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At least 289 records · Page 16Linked to original sources

Managing native predators: Evidence from a partial removal of raccoons ( Procyon lotor ) on the Outer Banks of North Carolina, USA

Raccoons ( Procyon lotor ) are important predators of ground-nesting species in coastal systems. They have been identified as a primary cause of nest failure for the American Oystercatcher ( Haematopus palliatus ) throughout its range. Concerns over the long-term effects of raccoon predation and increased nest success following a hurricane inspired a mark-resight study of the raccoon population on a barrier island off North Carolina, USA. Approximately half of the raccoons were experimentally removed in 2008. Nests ( n = 700) were monitored on two adjacent barrier islands during 2004–2013. Daily nest survival estimates were highest for 2004 (0.974 ± 0.005) and lowest for 2007 and 2008 (0.925 ± 0.009 and 0.925 ± 0.010, respectively). The only model in our candidate set that received any support included island and time of season, along with a diminishing effect of the hurricane and a constant, 5-year effect of the raccoon removal. For both hurricane and raccoon removal, however, the support for island-specific effects was weak (β = -0.204 ± 0.116 and 0.146 ± 0.349, respectively). We conclude that either the raccoon reduction was inadequate, or factors other than predation cause more variation in nest success than previously recognized. A multi-faceted approach to management aimed at reducing nest losses to storm overwash, predation, and human disturbance is likely to yield the largest population level benefits.

North Carolina

Apparent predation by cattle at grassland bird nests

We document the first cases of cattle behaving as avian predators, removing nestlings and eggs from three active ground nests in continuously grazed pastures in southwestern Wisconsin, 2000–2001. Cows removed three of four Savannah Sparrow ( Passerculus sandwichensis ) eggs from one nest (the fourth egg was damaged), all four Eastern Meadowlark ( Sturnella magna ) nestlings from another, and all three Savannah Sparrow nestlings from a third. We found only two of three missing eggs (intact) and one of seven missing nestlings (dead) near two of the nests. Cows may have eaten the egg and nestlings we were unable to account for; alternatively, the egg and nestlings may have been scavenged by predators or removed from the area by the adult birds. Without videotape documentation, we would have attributed nest failure to traditional predators and cattle would not have been implicated. We may be underestimating the impact of cattle on ground nests by not considering cattle as potential predators.

The Wilson Bulletin

Do predators control prey species abundance? An experimental test with brown treesnakes on Guam

The effect of predators on the abundance of prey species is a topic of ongoing debate in ecology; the effect of snake predators on their prey has been less debated, as there exists a general consensus that snakes do not negatively influence the abundance of their prey. However, this viewpoint has not been adequately tested. We quantified the effect of brown treesnake ( Boiga irregularis ) predation on the abundance and size of lizards on Guam by contrasting lizards in two 1-ha treatment plots of secondary forest from which snakes had been removed and excluded vs. two 1-ha control plots in which snakes were monitored but not removed or excluded. We removed resident snakes from the treatment plots with snake traps and hand capture, and snake immigration into these plots was precluded by electrified snake barriers. Lizards were sampled in all plots quarterly for a year following snake elimination in the treatment plots. Following the completion of this experiment, we used total removal sampling to census lizards on a 100-m 2 subsample of each plot. Results of systematic lizard population monitoring before and after snake removal suggest that the abundance of the skink, Carlia ailanpalai, increased substantially and the abundance of two species of gekkonids, Lepidodactylus lugubris and Hemidactylus frenatus , also increased on snake-free plots. No treatment effect was observed for the skink Emoia caeruleocauda . Mean snout–vent length of all lizard species only increased following snake removal in the treatment plots. The general increase in prey density and mean size was unexpected in light of the literature consensus that snakes do not control the abundance of their prey species. Our findings show that, at least where alternate predators are lacking, snakes may indeed affect prey populations.

Ecology

Small nonnative fishes as predators of larval razorback suckers

The razorback sucker (Xyrauchen texanus), an endangered big-river fish of the Colorado River basin, has demonstrated no sustainable recruitment in 4 decades, despite presence of spawning adults and larvae. Lack of adequate recruitment has been attributed to several factors, including predation by nonnative fishes. Substantial funding and effort has been expended on mechanically removing nonnative game fishes, typically targeting large predators. As a result, abundance of larger predators has declined, but the abundance of small nonnative fishes has increased in some areas. We conducted laboratory experiments to determine if small nonnative fishes would consume larval razorback suckers. We tested adults of three small species (threadfin shad, Dorosoma petenense; red shiner, Cyprinella lutrensis; fathead minnow, Pimephales promelas) and juveniles of six larger species (common carp, Cyprinus carpio; yellow bullhead, Ameiurus natalis; channel catfish, Ictalurus punctatus; rainbow trout, Oncorhynchus mykiss; green sunfish, Lepomis cyanellus; bluegill, L. macrochirus). These nonnative fishes span a broad ecological range and are abundant within the historical range of the razorback sucker. All nine species fed on larval razorback suckers (total length, 9-16 mm). Our results suggest that predation by small nonnative fishes could be responsible for limiting recovery of this endangered species.

Southwestern Naturalist

Factors determining Pochard nest predation along a wetland gradient

Waterfowl management on breeding grounds focuses on improving nest success, but few studies have compared waterfowl nest success and factors affecting nest survival along a wetland gradient and simultaneously identified nest predators. We monitored nests (n = 195) of common pochards ( Aythya ferina ) in Trebon Basin Biosphere Reserve, Czech Republic, during 1999–2002. Daily nest survival rates (DSRs, logistic-exposure) declined from island (0.985, 95% confidence interval, 0.978–0.991) to overwater (0.962, 0.950–0.971) and terrestrial (0.844, 0.759–0.904) nests. The most parsimonious model for DSRs included habitat class (DSRs: island > overwater > terrestrial) and nest visibility. Nest survival was improved by reduced nest visibility, increased water depth, and increased distance from the nest to habitat edge in littoral habitats. On islands, nest success increased with advancing date and increased distance to open water. A model of constant nest survival best explained the data for terrestrial nests. There were no observer effects on DSRs in any habitat. In 2003, artificial nests ( n = 180; 120 contained a wax-filled egg) were deployed on study plots. The model that best explained variation in DSRs for artificial nests included only 1 variable: habitat class (DSRs: island ≥ overwater > terrestrial). Mammalian predation of artificial nests (by foxes [ Vulpes vulpes ] and martens [ Martes spp.]) was more likely in terrestrial habitats than in littoral habitats or on islands. By contrast, corvids and marsh harriers ( Circus aeruginosus ) prevailed among predators of overwater and island nests. Our data indicate that artificial islands and wide strips of littoral vegetation may represent secure breeding habitats for waterfowl because those habitats allow nests to be placed in areas that are not accessible to, or that are avoided by, mammalian predators. Management actions should be aimed at preserving these habitats. This, along with creation of new artificial islands, could help to enhance breeding productivity of pochards and possibly other waterfowl species inhabiting man-made ponds.

Trebon Basin Biosphere Reserve

Does nonrandom nest placement imply nonrandom nest predation?: A reply

In response to the critique by Schmidt and Whelan (Condor 101(4):916-920, 1999), we find that the relationship between nest success and tree selectivity is dependent upon inclusion or exclusion of particular tree species, whether or not years are pooled, and the selectivity index used. We question their use of point estimates of nest success with extremely high variances, defend our index, question the application of the Chesson (1983) index to our data, and explain the need to analyze years separately. Bottomland hardwood forest systems are extremely variable; hydroperiods alter the suitability of nesting substrates, availability of alternative food, and behavior of predators and their prey. Given these features, actively searching for Acadian Flycatcher (Empidonax virescens) nests is seldom an efficient predator foraging strategy. Therefore, these predation events are best described as random; nests are principally encountered opportunistically by generalist predators while searching for other prey.

Condor

Upland duck nesting related to land use and predator reduction

Duck nesting was studied during 1971 in north-central South Dakota under four conditions: in idle, five or six year old fields of domestic grass-legume mixtures in an area where predators including the red fox ( Vulpes fulva ), raccoon ( Procyon lotor ), striped skunk ( Mephitis mephitis ), and badger ( Taxidea taxus ) were (1) reduced and (2) not reduced. Nesting was also studied in tracts of active agricultural land (primarily croplands and pastures) where predators were (3) reduced, and (4) not reduced. Under condition (1), 260 nests were found on 0.87 km 2 (299 nests/km 2 ), eggs hatched in 92 percent of the nests and production was 22.0 ducklings/hectare. Under condition (2), 187 nests were found on 2.22 km 2 (84 nests/km 2 ), nest success was 68 percent and 4.7 ducklings/hectare were produced. On active agricultural land subject to predator reduction (condition 3), 64 nests were found on 5.14 km 2 (12 nests/km 2 ). Eggs in 85 percent of the nests hatched and production was 0.7 duckling/hectare. On active agricultural land not subject to predator reduction (condition 4), 58 nests were found on 4.01 km 2 (14 nests/km 2 ), nest success was 51 percent and 0.5 duckling/hectare was produced. Idle, 16 to 65-hectare (40 to 160-acre) stands of cool-season, introduced grasses in combination with legumes produced maximum numbers of upland nesting ducks.

South Dakota

Relationship between snow depth and gray wolf predation on white-tailed deer

Survival of 203 yearling and adult white-tailed deer ( Odocoileus virginianus ) was monitored for 23,441 deer days from January through April 1975-85 in northeastern Minnesota. Gray wolf ( Canis lupus ) predation was the primary mortality cause, and from year to year during this period, the mean predation rate ranged from 0.00 to 0.29. The sum of weekly snow depths/month explained 51% of the variation in annual wolf predation rate, with the highest predation during the deepest snow.

Minnesota

Distilling professional opinion to gauge vulnerability of Guam avifauna to Brown Treesnake predation

The avifauna of Guam was devastated by the introduction of the Brown Treesnake, and the restoration of native birds would need to address the problem with eradication or suppression of BTS. With eradication of the snake unlikely in the near term, and suppression capabilities limited to specific finite areas, key information for reintroductions is how low BTS abundance will likely need to be for each bird species to be re-established based on their vulnerability to BTS predation. Here, we estimate vulnerability, which can no longer be measured directly, so biologists who are familiar with one or more of seven Guam birds were surveyed to obtain their knowledge and produce quantitative vulnerability estimates. As is typical of birds adapted to islands devoid of predators, respondents judged that our focal species exhibit few predator avoidance and tolerance traits, leaving body size as the prime determinant of vulnerability. Respondent opinion also holds that any behavior that reduces the likelihood of an encounter by BTS, e.g., roosting/nesting in palm crowns, cavity nesting, and in particular urban dwelling, substantially reduces vulnerability. Our results can help inform species-specific decisions about when it may be safe to consider the release of birds on Guam depending on the relative vulnerability of each species to predation by BTS.

Guam

Spatiotemporal risk avoidance varies seasonally, relative to risk intensity, in a reestablishing predator–prey system

Predation establishes risk, which can indirectly influence prey behavior and ecology. We evaluated the influence of Mexican gray wolves ( Canis lupus baileyi ) on habitat selection and spatiotemporal predator avoidance strategies of elk ( Cervus canadensis ). We fit 866 adult female elk with GPS collars across areas of varying wolf densities within the Mexican wolf experimental population area of eastern Arizona and western New Mexico between 2019−2021. Using step-selection functions we examined relative intensity of elk use in relation to landscape attributes, estimated predator/prey diel activity, and measures of risk. Risk metrics included predicted wolf presence, habitat openness, and predicted risky places modeled from attributes of locations where wolves killed elk. Wolf activity varied across seasons and increased midday and night in fall and monsoon seasons. Relative use by elk was best explained by incorporating an interaction between diel period and predicted risky places across all seasons. Elk utilized risky places more in times of nutritional deficit associated with high energetic demands of the third trimester pregnancy and lactation and when forage quality was best, during spring and monsoon season. Particularly, use of risky places increased at less risky times in areas with more established wolf presence, suggesting use of risky places varied relative to exposure to Mexican wolves. These behaviors highlight the importance of temporal avoidance when predators and prey are highly mobile and largely overlap in space. Our research suggests temporally responding to predictable and relatively static environmental characteristics associated with encounter and kill rates may better balance energetic trade-offs than anticipating changes in wolf activity or spatially avoiding areas with higher wolf presence. Thus, elk appear to be more willing to take chances and mitigate cursorial predation risk with a more immediate, reactive approach and make proactive trade-offs during the seasons they can best increase fitness.

Arizona, New Mexico

Behaviors of southwestern native fishes in response to introduced catfish predators

Native fishes reared in hatcheries typically suffer high predation mortality when stocked into natural environments. We evaluated the behavior of juvenile bonytail Gila elegans, roundtail chub Gila robusta, razorback sucker Xyrauchen texanus, and Sonora sucker Catostomus insignis in response to introduced channel catfish Ictalurus punctatus and flathead catfish Pylodictis olivaris. Our laboratory tests indicate these species did not inherently recognize catfish as a threat, but they can quickly (within 12 h) change their behavior in response to a novel predator paired with the sight and scent of a dead conspecific. Chubs appear to avoid predation by swimming away from the threat, whereas suckers reduced movement. Effects of antipredator conditioning on survival of fish reared in hatcheries is unknown; however, our results suggest some native fish can be conditioned to recognize introduced predators, which could increase poststocking survival.

Journal of Fish and Wildlife Management

Response of corvid nest predators to thinning: implications for balancing short- and long-term goals for restoration of forest habitat

Forest thinning on public lands in the Pacific Northwest USA is an important tool for restoring diversity in forest stands with a legacy of simplified structure from decades of intensive management for timber production. A primary application of thinning in young (< 50-year-old) stands is to accelerate forest development to mitigate loss of late-seral habitat to decades of logging. However, thinning may have short-term negative effects for some species associated with mature forest that are expected to benefit from the practice over the long term. An increased risk of nest predation is a primary concern to managers charged with stewardship of habitat for the federally threatened Marbled Murrelet ( Brachyramphus marmoratus ), a species that nests in older forests. Predation by corvids is the greatest cause of nest failure for the Marbled Murrelet, and corvids are known to respond positively to forest disturbance, but quantitative information is lacking on the potential impacts of thinning on risk of nest predation. We investigated the response of two common corvid nest predators, Steller’s Jay ( Cyanocitta stelleri ) and Canada Jay ( Perisoreus canadensis ), to variation in thinning intensity in young forest (< 50 years old) using data from a long-term silviculture experiment. We used a Before-After-Control-Impact (BACI) design, linear mixed modeling, and occupancy modeling to quantify differences in corvid observation rates among varying levels of thinning intensity, and to assess changes in jay response over more than a decade following thinning. We found an increase in observation rates of both species in the heavily thinned treatment during the first 5 to 7 years following thinning, and some evidence of a short-term increase in Steller’s Jay activity in the thinning-with-gaps treatment. Neither jay species responded to the least intensive thinning treatment, which reduced average canopy cover by < 30%. By approximately a decade after thinning, observation rates of jays did not differ between unthinned controls and any of the thinning treatments. Incorporating our quantitative information into landscape-level planning can help managers balance short- and long-term conservation goals.

Oregon

Reducing predation by common ravens on desert tortoises in the Mojave and Colorado Deserts

Conflicts between humans and natural populations often result from habitat fragmentation and degradation that accompanies human activities. Common raven populations in the Mojave Desert have benefited by human-provided resources; they've expanded precipitously in recent years. Because ravens prey on juveniles of the threatened desert tortoise, they have become the focus of management concerns to help recover dwindling tortoise populations. I have outlined herein a series of management recommendations designed to reduce raven predation on desert tortoises thereby facilitating juvenile tortoise recruitment into the population of reproductive adults. The recommendations are based on the best available scientific information and are intended to provide a basis for a long-term reduction in raven impacts. The recommendations fall into four basic categories. (1) Modify anthropogenic sources of food, water, and nesting substrates to reduce their use by ravens. This includes modifying landfill operations, septage containment practices, livestock management, and other commercial and private practices that help facilitate raven survival and dispersal by providing food and water. Most of these measures are long-term actions deigned to reduce the carrying capacity of the desert for ravens. This action is critical and must be done over very large areas. (2)Lethal removal of ravens by shooting or euthanizing following live trapping. Specific ravens known to prey on tortoises would be targeted as well as all ravens found foraging within specific high-priority desert tortoise management zones (e.g., Desert Tortoise Natural Areas, DTNA). These actions would primarily be deployed on a short-term emergency basis to give specific tortoise populations a necessary boost until other measures become fully implemented and achieve their goals. (3) Conduct research on raven ecology, raven behavior, and methods to reduce raven predation on tortoises. Results of these studies would be used to design future phases of the raven management program. (4) All actions should be approached within an adaptive management framework. As such monitor, actions should be designed as experiments so that monitoring of actions will yield reliable and scientifically sound results. Coordinating and oversight teams should be convened to facilitate cooperation and coordination among agencies and to ensure that the actions are being implemented effectively. Recommendations made herein were developed to help recover tortoise populations by reducing raven predation on juvenile tortoises. If the recommendations made are implemented in concert with actions reducing other causes of mortality, ill health, and lowered reproductive output, they should aid in the long-term recovery of desert tortoise populations. Many important aspects of raven population dynamics, raven predation on tortoises, and how to manage raven populations and behavior are as yet unknown. Because of this, any raven management program must be implemented within an adaptive management framework. Doing so would allow for sufficient flexibility to modify the program as new information is gained.

Report

An introduced predator alters Aleutian Island plant communities by thwarting nutrient subsidies

The ramifying effects of top predators on food webs traditionally have been studied within the framework of trophic cascades. Trophic cascades are compelling because they embody powerful indirect effects of predators on primary production. Although less studied, indirect effects of predators may occur via routes that are not exclusively trophic. We quantified how the introduction of foxes onto the Aleutian Islands transformed plant communities by reducing abundant seabird populations, thereby disrupting nutrient subsidies vectored by seabirds from sea to land. We compared soil and plant fertility, plant biomass and community composition, and stable isotopes of nitrogen in soil, plants, and other organisms on nine fox-infested and nine historically fox-free islands across the Aleutians. Additionally, we experimentally augmented nutrients on a fox-infested island to test whether differences in plant productivity and composition between fox-infested and fox-free islands could have arisen from differences in nutrient inputs between island types. Islands with historical fox infestations had soils low in phosphorus and nitrogen and plants low in tissue nitrogen. Soils, plants, slugs, flies, spiders, and bird droppings on these islands had low d15N values indicating that these organisms obtained nitrogen from internally derived sources. In contrast, soils, plants, and higher trophic level organisms on fox-free islands had elevated d15N signatures indicating that they utilized nutrients derived from the marine environment. Furthermore, soil phosphorus (but not nitrogen) and plant tissue nitrogen were higher on fox-free than fox-infested islands. Nutrient subsidized fox-free islands supported lush, high biomass plant communities dominated by graminoids. Fox-infested islands were less graminoid dominated and had higher cover and biomass of low-lying forbs and dwarf shrubs. While d15N profiles of soils and plants and graminoid biomass varied with island size and distance from shore, after accounting for these effects differences between fox-infested and fox-free islands still existed. Fertilization over four years caused a 24-fold increase in graminoid biomass and a shift toward a more graminoid dominated plant community typical of fox-free islands. These results indicate that apex predators can influence plant productivity and composition through complex interaction web pathways involving both top-down forcing and bottom-up nutrient exchanges across systems. ?? 2006 by the Ecological Society of America.

Ecological Monographs

Sitting ducklings: Timing of hatch, nest departure, and predation risk for dabbling duck broods

For ground‐nesting waterfowl, the timing of egg hatch and duckling departure from the nest may be influenced by the risk of predation at the nest and en route to wetlands and constrained by the time required for ducklings to imprint on the hen and be physically able to leave the nest. We determined the timing of hatch, nest departure, and predation on dabbling duck broods using small video cameras placed at the nests of mallard ( Anas platyrhynchos ; n = 26), gadwall ( Mareca strepera ; n = 24), and cinnamon teal ( Anas cyanoptera ; n = 5). Mallard eggs began to hatch throughout the day and night, whereas gadwall eggs generally started to hatch during daylight hours (mean 7.5 hr after dawn). Among all species, duckling departure from the nest occurred during daylight (98%), and 53% of hens typically left the nest with their broods 1–4 hr after dawn. For mallard and gadwall, we identified three strategies for the timing of nest departure: (a) 9% of broods left the nest the same day that eggs began to hatch (6–12 hr later), (b) 81% of broods left the nest the day after eggs began to hatch, and (c) 10% of broods waited 2 days to depart the nest after eggs began to hatch, leaving the nest just after the second dawn (27–42 hr later). Overall, eggs were depredated at 10% of nests with cameras in the 2 days prior to hatch and ducklings were depredated at 15% of nests with cameras before leaving the nest. Our results suggest that broods prefer to depart the nest early in the morning, which may best balance developmental constraints with predation risk both at the nest and en route to wetlands.

Ecology and Evolution

A spatially explicit modeling framework to guide management of subsidized avian predator densities

Anthropogenic resource subsidization across western ecosystems has contributed to widespread increases in generalist avian predators, including common ravens ( Corvus corax ; hereafter, raven). Ravens are adept nest predators and can negatively impact species of conservation concern. Predation effects from ravens are especially concerning for greater sage-grouse ( Centrocercus urophasianus ; hereafter, sage-grouse), which have experienced prolonged population decline. Our objectives were to quantify spatiotemporal patterns in raven density, evaluate sage-grouse nest success concurrent with fluctuating raven densities, and demonstrate a spatially explicit decision support tool to guide management applications to appropriate conflict areas. We combined ~28,000 raven point count surveys with data from more than 900 sage-grouse nests between 2009 and 2019 within the Great Basin, USA. We modeled variation in raven density using a Bayesian hierarchical distance sampling approach with environmental covariates on detection and abundance. Concurrently, we modeled sage-grouse nest survival using a hierarchical frailty model as a function of raven density and other environmental covariates that influence the risk of nest failure. Raven density commonly exceeded 0.5 ravens km −2 and increased at low elevations with more anthropogenic development and/or agriculture. Reduced sage-grouse nest survival was strongly associated with elevated raven density (e.g., >0.5 ravens km −2 ) and varied with topographic ruggedness, shrub cover, and burned areas. For conservation application, we developed a spatially explicit planning tool that predicts nest survival under current and reduced raven numbers within the Great Basin to help direct management actions to localized areas where sage-grouse nests are at highest risk of failure. Our modeling framework can be generalized to multiple species where spatially registered abundance and demographic data are available.

Nevada

Evaluating consumptive and nonconsumptive predator effects on prey density using field times series data

Determining the degree to which predation affects prey abundance in natural communities constitutes a key goal of ecological research. Predators can affect prey through both consumptive effects (CEs) and nonconsumptive effects (NCEs), although the contributions of each mechanism to the density of prey populations remain largely hypothetical in most systems. Common statistical methods applied to time series data cannot elucidate the mechanisms responsible for hypothesized predator effects on prey density (e.g., differentiate CEs from NCEs), nor provide parameters for predictive models. State space models (SSMs) applied to time series data offer a way to meet these goals. Here, we employ SSMs to assess effects of an invasive predatory zooplankter, Bythotrephes longimanus, on an important prey species, Daphnia mendotae, in Lake Michigan. We fit mechanistic models in a SSM framework to seasonal time series (1994-2012) using a recently developed, maximum likelihood-based optimization method, iterated filtering, which can overcome challenges in ecological data (e.g. nonlinearities, measurement error, and irregular sampling intervals). Our results indicate that B. longimanus strongly influences D. mendotae dynamics, with mean annual peak densities of B. longimanus observed in Lake Michigan estimated to cause a 61% reduction in D. mendotae population growth rate and a 59% reduction in peak biomass density. Further, the observed B. longimanus effect is most consistent with an NCE via reduced birth rates. The SSM approach also provided estimates for key biological parameters (e.g., demographic rates) and the contribution of dynamic stochasticity and measurement error. Our study therefore provides evidence derived directly from survey data that the invasive zooplankter B. longimanus is affecting zooplankton demographics and offer parameter estimates needed to inform predictive models that explore the effect of B. longimanus under different scenarios such as climate change.

Ecology

Informing management of recovering predators and their prey with ecological diffusion models

The reintroduction and recovery of predators can be ecologically beneficial as well as socially and economically controversial. However, the growth and expansion of predator populations, and thus their ecological, social, and economic impacts, are not static but rather they vary in space and time. We propose a spatiotemporal statistical modeling framework based on ecological diffusion to better inform the ecology and management of recovering predators and their prey. We demonstrate its utility by applying it to a recovering sea otter ( Enhydra lutris ) population in Southeast Alaska, where sea otters were reintroduced in the late 1960s and have exhibited unprecedented population growth. Estimated parameters yield inferences about movement and population ecology, and our approach provides useful derived quantities, such as local abundance and carrying capacity as well as a quantity we term the equilibrium differential. We used our model to examine how density dependence and carrying capacity of sea otters vary spatially across a region. The diffusion modeling approach we present can be generalized for use in other instances of (re)colonization across taxa to inform management and conservation efforts.

Alaska