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At least 307 records · Page 17Linked to original sources

Changes in conservation value from grasslands to savannas to forests: How a temperate canopy cover gradient affects butterfly community composition

Temperate savannas and grasslands are globally threatened. In the Midwest United States of America (USA), for example, oak savannas persist today at a small percentage of recent historic coverage. Therefore, restoration of habitats of low and intermediate canopy cover is a landscape conservation priority that often emphasizes returning tree density to a savanna-like target value. Understanding how animal species react to such changes in vegetation structure is important for assessing the value of these restoration plans. We examined how butterfly community attributes in northwest Indiana USA, including community composition, richness, and abundance responded to a grassland-to-forest gradient of canopy cover. Butterfly community composition under intermediate canopy cover differed significantly from community composition in the most open or closed-canopy habitats. Composition of the plant community in flower was a significant predictor of three assessed attributes of the butterfly community—composition, richness, and abundance. Phenology, expressed as day-of-the-year, was also a strong predictor of these butterfly community attributes. Few butterfly species were habitat specialists as adults although canopy cover was a more important predictor of adult community composition than of richness or abundance of butterflies. Therefore, adult butterfly community differences along the canopy cover gradient were less about butterfly communities filled with habitat specialists for different canopy-defined habitats and more about gradual changes in community composition along this gradient. Overall, butterfly community richness was predicted to peak at about 34% canopy cover, butterfly abundance at about 53% canopy cover, community conservation value at about 59% canopy cover, and a combination of desirable conservation attributes–high diversity, high abundance, and high conservation value–was predicted to reach a peak of co-occurrence at about 67% canopy cover suggesting that habitats of intermediate canopy cover might be particularly effective for butterfly conservation in this region.

Indiana

Conservation of temporary wetlands

Temporary wetlands are characterized by frequent drying resulting in a unique, highly specialized assemblage of often rare or specialized plant and animal species. They are found on all continents and in a variety of landscape settings. Although accurate estimates of the abundance of temporary wetlands are available in only a few countries, global estimations identify a decline in number and quality. The key environmental factors driving the structure of ecological communities in temporary wetlands are the duration, timing, frequency and predictability of the aquatic and dry phases, which varies greatly with region and hydrogeomorphic setting. Temporary wetlands have been historically neglected, but improved social awareness of the functions and values of, and increases in scientific interest, suggest that this is changing. They play an ecological role in both global cycles (i.e., CO 2 emissions) and biodiversity (in proportion to their size, they contribute disproportionately to regional and global biodiversity). Moreover, they provide valuable ecosystem services including wildlife habitat, nutrient flux to adjacent ecosystems, flood control, water filtration, and cultural services. Effective conservation of temporary wetlands requires addressing threats (i.e., inconsistent and inadequate regulatory protections; climate change; changes in land use) and management challenges (i.e., management at both local and landscape scales; incomplete understanding of the ecosystem services provided by them; the need to enhance inventories). The most suitable approaches for conserving temporary wetlands include (1) regulations or other forms of protection; (2) sustainable management; (3) restoration and creation; and (4) collaborative conservation.

Book chapter

Lack of observed movement response to lead exposure of California condors

Lead poisoning is an important conservation concern for wildlife, and scavenging birds are especially at risk from consumption of carcasses of animals killed with lead ammunition. Because current methods to identify lead exposure require animal capture and blood collection, management would benefit from the development of a less costly and noninvasive behavioral test for illness in wild animals. We attempted to design such a test to identify lead exposure in California condors ( Gymnogyps californianus ) that we tracked with global positioning system (GPS) telemetry in southern California, USA, 2013–2016. We measured blood-lead concentrations in tracked birds and expected that flight behavior would be influenced by lead exposure; thus, we measured the effect of blood-lead concentrations on 2 different types of movement rates and on the proportion of time condors spent in flight. We found no effect of lead exposure on any of these 3 behavioral metrics. Our work suggests that the measurements we took of flight behaviors were not a useful tool in predicting lead exposure in the mildly to moderately exposed birds we studied. Wild birds are effective at hiding illness, especially condors who have a strong social hierarchy in which showing weakness is a disadvantage. However, focusing on behaviors other than flight, expanding the sample studied to include birds with a wider range of lead concentration values, or analyzing tissues such as feathers (rather than, or in addition to, blood) may be more useful for identification of lead exposure and other diseases that may limit wildlife populations. © 2017 This article is a U.S. Government work and is in the public domain in the USA.

Journal of Wildlife Management

Predicting climate change impacts on poikilotherms using physiologically guided species abundance models

Poikilothermic animals comprise most species on Earth and are especially sensitive to changes in environmental temperatures. Species conservation in a changing climate relies upon predictions of species responses to future conditions, yet predicting species responses to climate change when temperatures exceed the bounds of observed data is fraught with challenges. We present a physiologically guided abundance (PGA) model that combines observations of species abundance and environmental conditions with laboratory-derived data on the physiological response of poikilotherms to temperature to predict species geographical distributions and abundance in response to climate change. The model incorporates uncertainty in laboratory-derived thermal response curves and provides estimates of thermal habitat suitability and extinction probability based on site-specific conditions. We show that temperature-driven changes in distributions, local extinction, and abundance of cold, cool, and warm-adapted species vary substantially when physiological information is incorporated. Notably, cold-adapted species were predicted by the PGA model to be extirpated in 61% of locations that they currently inhabit, while extirpation was never predicted by a correlative niche model. Failure to account for species-specific physiological constraints could lead to unrealistic predictions under a warming climate, including underestimates of local extirpation for cold-adapted species near the edges of their climate niche space and overoptimistic predictions of warm-adapted species.

Minnesota

Droughts reshape apex predator space use and intraguild overlap

1. Droughts are increasing in frequency and severity globally due to climate change, leading to changes in resource availability that may have cascading effects on animal ecology. Resource availability is a key driver of animal space use, which in turn influences interspecific interactions like intraguild competition. Understanding how climate-induced changes in resource availability influence animal space use, and how species-specific responses scale up to affect intraguild dynamics, is necessary for predicting broader community-level responses to climatic changes. 2. Although several studies have demonstrated the ecological impacts of drought, the behavioral responses of individuals that scale up to these broader-scale effects are not well known, particularly among animals in top trophic levels, such as large carnivores. Furthermore, we currently lack understanding of how the impacts of climate variability on individual carnivore behavior are linked to intraguild dynamics, in part because multi-species datasets collected at timescales relevant to climatic changes are rare. 3. Using 11 years of GPS data from four sympatric large carnivore species in southern Africa – lions ( Panthera leo ), leopards ( Panthera pardus ), African wild dogs ( Lycaon pictus) , and cheetahs ( Acinonyx jubatus ) – spanning 4 severe drought events, we test whether drought conditions impact 1) large carnivore space use, 2) broad-scale intraguild spatial overlap, and 3) fine-scale intraguild interactions. 4. Drought conditions expanded space use across species, with carnivores increasing their monthly home range sizes by 35% (wild dogs) to 66% (leopards). Drought conditions increased the amount of spatial overlap between lions and subordinate felids (cheetahs and leopards) by up to 119%, but only lion-cheetah encounter rates were affected by these changes, declining in response to drought. 5. Our findings reveal that drought has a clear signature on the space use of multiple sympatric large carnivore species, which can alter spatiotemporal partitioning between competing species. Our study thereby illuminates the links between environmental change, animal behavior, and intraguild dynamics. While fine-scale avoidance strategies may facilitate intraguild coexistence during periodic droughts, large carnivore conservation may require considerable expansion of protected areas or revised human-carnivore coexistence strategies to accommodate the likely long-term increased space demands of large carnivores under projected increases in drought intensity.

Okavango Delta

“Good” and “bad”: Human perceptions of and interactions with urban wildlife

Urban environments offer habitat for many species of animals. Although some of those are ubiquitous and/or undesirable, others are native and in some cases, of conservation value. In many cases, urban wildlife populations are a source of enjoyment for human residents, who sometimes invest considerable amounts in attracting them to yards and public spaces. Their presence there can serve an important educational role that helps protect non-urban habitats and species. Nonetheless, urban wildlife must survive what has been termed a “landscape of fear.” Although some of the urban wildlife that do well in this environment are benign, other populations – sometimes of a species that, in other locations, is iconic and desirable – can become problematic. Some species can serve as vectors that carry important zoonosis, such as the plague or diseases that affect other wildlife. Others can create noise or olfactory nuisances and degrade structures or usability of public spaces. Some pose hazards at busy airports, whereas still others may present an envenomation or predation risk on unwary humans. Here, we review the role that reptiles, birds, and mammals play in urban environments and discuss how urban wildlife rehabilitation centers help address some related issues. We close by looking ahead and trying to predict how global patterns such as increased urbanization and population growth may affect urban wildlife and its value for conservation.

Book chapter

Distribution and habitat use by manatees (Trichechus manatus manatus) in Belize and Chetumal Bay, Mexico

The nearshore coastal areas of Belize and of Chetumal Bay, Mexico, support one of the largest populations of manatees ( Trichechus manatus manatus ) in the Caribbean. In order to further document the distribution, relative abundance, habitat associations, and status of this population, we conducted three aerial surveys. The flights were done in January 1994, May 1994, and January 1995. Total manatee counts for each survey were 266, 207, and 171, respectively. Calves represented 7.4% of the grand total. Solitary manatees represented 62.8% of the total count. Manatees were numerous around the cays east of Belize City, in Placentia and Indian Hill lagoons, and in Chetumal Bay. Most manatees were observed in lagoons and rivers. Growing tourist activities should be assessed to avoid undesirable effects on manatees. Poaching and marketing of manatee products continues to jeopardize the animals in some areas. Current research and management activities are part of a cooperative international program to conserve the manatee in the Caribbean region.

Chetumal Bay

Space use of killdeer at a Great Basin breeding area

Wetland conservation efforts require knowledge of space use by a diversity of waterbirds. However, determining space use of animals requires intensive monitoring of individual organisms. Often, activity patterns during much of the annual cycle are neglected in analyses of home range and habitat use. From 1995-97, we monitored space use in a population of individually marked killdeer ( Charadrius vociferus ) that breed, and reside for a number of additional months each year, in the western Great Basin. We used linear distance measures and home range-area estimates, derived by fixed-kernel methods, to examine patterns of space use of adults prior to, during, and following nesting. Overall, killdeer used a local area of approximately 6 ha. Birds remained closer to nests while tending eggs than either before or after nesting, although extensive movements away from the nest were observed during all time periods. Females tended to move farther from nests than did males. Birds nesting farther from water bodies were generally observed at greater distances from nests than those nesting closer to shorelines during all time periods. Twenty-seven percent of individuals were observed greater than 1 km from nest locations, particularly during postnesting periods. During nesting periods, males were less likely to be observed at longer distances from nests than were females. There were no differences in home range size based upon sex, time period, or distance from water. We suggest that home range size may not always accurately measure differences in space use and that multiple measures, including distance from nests, should be considered. Understanding the sedentary nature of killdeer during much of the annual cycle and their intense use of local areas is important for making management decisions. Coupled with data on other shorebird species, this information also has broader implications for management of wetland systems by indicating large scale spatial and habitat requirements and the connectivity of geographically distinct habitat patches.

California, Idaho, Nevada, Oregon, Washington

Northern Prairie Wildlife Research Center

The Northern Prairie Wildlife Research Center (NPWRC) conducts integrated research to fulfill the Department of the Interior's responsibilities to the Nation's natural resources. Located on 600 acres along the James River Valley near Jamestown, North Dakota, the NPWRC develops and disseminates scientific information needed to understand, conserve, and wisely manage the Nation's biological resources. Research emphasis is primarily on midcontinental plant and animal species and ecosystems of the United States. During the center's 40-year history, its scientists have earned an international reputation for leadership and expertise on the biology of waterfowl and grassland birds, wetland ecology and classification, mammalian behavior and ecology, grassland ecosystems, and application of statistics and geographic information systems. To address current science challenges, NPWRC scientists collaborate with researchers from other U.S. Geological Survey centers and disciplines (Biology, Geography, Geology, and Water) and with biologists and managers in the Department of the Interior (DOI), other Federal agencies, State agencies, universities, and nongovernmental organizations. Expanding upon its scientific expertise and leadership, the NPWRC is moving in new directions, including invasive plant species, restoration of native habitats, carbon sequestration and marketing, and ungulate management on DOI lands.

Fact Sheet

Monitoring the southwestern Wyoming landscape—A foundation for management and science

Natural resource monitoring involves repeated collections of resource condition data and analyses to detect possible changes and identify underlying causes of changes. For natural resource agencies, monitoring provides the foundation for management and science. Specifically, analyses of monitoring data allow managers to better understand effects of land-use and other changes on important natural resources and to achieve their conservation and management goals. Examples of natural resources monitored on public lands include wildlife habitats, plant productivity, animal movements and population trends, soil chemistry, and water quality and quantity. Broader definitions of monitoring also recognize the need for scientifically valid data to help support planning efforts and informed decisions, to develop adaptive management strategies, and to provide the means for evaluating management outcomes.

Wyoming

An animal location-based habitat suitability model for bighorn sheep and wild horses in Bighorn Canyon National Recreation Area and the Pryor Mountain Wild Horse Range, Montana, and Wyoming

The purpose of this habitat suitability model is to provide a tool that will help managers and researchers better manage bighorn sheep and wild horses in the Bighorn Canyon National Recreation Area (BICA) and Pryor Mountain Wild Horse Range (PMWHR). A concern in the management of the Pryor Mountain wild horse population is whether or not the wild horses compete with bighorn sheep for available forage or available space. Two studies have been conducted that have shown no obvious, convincing competition between the two species. A study of diets and habitat-use of both species revealed substantial diet overlap only during some seasons, but there were considerable spatial and habitat separations between wild horses and bighorns during all seasons (Kissell and others, 1996). This empirical data was then used in a modeling exercise that predicted that neither the current (about 160 horses at the time of the analysis) nor larger numbers of wild horses on the area (e.g., about 200 horses) would result in reduced numbers or condition of bighorn sheep (Coughenour 1999). But competition is a very complex biological process to document. Bighorns might have already been spatially avoiding wild horses when these studies were conducted. A second concern for managers is that earlier studies suggest both species are not using many areas of the range that appear to be suitable (Gudorf and others, 1996; Kissell and others, 1996). A primary goal for the management of both species is to increase their numbers for purposes of genetic conservation and viability. The bighorn sheep population declined during the mid-1990’s from a peak of about 211 animals to ~ 100 animals at present. Absolute minimum goals for genetic viability in the bighorn sheep herd (genetic effective population size of N >50) suggest at least 150 animals should be present, while studies of persistence suggest populations of 250+ are e more likely to recover rapidly and persist should the population experience an epizootic die-off (Singer and others, 2001). Since all bighorn sheep populations are potentially vulnerable to disease epizootics, managing for larger populations of 200–300 animals appears to increase the potential for long-term persistence (Berger, 1990; Singer and others, 2001). Wild horses are not prone to rapid disease die-offs. However, minimum goals for genetic viability in the Pryor Mountain wild horses ( Ne > 50) require that at least 160 animals be present on the range (Singer and others, 2000). Since the Ne > 50 goal is set for the breeding of domestic animals, and since the vagaries of drought, severe winters, predation, and other stochastic events cause stress in wild animals, larger goals for Ne (e.g. Ne > 100) for wild horses are even more desirable (USDI, BLM, 1999; Gross, 2000). Expanding the area of the wild horse range is one option, but the prospects for expanding the range do not appear to be great (L. Coates-Markle, BLM, oral comm.). A second option would be to increase the amount of useable habitat for horses on the existing range. One goal of this modeling effort was to use GIS-based habitat analyses to determine the reason wild horses are not using some areas of the range, and to explore the potential for making some of these areas useable. The National Park Service (NPS) has shown considerable interest in management actions within BICA that will increase the range, useable habitat, and population size of bighorn sheep. There has also been interest expressed by the Bureau of Land Management (BLM) and wild horse advocates to improve the useable habitat for wild horses and to possibly increase the size of the horse range.

Report

Effects of prescribed fire on prenesting movements of wild turkeys in Arkansas

The restoration of historic disturbance regimes is an increasingly common management strategy to conserve disturbance-dependent communities and species, and enhance resilience of ecosystems to climate change or plant and animal invasions. However, the reintroduction of frequent and wide-scale disturbance may have unexpected consequences on species that are accustomed to an environment absent of frequent disturbance. Implementation of frequent prescribed fire for community restoration has coincided with population declines of eastern wild turkeys ( Meleagris gallopavo silvestris ) in the Ozark Highlands of Arkansas. We investigated whether there was evidence of linkage between wild turkey movement behavior and habitat use and the widespread use of prescribed fire that began in 2002 and continued through the end of our study in 2013. We fitted 67 female wild turkeys with satellite transmitters in 2012 and 2013 on the White Rock Ecosystem Restoration Area to estimate annual and seasonal home ranges and examine pre-nesting habitat use at multiple spatial scales. Home range was larger for adults in 2013 (4,750 ha, SE = 946.2, n = 12) than in 2012 (2,703 ha, SE = 283.4, n = 19). Mean pre-nesting ranges for adults were 1,633 ha (SE = 238.9, n = 23) in 2012 and 1,118 ha (SE = 158.8, n = 18) in 2013, smaller than those of sub-adults, 1,761 ha (SE = 581.2, n = 3) in 2012 and 5,576 ha (SE = 2260, n = 8) in 2013. Habitat selection analyses for the pre-nesting period indicated that wild turkeys used locations with more variability in canopy cover and vegetation height and in smaller patch sizes than expected based on availability. However, female wild turkeys selected nesting cover that had more variability in canopy cover and vegetation height but in larger patches. Differences in sub-adults and adult home and seasonal ranges suggest that other factors such as density dependence or social hierarchy could be driving movements as opposed to prescribed fire application. Wild turkey females selected variable vegetation structure during the pre-nesting and nesting periods with characteristics that could be a result of frequent prescribed fire or forest succession. However, we documented no direct evidence these characteristics or other aspects of the prescribed fire program were causing wild turkey population declines.

Arkansas

Association, roost use and simulated disruption of Myotis septentrionalis maternity colonies

How wildlife social and resource networks are distributed on the landscape and how animals respond to resource loss are important aspects of behavioral ecology. For bats, understanding these responses may improve conservation efforts and provide insights into adaptations to environmental conditions. We tracked maternity colonies of northern bats ( Myotis septentrionalis ) at Fort Knox, Kentucky, USA to evaluate their social and resource networks and space use. Roost and social network structure differed between maternity colonies. Overall roost availability did not appear to be strongly related to network characteristics or space use. In simulations for our two largest networks, roost removal was related linearly to network fragmentation; despite this, networks were relatively robust, requiring removal of >20% of roosts to cause network fragmentation. Results from our analyses indicate that northern bat behavior and space use may differ among colonies and potentially across the maternity season. Simulation results suggest that colony social structure is robust to fragmentation caused by random loss of small numbers of roosts. Flexible social dynamics and tolerance of roost loss may be adaptive strategies for coping with ephemeral conditions in dynamic forest habitats.

Behavioural Processes

Use of spatial capture-recapture modeling and DNA data to estimate densities of elusive animals

Assessment of abundance, survival, recruitment rates, and density (i.e., population assessment) is especially challenging for elusive species most in need of protection (e.g., rare carnivores). Individual identification methods, such as DNA sampling, provide ways of studying such species efficiently and noninvasively. Additionally, statistical methods that correct for undetected animals and account for locations where animals are captured are available to efficiently estimate density and other demographic parameters. We collected hair samples of European wildcat ( Felis silvestris ) from cheek-rub lure sticks, extracted DNA from the samples, and identified each animals' genotype. To estimate the density of wildcats, we used Bayesian inference in a spatial capture-recapture model. We used WinBUGS to fit a model that accounted for differences in detection probability among individuals and seasons and between two lure arrays. We detected 21 individual wildcats (including possible hybrids) 47 times. Wildcat density was estimated at 0.29/km 2 (SE 0.06), and 95% of the activity of wildcats was estimated to occur within 1.83 km from their home-range center. Lures located systematically were associated with a greater number of detections than lures placed in a cell on the basis of expert opinion. Detection probability of individual cats was greatest in late March. Our model is a generalized linear mixed model; hence, it can be easily extended, for instance, to incorporate trap- and individual-level covariates. We believe that the combined use of noninvasive sampling techniques and spatial capture-recapture models will improve population assessments, especially for rare and elusive animals.

Conservation Biology

Great gray owls ( Strix nebulosa ) in Yosemite National Park: on the importance of food, forest structure, and human disturbance

We studied great gray owls (Strix nebulosa Forster) in Yosemite National Park, California, measuring variables that could potentially influence patterns of occurrence and conservation of this stateendangered species. We found that owl presence was closely tied to habitat (red fir (Abies magnified A. Murray) and the abundance of meadows), prey, and snags across the landscape. We also found that indicators of human recreational activities negatively influenced owl distribution and habitat use. Great gray owls appear to prefer mid-elevation red fir forest with meadows that are drier and more productive in terms of small mammal populations. That these areas also have the highest human activity presents a paradox, both for individual owls and for the future conservation and management of this California endangered species. The extent to which human recreation in natural areas affects animal behavior, species distribution, and productivity is a growing issue in natural area management. We present information that will allow land managers to better understand how existing natural resources, coupled with human recreation, influence the distribution and habitat use of the great gray owl.

California

Fulfilling a paradoxical mandate: can the Environmental Water Account ensure the reliability of freshwater exports from the Sacramento-San Joaquin Delta and simultaneously protect delta smelt (Hypomesus transpacificus) from excessive entrainment?

The San Francisco Estuary (SFE) is often defined by its extremes. It is considered one of the most urbanized estuaries in the world (Conomos 1979, Nichols et al. 1986), and one of the most invaded estuaries in the United States, with hundreds of aquatic nonindigenous species established throughout the system (Cohen and Carlton 1995, Dill and Cordone 1997, Kimmerer and Orsi 1996). It is also one of the most managed estuaries, particularly in relation to freshwater inflow, water circulation, and water quality (Jassby and Powell 1994, CSWRCB 1995, Arthur et al. 1996, Kimmerer 2002). Despite this high level of disturbance, the SFE is one of the most valuable natural resources in the western United States (CALFED 2000). The SFE provides important habitat for numerous native plant and animal species, many of special concern, as well as several species with sport and commercial value (CALFED 2000). Conserving and restoring estuarine habitat and natural resources is a pressing and complex challenge for the responsible government agencies because human water needs continue to increase in concert with continuing urbanization of the watershed.

Interagency Ecological Program Newsletter

Targeting ecosystem features for conservation: Standing crops in the Florida Everglades

The Everglades in southern Florida, U.S.A., is a major focus of conservation activities. The freshwater wetlands of the Everglades do not have high species richness, and no species of threatened aquatic animals or plants live there. We have, however, identified a distinctive ecological feature of the Everglades that is threatened by canal construction, draining, and nutrient enrichment from agricultural runoff compared to values reported from other freshwater systems, standing stocks of periphyton in relatively undisturbed areas of the Everglades were unusually high, and standing stocks of invertebrates and fish were unusually low. Averaging data gathered from nine sites and five sampling periods spanning I year, we found that periphyton standing crop was 88.2 g/m2 (ash-free dry mass), invertebrate standing stock was 0.64 g/m2 (dry mass), and fish standing stock was 1.2 g/m2 (dry mass of large and small species combined). We found that fish standing stocks were much higher in phosphorus-enriched sites than in nearby reference sites but that invertebrate standing stocks were similar in enriched and reference sites. Our results support the notion that oligotrophy is at least partially responsible for the low standing stocks of fish, but they also suggest that species interactions and a paucity of deep-water refugia are important. Anthropogenic eutrophication in Everglades marshes will lead to the loss of distinctive ecosystem features. A focus on species richness and 'hot spots' of threatened species provides no basis for conservation of ecosystems like the Everglades. If oligotrophic ecosystems often have low species richness, they will be underrepresented in preservation networks based on some common criteria for establishing conservation priorities.

Conservation Biology

Selecting islands and shoals for conservation based on biological and aesthetic criteria

Consideration of biological quality has long been an important component of rating areas for conservation. Often these same areas are highly valued by people for aesthetic reasons, creating demands for housing and recreation that may conflict with protection plans for these habitats. Most methods of selecting land for conservation purposes use biological factors alone. For some land areas, analysis of aesthetic qualities is also important in describing the scenic value of undisturbed land. A method for prioritizing small islands and shoals based on both biological and visual quality factors is presented here. The study included 169 undeveloped islands and shoals a??0.8 ha in the Thousand Islands Region of the St. Lawrence River, New York. Criteria such as critical habitat for uncommon plant and animal species were considered together with visual quality and incorporated into a rating system that ranked the islands and shoals according to their priority for conservation management and protection from development. Biological factors were determined based on previous research and a field survey. Visual quality was determined by visual diagnostic criteria developed from public responses to photographs of a sample of islands. Variables such as elevation, soil depth, and type of plant community can be used to classify islands into different categories of visual quality but are unsuccessful in classifying islands into categories of overall biological quality.

Environmental Management