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At least 37 records · Page 2Linked to original sources

The use (and misuse) of sediment traps in coral reef environments: Theory, observations, and suggested protocols

Sediment traps are commonly used as standard tools for monitoring “sedimentation” in coral reef environments. In much of the literature where sediment traps were used to measure the effects of “sedimentation” on corals, it is clear from deployment descriptions and interpretations of the resulting data that information derived from sediment traps has frequently been misinterpreted or misapplied. Despite their widespread use in this setting, sediment traps do not provide quantitative information about “sedimentation” on coral surfaces. Traps can provide useful information about the relative magnitude of sediment dynamics if trap deployment standards are used. This conclusion is based first on a brief review of the state of knowledge of sediment trap dynamics, which has primarily focused on traps deployed high above the seabed in relatively deep water, followed by our understanding of near-bed sediment dynamics in shallow-water environments that characterize coral reefs. This overview is followed by the first synthesis of near-bed sediment trap data collected with concurrent hydrodynamic information in coral reef environments. This collective information is utilized to develop nine protocols for using sediment traps in coral reef environments, which focus on trap parameters that researchers can control such as trap height (H), trap mouth diameter (D), the height of the trap mouth above the substrate (z o ), and the spacing between traps. The hydrodynamic behavior of sediment traps and the limitations of data derived from these traps should be forefront when interpreting sediment trap data to infer sediment transport processes in coral reef environments.

Coral Reefs

Rearing environment influences boldness and prey acquisition behavior, and brain and lens development of bull trout

Animals reared in barren captive environments exhibit different developmental trajectories and behaviors than wild counterparts. Hence, the captive phenotypes may influence the success of reintroduction and recovery programs for threatened and endangered species. We collected wild bull trout embryos from the Metolius River Basin, Oregon and reared them in differing environments to better understand how captivity affects the bull trout Salvelinus confluentus phenotype. We compared the boldness and prey acquisition behaviors and development of the brain and eye lens of bull trout reared in conventional barren and more structurally complex captive environments with that of wild fish. Wild fish and captive reared fish from complex habitats exhibited a greater level of boldness and prey acquisition ability, than fish reared in conventional captive environments. In addition, the eye lens of conventionally reared bull trout was larger than complex reared captive fish or same age wild fish. Interestingly, we detected wild fish had a smaller relative cerebellum than either captive reared treatment. Our results suggest that rearing fish in more complex captive environments can create a more wild-like phenotype than conventional rearing practices. A better understanding of the effects of captivity on the development and behavior of bull trout can inform rearing and reintroduction programs though prediction of the performance of released individuals.

Environmental Biology of Fishes

Limnological and climatic environments at Upper Klamath Lake, Oregon during the past 45 000 years

Upper Klamath Lake, in south-central Oregon, contains long sediment records with well-preserved diatoms and lithological variations that reflect climate-induced limnological changes. These sediment archives complement and extend high resolution terrestrial records along a north-south transect that includes areas influenced by the Aleutian Low and Subtropical High, which control both marine and continental climates in the western United States. The longest and oldest core collected in this study came from the southwest margin of the lake at Caledonia Marsh, and was dated by radiocarbon and tephrochronology to an age of about 45 ka. Paleolimnological interpretations of this core, based upon geochemical and diatom analyses, have been augmented by data from a short core collected from open water environments at nearby Howards Bay and from a 9-m core extending to 15 ka raised from the center of the northwestern part of Upper Klamath Lake. Pre- and full-glacial intervals of the Caledonia Marsh core are characterized and dominated by lithic detrital material. Planktic diatom taxa characteristic of cold-water habitats (Aulacoseira subarctica and A. islandica) alternate with warm-water planktic diatoms (A. ambigua) between 45 and 23 ka, documenting climate changes at millennial scales during oxygen isotope stage (OIS) 3. The full-glacial interval contains mostly cold-water planktic, benthic, and reworked Pliocene lacustrine diatoms (from the surrounding Yonna Formation) that document shallow water conditions in a cold, windy environment. After 15 ka, diatom productivity increased. Organic carbon and biogenic silica became significant sediment components and diatoms that live in the lake today, indicative of warm, eutrophic water, became prominent. Lake levels fell during the mid-Holocene and marsh environments extended over the core site. This interval is characterized by high levels of organic carbon from emergent aquatic vegetation (Scirpus) and by the Mazama ash (7.55 ka), generated by the eruption that created nearby Crater Lake. For a brief time the ash increased the salinity of Upper Klamath Lake. High concentrations of molybdenum, arsenic, and vanadium indicate that Caledonia Marsh was anoxic from about 7 to 5 ka. After the mid-Holocene, shallow, but open-water environments returned to the core site. The sediments became dominated (>80%) by biogenic silica. The open-water cores show analogous but less extreme limnological and climatic changes more typical of mid-lake environments. Millennial-scale lake and climate changes during OIS 3 at Upper Klamath Lake contrast with a similar record of variation at Owens Lake, about 750 km south. When Upper Klamath Lake experienced cold-climate episodes during OIS 3, Owens Lake had warm but wet episodes; the reverse occurred during warmer intervals at Upper Klamath Lake. Such climatic alternations apparently reflect the variable position and strength of the Aleutian Low during the mid-Wisconsin.

Oregon

Water temperatures in select nearshore environments of the Colorado River in Grand Canyon, Arizona, during the Low Steady Summer Flow experiment of 2000

Water releases from Glen Canyon Dam, Arizona, are the primary determinant of streamflow, sediment transport, water quality, and aquatic and riparian habitat availability in the Colorado River downstream of the dam in Grand Canyon. The presence and operation of the dam have transformed the seasonally warm Colorado River into a consistently cold river because of hypolimnetic, or deep-water, releases from the penstock withdrawal structures on the dam. These releases have substantially altered the thermal regime of the downstream riverine environment. This, in turn, has affected the biota of the river corridor, particularly native and nonnative fish communities and the aquatic food web. In the spring and summer of 2000, a Low Steady Summer Flow experiment was conducted by the U.S. Geological Survey and the Bureau of Reclamation to evaluate the effects of the experimental flow on physical and biological resources of the Colorado River ecosystem downstream from Glen Canyon Dam to Lake Mead on the Arizona-Nevada border. This report describes the water temperatures collected during the experimental flow from 14 nearshore sites in the river corridor in Grand Canyon to assess the effects of steady releases on the thermal dynamics of nearshore environments. These nearshore areas are characterized by low-velocity flows with some degree of isolation from the higher velocity flows in the main channel and are hypothesized to be important rearing environments for young native fish. Water-temperature measurements were made at 14 sites, ranging from backwater to open-channel environments. Warming during daylight hours, relative to main-channel temperatures, was measured at all sites in relation to the amount of isolation from the main-channel current. Boat traffic, amount of direct solar radiation, and degree of isolation from the main-channel current appear to be the primary factors affecting the differential warming of the nearshore environment.

Arizona

Sea-floor morphology and sedimentary environments of western Block Island Sound, northeast of Gardiners Island, New York

Multibeam-echosounder data, collected during survey H12299 by the National Oceanic and Atmospheric Administration in a 162-square-kilometer area of Block Island Sound, northeast of Gardiners Island, New York, are used along with sediment samples and bottom photography, collected at 37 stations in this area by the U.S. Geological Survey during cruise 2013-005-FA, to interpret sea-floor features and sedimentary environments. These data and interpretations provide important base maps for future studies of the sea floor, focused, for example, on benthic ecology and resource management. The features and sedimentary environments on the sea floor are products of the glacial history and modern tidal regime. Features include bedforms such as sand waves and megaripples, boulders, a large current-scoured depression, exposed glaciolacustrine sediments, and areas of modern marine sediment. Sand covers much of the study area and is often in the form of sand waves and megaripples, which indicate environments characterized by coarse-grained bedload transport. Boulders and gravelly lag deposits, which indicate environments of erosion or nondeposition, are found off the coast of Gardiners Island and on bathymetric highs, probably marking areas where deposits associated with recessional ice-front positions, the northern flank of the terminal moraine, or coastal-plain sediments covered with basal till are exposed. Bottom photographs and video of boulders show that they are commonly covered with sessile fauna. Strong tidal currents have produced the deep scour depression along the northwestern edge of the study area. The eastern side of this depression is armored with a gravel lag. Sea-floor areas characterized by modern marine sediments appear featureless at the 2-meter resolution of the bathymetry and flat to current rippled in the photography. These modern environments are indicative of sediment sorting and reworking.

New York

Sea-floor morphology and sedimentary environments in southern Narragansett Bay, Rhode Island

Multibeam echosounder data collected by the National Oceanic and Atmospheric Administration along with sediment samples and still and video photography of the sea floor collected by the U.S. Geological Survey were used to interpret sea-floor features and sedimentary environments in southern Narragansett Bay, Rhode Island, as part of a long-term effort to map the sea floor along the northeastern coast of the United States. Sea-floor features include rocky areas and scour depressions in high-energy environments characterized by erosion or nondeposition, and sand waves and megaripples in environments characterized by coarse-grained bedload transport. Two shipwrecks are also located in the study area. Much of the sea floor is relatively featureless within the resolution of the multibeam data; sedimentary environments in these areas are characterized by processes associated with sorting and reworking. This report releases bathymetric data from the multibeam echosounder, grain-size analyses of sediment samples, and photographs of the sea floor and interpretations of the sea-floor features and sedimentary environments. It provides base maps that can be used for resource management and studies of topics such as benthic ecology, contaminant inventories, and sediment transport.

Rhode Island

Optimal exploitation strategies for an animal population in a stochastic serially correlated environment

Optimal exploitation strategies were studied for an animal population in a stochastic, serially correlated environment. This is a general case and encompasses a number of important cases as simplifications. Data on the mallard (Anas platyrhynchos) were used to explore the exploitation strategies and test several hypotheses because relatively much is known concerning the life history and general ecology of this species and extensive empirical data are available for analysis. The number of small ponds on the central breeding grounds was used as an index to the state of the environment. Desirable properties of an optimal exploitation strategy were defined. A mathematical model was formulated to provide a synthesis of the existing literature, estimates of parameters developed from an analysis of data, and hypotheses regarding the specific effect of exploitation on total survival. Both the literature and the analysis of data were inconclusive concerning the effect of exploitation on survival. Therefore, alternative hypotheses were formulated: (1) exploitation mortality represents a largely additive form of mortality, or (2 ) exploitation mortality is compensatory with other forms of mortality, at least to some threshold level. Models incorporating these two hypotheses were formulated as stochastic dynamic programming models and optimal exploitation strategies were derived numerically on a digital computer. Optimal exploitation strategies were found to exist under rather general conditions. Direct feedback control was an integral component in the optimal decision-making process. Optimal exploitation was found to be substantially different depending upon the hypothesis regarding the effect of exploitation on the population. Assuming that exploitation is largely an additive force of mortality, optimal exploitation decisions are a convex function of the size of the breeding population and a linear or slightly concave function of the environmental conditions. Optimal exploitation under this hypothesis tends to reduce the variance of the size of the population. Under the hypothesis of compensatory mortality forces, optimal exploitation decisions are approximately linearly related to the size of the breeding population. Environmental variables may be somewhat more important than the size of the breeding population to the production of young mallards. In contrast, the size of the breeding population appears to be more important in the exploitation process than is the state of the environment. The form of the exploitation strategy appears to be relatively insensitive to small changes in the production rate. In general, the relative importance of the size of the breeding population may decrease as fecundity increases. The optimal level of exploitation in year t must be based on the observed size of the population and the state of the environment in year t unless the dynamics of the population, the state of the environment, and the result of the exploitation decisions are completely deterministic. Exploitation based on an average harvest, harvest rate, or designed to maintain a constant breeding population size is inefficient.

Book

Optimal exploitation strategies for an animal population in a stochastic serially correlated environment

Optimal exploitation strategies were studied for an animal population in a stochastic, serially correlated environment. This is a general case and encompasses a number of important cases as simplifications. Data on the mallard (Anas platyrhynchos) were used to explore the exploitation strategies and test several hypotheses because relatively much is known concerning the life history and general ecology of this species and extensive empirical data are available for analysis. The number of small ponds on the central breeding grounds was used as an index to the state of the environment. Desirable properties of an optimal exploitation strategy were defined. A mathematical model was formulated to provide a synthesis of the existing literature, estimates of parameters developed from an analysis of data, and hypotheses regarding the specific effect of exploitation on total survival. Both the literature and the analysis of data were inconclusive concerning the effect of exploitation on survival. Therefore, alternative hypotheses were formulated: (1) exploitation mortality represents a largely additive form of mortality, or (2 ) exploitation mortality is compensatory with other forms of mortality, at least to some threshold level. Models incorporating these two hypotheses were formulated as stochastic dynamic programming models and optimal exploitation strategies were derived numerically on a digital computer. Optimal exploitation strategies were found to exist under rather general conditions. Direct feedback control was an integral component in the optimal decision-making process. Optimal exploitation was found to be substantially different depending upon the hypothesis regarding the effect of exploitation on the population. Assuming that exploitation is largely an additive force of mortality, optimal exploitation decisions are a convex function of the size of the breeding population and a linear or slightly concave function of the environmental conditions. Optimal exploitation under this hypothesis tends to reduce the variance of the size of the population. Under the hypothesis of compensatory mortality forces, optimal exploitation decisions are approximately linearly related to the size of the breeding population. Environmental variables may be somewhat more important than the size of the breeding population to the production of young mallards. In contrast, the size of the breeding population appears to be more important in the exploitation process than is the state of the environment. The form of the exploitation strategy appears to be relatively insensitive to small changes in the production rate. In general, the relative importance of the size of the breeding population may decrease as fecundity increases. The optimal level of exploitation in year t must be based on the observed size of the population and the state of the environment in year t unless the dynamics of the population, the state of the environment, and the result of the exploitation decisions are completely deterministic. Exploitation based on an average harvest, harvest rate, or designed to maintain a constant breeding population size is inefficient.

Dissertation Abstracts International

Depositional environments as a guide to uranium mineralization in the Chinle Formation, San Rafael Swell, Utah

Uranium deposits in the San Rafael Swell are related to sedimentary depositional environments in the Upper Triassic Chinle Formation. The sedimentary textures resulting from depositional processes operating in low-energy environments appear to have influenced uranium mineralization. The Ohinle consists of three fining-upward, fluvial-lacustrine sequences. Uranium minerals are concentrated in the lower part of the lowest sequence in areas where sediments of low-energy environments are complexly interbedded with sediments of other environments. Areas favorable for uranium exploration exist in the subsurface to the north, west, and south of the Chinle outcrop in the Swell. This determination is based on the spatial distribution of depositional environments and the pattern of Chinle deposition through time.

Utah

Observations on the use of membrane filtration and liquid impingement to collect airborne microorganisms in various atmospheric environments

The influence of sample-collection-time on the recovery of culturable airborne microorganisms using a low-flow-rate membrane-filtration unit and a high-flow-rate liquid impinger were investigated. Differences in recoveries were investigated in four different atmospheric environments, one mid-oceanic at an altitude of ~10.0 m, one on a mountain top at an altitude of ~3,000.0 m, one at ~1.0 m altitude in Tallahassee, Florida, and one at ~1.0 m above ground in a subterranean-cave. Regarding use of membrane filtration, a common trend was observed: the shorter the collection period, the higher the recovery of culturable bacteria and fungi. These data also demonstrated that lower culturable counts were common in the more remote mid-oceanic and mountain-top atmospheric environments with bacteria, fungi, and total numbers averaging (by sample time or method categories) <3.0 colony-forming units (CFU) m −3 . At the Florida and subterranean sites, the lowest average count noted was 3.5 bacteria CFU m −3 , and the highest averaged 140.4 total CFU m −3 . When atmospheric temperature allowed use, the high-volume liquid impinger utilized in this study resulted in much higher recoveries, as much as 10× greater in a number of the categories (bacterial, fungal, and total CFU). Together, these data illustrated that (1) the high-volume liquid impinger is clearly superior to membrane filtration for aeromicrobiology studies if start-up costs are not an issue and temperature permits use; (2) although membrane filtration is more cost friendly and has a ‘typically’ wider operational range, its limits include loss of cell viability with increased sample time and issues with effectively extracting nucleic acids for community-based analyses; (3) the ability to recover culturable microorganisms is limited in ‘extreme’ atmospheric environments and thus the use of a ‘limited’ methodology in these environments must be taken into account; and (4) the atmosphere culls, i.e., everything is not everywhere.

Aerobiologia

A new approach for representing agent-environment feedbacks: Coupled agent-based and state-and-transition simulation models

Context Agent-based models (ABMs) and state-and-transition simulation models (STSMs) have proven useful for understanding processes underlying social-ecological systems and evaluating practical questions about how systems might respond to different scenarios. ABMs can simulate a variety of agents (autonomous units, such as wildlife or people); agent characteristics, decision-making, adaptive behavior, and mobility; and agent-environment interactions. STSMs are flexible and intuitive stochastic landscape models that can track scenarios and integrate diverse data. Both can be run spatially and track metrics of management success. Objectives Due to the complementarity of these approaches, we sought to couple them through a dynamic linkage and demonstrate the relevance of this advancement for modeling landscape processes and patterns. Methods We developed analytical techniques and software tools to couple these modeling approaches using NetLogo, R, and the ST-Sim package for SyncroSim. We demonstrated the capabilities and value of this coupled approach through a proof-of-concept case study of bison-vegetation interactions in Badlands National Park. Results The coupled ABM-STSM: (1) streamlined handling of model inputs and outputs; (2) allowed representation of processes at multiple temporal scales; (3) minimized assumptions; and (4) generated spatial and temporal patterns that better reflected agent-environment interactions. Conclusions These developments constitute a new approach for representing agent-environment feedbacks; modelers can now use output from an ABM to dictate landscape changes within an STSM that in turn influence agents. This facilitates experimentation across domains (agent and environment) and creation of more realistic and management-relevant projections, and opens new opportunities for communicating models and linking to other methods.

Landscape Ecology

Modern sedimentary environments on the Rhode Island inner shelf, off the eastern United States

Analyses of side-scan sonar records along with previously published bathymetric, textural and subbottom data reveal the sedimentary environments on the inner Continental Shelf south of Narragansett Bay, Rhode Island. The bottom topography in this area is characterized by a broad central depression bordered by shallow, irregular sea floor on the north and east and by a discontinuous, curvilinear ridge on the south and west. Four distinct environments were identified: (1) Pre-Mesozoic coastal rocks are exposed on the sea floor at isolated locations near the shore (waterdepths < 32 m). These exposures have pronounced, irregular topographic relief and produce blotchy patterns op side-scan sonographs. (2) Glacial moraine deposits form the discontinuous offshore ridge. These deposits have hummocky sea-floor relief, are covered by lag gravel and boulders, and appear as predominantly black (strongly reflective) patterns on the side-scan records. (3) Over most of the shallow, irregular bottom in the northeast, on the flanks of the morainal ridge, and atop bathymetric highs, the sea floor is characterized as a mosaic of light and dark patches and lineations. The dark (more reflective) zones are areas of coarse sands and megaripples (wavelengths = 0.8--1.2 m) that either have no detectable relief or are slightly depressed relative to surrounding (fight) areas of finer-grained sands. (4) Smooth beds that produce nearly featureless patterns on the sonographs occupy the broad central bathymetric depression as well as smaller depressions north and east of Block Island. Within the broad depression, sonographs having practically no shading indicate a central zone of modern sandy silt, whereas records having moderate tonality define a peripheral belt of silty sand. The sedimentary environments that are outlined range from erosional or nondepositional (bedrock, glacial moraine) to depositional (featureless beds), and include areas that may reflect a combination of erosional and depositional processes (textural patchiness). The distribution and characteristics of the environments reveal the general post-glacial sedimentary history of this area and provide a guide to future utilization of the shelf surface.

Rhode Island

Ground level environmental protein concentrations in various ecuadorian environments: potential uses of aerosolized protein for ecological research

Large quantities of free protein in the environment and other bioaerosols are ubiquitous throughout terrestrial ground level environments and may be integrative indicators of ecosystem status. Samples of ground level bioaerosols were collected from various ecosystems throughout Ecuador, including pristine humid tropical forest (pristine), highly altered secondary humid tropical forest (highly altered), secondary transitional very humid forest (regrowth transitional), and suburban dry montane deforested (suburban deforested). The results explored the sensitivity of localized aerosol protein concentrations to spatial and temporal variations within ecosystems, and their value for assessing environmental change. Ecosystem specific variations in environmental protein concentrations were observed: pristine 0.32 ± 0.09 μg/m3, highly altered 0.07 ± 0.05 μg/m 3 , regrowth transitional 0.17 ± 0.06 μg/m 3 , and suburban deforested 0.09 ± 0.04 μg/m 3 . Additionally, comparisons of intra-environmental differences in seasonal/daily weather (dry season 0.08 ± 0.03 μg/m 3 and wet season 0.10 ± 0.04 μg/m 3 ), environmental fragmentation (buffered 0.19 ± 0.06 μg/m 3 and edge 0.15 ± 0.06 μg/m 3 ), and sampling height (ground level 0.32 ± 0.09 μg/m 3 and 10 m 0.24 ± 0.04 μg/m 3 ) demonstrated the sensitivity of protein concentrations to environmental conditions. Local protein concentrations in altered environments correlated well with satellite-based spectral indices describing vegetation productivity: normalized difference vegetation index (NDVI) (r 2 = 0.801), net primary production (NPP) (r 2 = 0.827), leaf area index (LAI) (r 2 = 0.410). Moreover, protein concentrations distinguished the pristine site, which was not differentiated in spectral indices, potentially due to spectral saturation typical of highly vegetated environments. Bioaerosol concentrations represent an inexpensive method to increase understanding of environmental changes, especially in densely vegetated ecosystems with high canopies or in areas needing high spatial and temporal resolution. Further research to expand understanding of the applicability of bioaerosol concentrations for environmental monitoring is supported by this pilot study.

Ecological Indicators

Influence of history and environment on the sediment dynamics of intertidal flats

Morphological trends of three distinct intertidal environments in South San Francisco Bay were investigated using a combination of measurement and modeling tools. Because of the inherent relationship between the physical environment and the sediment properties, the sediment properties provide a good indicator of morphologic trends. A significant finding of this study is that surface sediment erodibility increases as the energy level in the environment increases. Conversely subsurface sediment erodibility shows a strong relationship to the long-term history of the site. The combination of the measured sediment properties, the history of deposition and erosion, and simple modeling of the physical environment illustrate the interaction of these properties such that an understanding of intertidal flat behavior is developed.

California

The influence of tectonic environment on dynamic earthquake triggering: A review and case study on Alaskan volcanoes

The phenomenon of dynamic earthquake triggering, when seismic waves from an earthquake trigger seismicity at distant sites, has been recognized for over 25 years, yet knowledge of the global distribution of dynamic triggering remains far from complete. Because occurrences of dynamic triggering provide in-situ information of the stress-state of a responding site, a more complete global picture of susceptible crustal environments becomes an important component of seismic hazard evaluations. Here we 1) review evidence for tectonic regime dependence of dynamic triggering susceptibility in the shallow brittle crust, and 2) explore triggering susceptibility at Alaskan volcanoes. We search for significant increases in seismicity rates at 19 Alaskan volcanic areas between 2006 and 2013 within 3 days following regional and teleseismic earthquakes of magnitude 7 and greater. We find evidence for 12 triggered responses at 9 volcanoes, but no evidence for triggered responses following hundreds of other earthquakes. The most impressive response was that of Pavlof Volcano to the 2011 M W 9.0 Tohoku-Oki earthquake. Our results suggest that triggered responses are difficult to predict, as they do not depend on background seismicity rates or amplitude of incident seismic waves from distant earthquakes. Taken together, observational and theoretical evidence suggests that dynamic triggering occurs in all tectonic environments. So far however, it appears to be more common in extensional and transtensional environments and regions with high pore-fluid pressure than in compressional and transpressional environments. Although some volcanic and geothermal areas may be primed for dynamic earthquake triggering, volcanoes in Alaska are relatively insensitive. The weak and variable responses of Alaskan volcanoes suggest that triggering susceptibility depends on time varying crustal conditions, such as the state of stress and distribution of pore pressure. Results also indirectly suggest that eruption triggering by distant earthquakes is unlikely in Alaska over the time scale of days.

Alaska

The uniqueness of humic substances in each of soil, stream and marine environments

Definitive compositional differences are shown to exist for both fulvic acids and humic acids from soil, stream and marine environments by five different methods (1H and 13C NMR spectroscopy, 14C age and ?? 13C isotopic analyses, amino acid analyses and pyrolysis-mass spectrometry). Definitive differences are also found between fulvic acids and humic acids within each environment. These differences among humic substances from various sources are more readily discerned because the method employed for the isolation of humic substances from all environments excludes most of the non-humic components and results in more purified humic isolates from water and soils. The major compositional aspects of fulvic acids and humic acids which determine the observed characteristic differences in each environment are the amounts and compositions of saccharide, phenolic, methoxyl, aromatic, hydrocarbon, amino acid and nitrogen moieties.

Analytica Chimica Acta

Carrying capacity in a heterogeneous environment with habitat connectivity

A large body of theory predicts that populations diffusing in heterogeneous environments reach higher total size than if non-diffusing, and, paradoxically, higher size than in a corresponding homogeneous environment. However, this theory and its assumptions have not been rigorously tested. Here, we extended previous theory to include exploitable resources, proving qualitatively novel results, which we tested experimentally using spatially diffusing laboratory populations of yeast. Consistent with previous theory, we predicted and experimentally observed that spatial diffusion increased total equilibrium population abundance in heterogeneous environments, with the effect size depending on the relationship between r and K . Refuting previous theory, however, we discovered that homogeneously distributed resources support higher total carrying capacity than heterogeneously distributed resources, even with species diffusion. Our results provide rigorous experimental tests of new and old theory, demonstrating how the traditional notion of carrying capacity is ambiguous for populations diffusing in spatially heterogeneous environments.

Ecology Letters

Depositional environments of the Cache, Lower Lake, and Kelseyville Formations, Lake County, California

We describe the depositional environments of the Cache, Lower Lake, and Kelseyville Formations in light of habitat preferences of recovered mollusks, ostracodes, and diatoms. Our reconstruction of paleoenvironments for these late Cenozoic deposits provides a framework for an understanding of basin evolution and deposition in the Clear Lake region. The Pliocene and Pleistocene Cache Formation was deposited primarily in stream and debris flow environments; fossils from fine-grained deposits indicate shallow, fresh-water environments with locally abundant aquatic vegetation. The fine-grained sediments (mudstone and siltstone) were probably deposited in ponds in abandoned channels or shallow basins behind natural levees. The abandoned channels and shallow basins were associated with the fluvial systems responsible for deposition of the bulk of the technically controlled Cache Formation. The Pleistocene Lower Lake Formation was deposited in a water mass large enough to contain a variety of local environments and current regimes. The recovered fossils imply a lake with water depths of 1 to 5 m. However, there is strong support from habitat preferences of the recovered fossils for inferring a wide range of water depths during deposition of the Lower Lake Formation; they indicate a progressively shallowing system and the culmination of a desiccating lacustrine system. The Pleistocene Kelseyville Formation represents primarily lacustrine deposition with only minor fluvial deposits around the margins of the basin. Local conglomerate beds and fossil tree stumps in growth position within the basin indicate occasional widespread fluvial incursions and depositional hiatuses. The Kelseyville strata represent a large water mass with a muddy and especially fluid substrate having permanent or sporadic periods of anoxia. Central-lake anoxia, whether permanent or at irregular intervals, is the simplest way to account for the low numbers of benthic organisms recovered from the Kelseyville Formation. Similar low-oxygen conditions for benthic life are represented throughout the sedimentary history of Clear Lake. Water depths for the Kelseyville Formation of 10 to 30 m and 12 m near the margins of the basin are inferred both before and after fluvial incursions. These water-depth fluctuations cannot be correlated with major climatic changes as indicated by pollen and fossil leaves and cones; they may be due to faulting in this technically active region.

GSA Special Papers