involvement from multiple governing bodies within the community that could include the Tribal Council, city government, and village corpora- tion. The project title, with its references to the future (Looking Forward), past (Looking Back), and present (Building Resilience Today), refers to the idea that adaptation planning relies on all three perspectives. Equally important, how- ever, is the dialogue to exchange past and present information, context, and what we expect in the future. According- ly, two training sessions held at the International Arctic Research Center in Fairbanks, Alaska at the beginning and near the end of the project were developed to provide community team interaction with each other and with university and federal science partners. The project team also traveled to the partner communities and held a series of onsite events with community members to document locally-relevant information and share climate science tailored to the needs and conditions of each community. This report represents the community information shared during those onsite events. The Meeting Announcement (page 5) shows the date and description of the outreach events. The purpose of these events was to: 1) facilitate mapping of a Traditional Use Area to refine an area for climate pro- jections; 2) construct current and past seasonal Subsistence Calendars to identify important species and times of the year; 3) document Indigenous and local knowledge from current community members about environmental changes they have observed over their lifetimes; and 4) assist with documenting what the community perceived to be climate-related issues through photos and interviews. The agenda of the visits was co-produced with the commu- nity team. In each community, the community team and the project team co-hosted an open-to-the-public meeting and met with various groups. The community team advertised the meetings by posting community fliers, making announcements on the community radio, and reaching out to individuals that would contribute to the engagement discussions. Each community meeting focused on activities to develop seasonal Subsistence Calendars, map Tradition- al Use Areas, and document observed environmental changes. Community members spent time at stations dedicated to each of these activities working with project team members. The project team also met with various groups of indi- viduals that included village corporation, tribal council, and city representatives where additional information about observed environmental changes was gathered. This community report presents some of the information developed in these activities.

Alaska

Looking forward, looking back: Building resilience today community report: Kwigillingok, AK

The Alaska Climate Adaptation Science Center (AK CASC), in partnership with the Aleutian Pribilof Islands Association (APIA), designed the Looking Forward, Looking Back: Building Resilience Today (hereafter ‘BRT’) project as a series of trainings and workshops with tribal community leadership and members. The overarching goal of the project was to collaboratively develop the Indigenous knowledge and western science knowledge for adaptation planning. We worked with five community teams consisting of up to four leaders from communities that chose to participate in the project: Iliamna, Kotlik, Kwigillingok, Quinhagak, and St. Michael. Community teams were developed through the application process and the project duration. Community teams were encouraged to have involvement from multiple governing bodies within the community that could include the Tribal Council, the city government, and the village corporation. The project title, with its references to the future (Looking Forward), past (Looking Back), and present (Building Resilience Today), refers to the idea that adaptation planning relies on all three perspectives. Equally import- ant, however, is the dialogue to exchange past and present information, context, and what we expect in the future. Ac- cordingly, two training sessions held at the International Arctic Research Center in Fairbanks, Alaska at the beginning and near the end of the project were developed to provide community team interaction with each other and with university and federal science partners. The project team also traveled to the partner communities and held a series of onsite events with community members to document locally-relevant information and share climate science tailored to the needs and conditions of each community. This report represents the community information shared during those onsite events. The Meeting Announcement (page 5) shows the date and description of the outreach events. The purpose of these events was to: 1) facilitate mapping of a Traditional Use Area to refine an area for climate pro- jections; 2) construct current and past seasonal Subsistence Calendars to identify important species and times of the year; 3) document Indigenous and local knowledge from current community members about environmental changes they have observed over their lifetimes; and 4) assist with documenting what the community perceived to be climate-related issues through photos and interviews. The agenda of the visits was co-produced with the commu- nity team. In each community, the community team and the project team co-hosted an open-to-the-public meeting and met with various groups. The community team advertised the meetings by posting community fliers, making announcements on the community radio, and reaching out to individuals that would contribute to the engagement discussions. Each community meeting focused on activities to develop seasonal Subsistence Calendars, map Tradition- al Use Areas, and document observed environmental changes. Community members spent time at stations dedicated to each of these activities working with project team members. The project team also met with various groups of indi- viduals that included village corporation, tribal council, and city representatives where additional information about observed environmental changes was gathered. This community report presents some of the information developed in these activities.

Alaska

Looking forward, looking back: Building resilience today community report: Kotlik, AK

The Alaska Climate Adaptation Science Center (AK CASC), in partnership with the Aleutian Pribilof Islands Association (APIA), designed the Looking Forward, Looking Back: Building Resilience Today (hereafter ‘BRT’) project as a series of trainings and workshops with tribal community leadership and members. The overarching goal of the project was to collaboratively develop the Indigenous knowledge and western science knowledge for adaptation planning. We worked with five community teams consisting of up to four leaders from communities that chose to participate in the project: Iliamna, Kotlik, Kwigillingok, Quinhagak, and St. Michael. Community teams were developed through the application process and the project duration. Community teams were encouraged to have involvement from multiple governing bodies within the community that could include the Tribal Council, the city governments, and the village corporations. The project title, with its references to the future (Looking Forward), past (Looking Back), and present (Building Resil- ience Today), refers to the idea that adaptation planning relies on all three perspectives. Equally important, however, is the dialogue to exchange past and present information, context, and what we expect in the future. Accordingly, two training sessions held at the International Arctic Research Center in Fairbanks, Alaska at the beginning and near the end of the project were developed to provide community team interaction with each other and with university and feder- al science partners. The project team also traveled to the partner communities and held a series of onsite events with community members to document locally-relevant information and share climate science tailored to the needs and conditions of each community. This report represents the community information shared during those onsite events. The Meeting Announcement (page 5) shows the date and description of the outreach events. The purpose of these events was to: 1) facilitate mapping of a Traditional Use Area to refine an area for climate projec- tions; 2) construct current and past seasonal Subsistence Calendars to identify important species and times of the year; 3) document Indigenous and local knowledge from current community members about environmental changes they have observed over their lifetimes; and 4) assist with documenting what the community perceived to be climate-relat- ed issues through photos and interviews. The agenda of the visits was co-produced with the community team. In each community, the community team and the project team co-hosted an open-to-the-public meeting and met with various groups. The community team advertised the meetings by posting community fliers, making announcements on the community radio, and reaching out to individuals that would contribute to the engagement discussions. Each commu- nity meeting focused on activities to develop seasonal Subsistence Calendars, map Traditional Use Areas, and document observed environmental changes. Community members spent time at stations dedicated to each of these activities working with project team members. The project team also met with various groups of individuals that included village corporation, tribal council, and city representatives where additional information about observed environmental chang- es was gathered. This community report presents some of the information developed in these activities.

Alaska

Results and evaluation of a survey to estimate Pacific walrus population size, 2006

In spring 2006, we conducted a collaborative U.S.-Russia survey to estimate abundance of the Pacific walrus ( Odobenus rosmarus divergens ). The Bering Sea was partitioned into survey blocks, and a systematic random sample of transects within a subset of the blocks was surveyed with airborne thermal scanners using standard strip-transect methodology. Counts of walruses in photographed groups were used to model the relation between thermal signatures and the number of walruses in groups, which was used to estimate the number of walruses in groups that were detected by the scanner but not photographed. We also modeled the probability of thermally detecting various-sized walrus groups to estimate the number of walruses in groups undetected by the scanner. We used data from radio-tagged walruses to adjust on-ice estimates to account for walruses in the water during the survey. The estimated area of available habitat averaged 668,000 km 2 and the area of surveyed blocks was 318,204 km 2 . The number of Pacific walruses within the surveyed area was estimated at 129,000 with 95% confidence limits of 55,000 to 507,000 individuals. This value can be used by managers as a minimum estimate of the total population size.

Bering Sea

Resource selection of the southern fox squirrel (Sciurus niger niger) in the coastal plain of Virginia

The southern fox squirrel ( Sciurus niger niger ) is a subspecies of fox squirrel that ranges from southeastern Virginia to northern Florida. Throughout its Southeastern range, southern fox squirrel habitat of natural mixed pine-hardwood forests has been fragmented by agriculture and short-rotation pine plantations. In Virginia, remaining habitat has been transformed by fire suppression on the landscape and has in turn resulted in small, disjunct populations of southern fox squirrels. We sought to identify the home range and resource utilization of southern fox squirrels in the southern Coastal Plain of Virginia. From established nest boxes, we captured, radio collared and tracked four individuals at Big Woods Wildlife Management area and Piney Grove Complex in Sussex County in 2022. We observed an average male 95% and 50% adaptive kernel home range 173.49 ha (SE = 25.73, N = 2) and 40.62 ha (SE = 5.87, N = 2), respectively and an average female 95% and 50% adaptive kernel home range of 28.51 ha (SE = 0.49, N = 2) and 4.71 ha (SE = 0.34, N = 2), respectively. At both the second and third orders of habitat selection, squirrels selected for pine savanna cover types and short fire-return intervals.

Virginia

Application of a hybrid model to reduce bias and improve precision in population estimates for elk (Cervus elaphus) inhabiting a cold desert ecosystem

Accurately estimating the size of wildlife populations is critical to wildlife management and conservation of species. Raw counts or “minimum counts” are still used as a basis for wildlife management decisions. Uncorrected raw counts are not only negatively biased due to failure to account for undetected animals, but also provide no estimate of precision on which to judge the utility of counts. We applied a hybrid population estimation technique that combined sightability modeling, radio collar-based mark-resight, and simultaneous double count (double-observer) modeling to estimate the population size of elk in a high elevation desert ecosystem. Combining several models maximizes the strengths of each individual model while minimizing their singular weaknesses. We collected data with aerial helicopter surveys of the elk population in the San Luis Valley and adjacent mountains in Colorado State, USA in 2005 and 2007. We present estimates from 7 alternative analyses: 3 based on different methods for obtaining a raw count and 4 based on different statistical models to correct for sighting probability bias. The most reliable of these approaches is a hybrid double-observer sightability model (model M H ), which uses detection patterns of 2 independent observers in a helicopter plus telemetry-based detections of radio collared elk groups. Data were fit to customized mark-resight models with individual sighting covariates. Error estimates were obtained by a bootstrapping procedure. The hybrid method was an improvement over commonly used alternatives, with improved precision compared to sightability modeling and reduced bias compared to double-observer modeling. The resulting population estimate corrected for multiple sources of undercount bias that, if left uncorrected, would have underestimated the true population size by as much as 22.9%. Our comparison of these alternative methods demonstrates how various components of our method contribute to improving the final estimate and demonstrates why each is necessary.

Colorado