Enhydra and Enhydriodon from the Pacific Coast of North America

Abstract

Two lineages of the "crab-eating" otter Erihydriodon , from the Old World Miocene and Pliocene, are suggested by the fossil record. One appears to lead to the late Pliocene Enhydriodon sivalensis from Villafranchian-equivalent beds in India and can be characterized by the presence of a parastyle on P 4 and by the location of the protocone of this tooth, which is located as far lingually as is the hypocone. In the other lineage no parastyle is developed on P 4 , the anteroposterior length of the carnassial blade is progressively reduced while the transverse width is increased, and the protocone of P 4 remains in its primitive position anterolateral to the hypocone. The second lineage, insofar as it is known, seems to lead toward the living sea otter Enhydra , particularly so because of the evolutionary direction shown by two species of Enhydriodon from the late Miocene and late Pliocene of California.