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Research about Yellowstone National Park

Source-linked reports with geographic coverage including Yellowstone National Park.

At least 127 records · Page 7Linked to original sources

Spatial population structure of Yellowstone bison

Increases in Yellowstone National Park, USA, bison ( Bison bison ) numbers and shifts in seasonal distribution have resulted in more frequent movements of bison beyond park boundaries and development of an interagency management plan for the Yellowstone bison population. Implementation of the plan under the adaptive management paradigm requires an understanding of the spatial and temporal structure of the population. We used polythetic agglomerative hierarchical cluster analysis of radiolocations obtained from free-ranging bison to investigate seasonal movements and aggregations. We classified radiolocations into 4 periods: annual, peak rut (15 Jul–15 Sep), extended rut (1 Jun–31 Oct), and winter (1 Nov–31 May). We documented spatial separation of Yellowstone bison into 2 segments, the northern and central herds, during all periods. The estimated year-round exchange rate (4.85–5.83%) of instrumented bison varied with the fusion strategy employed. We did not observe exchange between the 2 segments during the peak rut and it varied during the extended rut (2.15–3.23%). We estimated a winter exchange of 4.85–7.77%. The outcome and effectiveness of management actions directed at Yellowstone bison may be affected by spatial segregation and herd affinity within the population. Reductions based on total population size, but not applied to the entire population, may adversely affect one herd while having little effect on the other. Similarly, management actions targeting a segment of the population may benefit from the spatial segregation exhibited.

Wyoming↗

A simple solar radiation index for wildlife habitat studies

Solar radiation is a potentially important covariate in many wildlife habitat studies, but it is typically addressed only indirectly, using problematic surrogates like aspect or hillshade. We devised a simple solar radiation index (SRI) that combines readily available information about aspect, slope, and latitude. Our SRI is proportional to the amount of extraterrestrial solar radiation theoretically striking an arbitrarily oriented surface during the hour surrounding solar noon on the equinox. Because it derives from first geometric principles and is linearly distributed, SRI offers clear advantages over aspect-based surrogates. The SRI also is superior to hillshade, which we found to be sometimes imprecise and ill-behaved. To illustrate application of our SRI, we assessed niche separation among 3 ungulate species along a single environmental axis, solar radiation, on the northern Yellowstone winter range. We detected no difference between the niches occupied by bighorn sheep ( Ovis canadensis ) and elk ( Cervus elaphus; P = 0.104), but found that mule deer ( Odocoileus hemionus ) tended to use areas receiving more solar radiation than either of the other species ( P < 0.001). Overall, our SRI provides a useful metric that can reduce noise, improve interpretability, and increase parsimony in wildlife habitat models containing a solar radiation component.

Wyoming↗

Protocols for geologic hazards response by the Yellowstone Volcano Observatory to activity within the Yellowstone Volcanic System

Executive Summary The Yellowstone Plateau hosts an active volcanic system, with subterranean magma (molten rock), boiling, pressurized waters, and a variety of active faults with significant earthquake hazards. Within the next few decades, light-to-moderate earthquakes and steam explosions are certain to occur. Volcanic eruptions are less likely, but are ultimately inevitable in this active volcanic region. This document summarizes protocols, policies, and tools to be used by the Yellowstone Volcano Observatory (YVO) during earthquakes, hydrothermal explosions, or any geologic activity that could lead to a volcanic eruption. Yellowstone National Park is home to Yellowstone Caldera, the largest volcanic system by volume in the United States, as well as a vigorous hydrothermal system composed of pressurized subsurface boiling waters and active faults capable of generating substantial seismicity. The region is subject to hazards spanning a wide range of intensities, magnitudes, likelihood of occurrence, and geographic extent of impact. These hazards include small and comparatively common hydrothermal explosions, occasional strong earthquakes, rare relatively non-explosive lava flows, and very rare large explosive volcanic eruptions. Addressing the broad style of potential hazards and the vast spatial and temporal scales over which these hazards can occur requires a general plan that outlines the Yellowstone Volcano Observatory (YVO) response to a hazardous or potentially hazardous geological event or unrest (defined as departure from normal activity levels). The U.S. Geological Survey (USGS) Volcano Science Center (VSC) Response Plan for Significant Volcanic Events in the United States (Moran and others, 2024) forms the basis of any response by YVO but will be modified to suit the specific characteristics of the observatory, which operates as a consortium of nine federal, state, and academic institutions. Decisions on declaring an event response or “activity with potential” (defined as unrest that is not immediately hazardous but that may evolve into a hazardous event), as well as any changes in Volcano Alert Level and Aviation Color Code or the release of formal Information Statements, will be made by the USGS via the YVO Scientist-in-Charge (SIC) in consultation with the leads of the YVO member agencies. The YVO response to hazardous or potentially hazardous geological activity in or around Yellowstone National Park will focus on the collection and analysis of data relevant to the location and style of the activity. Those data will be interpreted within the existing geological framework for the region to develop probabilistic assessments of potential outcomes. These interpretations and assessments will be used to support decision making by emergency management officials including Yellowstone National Park managers or within the National Incident Management System if an Incident Command System (ICS) is activated. YVO will also convene a communications group open to each member agency to ensure consistent internal and external messaging and that the public is kept informed of the unrest through formal notifications, social media posts, online content, traditional media interviews, and community meetings. This response plan will be evaluated and updated as needed by the observatory and will be available through the YVO and USGS public websites. Responses to volcanic eruptions and responses outside of the Yellowstone region, but within the YVO area of responsibility (including Arizona, Utah, New Mexico, and Colorado), will follow the U.S. Geological Survey Volcano Science Center Response Plan for Significant Volcanic Events in the United States (Moran and others, 2024).

Idaho, Montana, Wyoming↗

Source and fate of inorganic solutes in the Gibbon River, Yellowstone National Park, Wyoming, USA: I. Low-flow discharge and major solute chemistry

The Gibbon River in Yellowstone National Park (YNP) is an important natural resource and habitat for fisheries and wildlife. However, the Gibbon River differs from most other mountain rivers because its chemistry is affected by several geothermal sources including Norris Geyser Basin, Chocolate Pots, Gibbon Geyser Basin, Beryl Spring, and Terrace Spring. Norris Geyser Basin is one of the most dynamic geothermal areas in YNP, and the water discharging from Norris is much more acidic (pH 3) than other geothermal basins in the upper-Madison drainage (Gibbon and Firehole Rivers). Water samples and discharge data were obtained from the Gibbon River and its major tributaries near Norris Geyser Basin under the low-flow conditions of September 2006. Surface inflows from Norris Geyser Basin were sampled to identify point sources and to quantify solute loading to the Gibbon River. The source and fate of the major solutes (Ca, Mg, Na, K, SiO 2 , Cl, F, HCO 3 , SO 4 , NO 3 , and NH 4 ) in the Gibbon River were determined in this study and these results may provide an important link in understanding the health of the ecosystem and the behavior of many trace solutes. Norris Geyser Basin is the primary source of Na, K, Cl, SO 4 , and N loads (35–58%) in the Gibbon River. The largest source of HCO 3 and F is in the lower Gibbon River reach. Most of the Ca and Mg originate in the Gibbon River upstream from Norris Geyser Basin. All the major solutes behave conservatively except for NH 4 , which decreased substantially downstream from Gibbon Geyser Basin, and SiO 2 , small amounts of which precipitated on mixing of thermal drainage with the river. As much as 9–14% of the river discharge at the gage is from thermal flows during this period.

Wyoming↗

Water-chemistry data for selected springs, geysers, and streams in Yellowstone National Park, Wyoming, 2006-2008

Water analyses are reported for 104 samples collected from numerous thermal and non-thermal features in Yellowstone National Park (YNP) during 2006-2008. Water samples were collected and analyzed for major and trace constituents from 10 areas of YNP including Apollinaris Spring and Nymphy Creek along the Norris-Mammoth corridor, Beryl Spring in Gibbon Canyon, Norris Geyser Basin, Lower Geyser Basin, Crater Hills, the Geyser Springs Group, Nez Perce Creek, Rabbit Creek, the Mud Volcano area, and Washburn Hot Springs. These water samples were collected and analyzed as part of research investigations in YNP on arsenic, antimony, iron, nitrogen, and sulfur redox species in hot springs and overflow drainages, and the occurrence and distribution of dissolved mercury. Most samples were analyzed for major cations and anions, trace metals, redox species of antimony, arsenic, iron, nitrogen, and sulfur, and isotopes of hydrogen and oxygen. Analyses were performed at the sampling site, in an on-site mobile laboratory vehicle, or later in a U.S. Geological Survey laboratory, depending on stability of the constituent and whether it could be preserved effectively. Water samples were filtered and preserved on-site. Water temperature, specific conductance, pH, emf (electromotive force or electrical potential), and dissolved hydrogen sulfide were measured on-site at the time of sampling. Dissolved hydrogen sulfide was measured a few to several hours after sample collection by ion-specific electrode on samples preserved on-site. Acidity was determined by titration, usually within a few days of sample collection. Alkalinity was determined by titration within 1 to 2 weeks of sample collection. Concentrations of thiosulfate and polythionate were determined as soon as possible (generally a few to several hours after sample collection) by ion chromatography in an on-site mobile laboratory vehicle. Total dissolved iron and ferrous iron concentrations often were measured on-site in the mobile laboratory vehicle. Concentrations of dissolved aluminum, arsenic, boron, barium, beryllium, calcium, cadmium, cobalt, chromium, copper, iron, potassium, lithium, magnesium, manganese, molybdenum, sodium, nickel, lead, selenium, silica, strontium, vanadium, and zinc were determined by inductively coupled plasma-optical emission spectrometry. Trace concentrations of dissolved antimony, cadmium, cobalt, chromium, copper, lead, and selenium were determined by Zeeman-corrected graphite-furnace atomic-absorption spectrometry. Dissolved concentrations of total arsenic, arsenite, total antimony, and antimonite were determined by hydride generation atomic-absorption spectrometry using a flow-injection analysis system. Dissolved concentrations of total mercury and methylmercury were determined by cold-vapor atomic fluorescence spectrometry. Concentrations of dissolved chloride, fluoride, nitrate, bromide, and sulfate were determined by ion chromatography. For many samples, concentrations of dissolved fluoride also were determined by ion-specific electrode. Concentrations of dissolved ferrous and total iron were determined by the FerroZine colorimetric method. Concentrations of dissolved ammonium were determined by ion chromatography, with reanalysis by colorimetry when separation of sodium and ammonia peaks was poor. Dissolved organic carbon concentrations were determined by the wet persulfate oxidation method. Hydrogen and oxygen isotope ratios were determined using the hydrogen and CO 2 equilibration techniques, respectively.

Wyoming↗

Is the track of the Yellowstone hotspot driven by a deep mantle plume? - Review of volcanism, faulting, and uplift in light of new data

Geophysical imaging of a tilted mantle plume extending at least 500 km beneath the Yellowstone caldera provides compelling support for a plume origin of the entire Yellowstone hotspot track back to its inception at 17 Ma with eruptions of flood basalts and rhyolite. The widespread volcanism, combined with a large volume of buoyant asthenosphere, supports a plume head as an initial phase. Estimates of the diameter of the plume head suggest it completely spanned the upper mantle and was fed from sources beneath the transition zone, We consider a mantle–plume depth to at least 1,000 km to best explain the large scale of features associated with the hotspot track. The Columbia River–Steens flood basalts form a northward-migrating succession consistent with the outward spreading of a plume head beneath the lithosphere. The northern part of the inferred plume head spread (pancaked) upward beneath Mesozoic oceanic crust to produce flood basalts, whereas basalt melt from the southern part intercepted and melted Paleozoic and older crust to produce rhyolite from 17 to 14 Ma. The plume head overlapped the craton margin as defined by strontium isotopes; westward motion of the North American plate has likely “scraped off” the head from the plume tail. Flood basalt chemistries are explained by delamination of the lithosphere where the plume head intersected this cratonic margin. Before reaching the lithosphere, the rising plume head apparently intercepted the east-dipping Juan de Fuca slab and was deflected ~ 250 km to the west; the plume head eventually broke through the slab, leaving an abruptly truncated slab. Westward deflection of the plume head can explain the anomalously rapid hotspot movement of 62 km/m.y. from 17 to 10 Ma, compared to the rate of ~ 25 km/m.y. from 10 to 2 Ma. A plume head-to-tail transition occurred in the 14-to-10-Ma interval in the central Snake River Plain and was characterized by frequent (every 200–300 ka for about 2 m.y. from 12.7 to 10.5 Ma) “large volume (> 7000 km 3 )”, and high temperature rhyolitic eruptions (> 1000 °C) along a ~ 200–km-wide east–west band. The broad transition area required a heat source of comparable area. Differing characteristics of the volcanic fields here may in part be due to variations in crustal composition but also may reflect development in differing parts of an evolving plume where the older fields may reflect the eruption from several volcanic centers located above very large and extensive rhyolitic magma chamber(s) over the detached plume head while the younger fields may signal the arrival of the plume tail intercepting and melting the lithosphere and generating a more focused rhyolitic magma chamber. The three youngest volcanic fields of the hotspot track started with large ignimbrite eruptions at 10.21, 6.62, and 2.05 Ma. They indicate hotspot migration N55° E at ~ 25 km/m.y. compatible in direction and velocity with the North American Plate motion. The Yellowstone Crescent of High Terrain (YCHT) flares outward ahead of the volcanic progression in a pattern similar to a bow-wave, and thus favors a sub-lithospheric driver. Estimates of YCHT-uplift rates are between 0.1 and 0.4 mm/yr. Drainage divides have migrated northeastward with the hotspot. The Continental Divide and a radial drainage pattern now centers on the hotspot. The largest geoid anomaly in the conterminous U.S. is also centered on Yellowstone and, consistent with uplift above a mantle plume. Bands of late Cenozoic faulting extend south and west from Yellowstone. These bands are subdivided into belts based both on recency of offset and range-front height. Fault history within these belts suggests the following pattern: Belt I — starting activity but little accumulated offset; Belt II — peak activity with high total offset and activity younger than 14 ka; Belt III — waning activity with large offset and activity younger than 140 ka; and Belt IV — apparently dead on substantial range fronts (south side of the eastern Snake River Plain only). These belts of fault activity have migrated northeast in tandem with the adjacent hotspot volcanism. On the southern arm of the YCHT, fault activity occurs on the inner, western slope consistent with driving by gravitational potential energy, whereas faulting has not started on the eastern, outer, more compressional slope. Range fronts increase in height and steepness northeastward along the southern-fault band. Both the belts of faulting and the YCHT are asymmetrical across the volcanic hotspot track, flaring out 1.6 times more on the south than the north side. This and the southeast tilt of the Yellowstone plume may reflect southeast flow of the upper mantle.

Wyoming↗

Methylmercury enters an aquatic food web through acidophilic microbial mats in Yellowstone National Park, Wyoming

Microbial mats are a visible and abundant life form inhabiting the extreme environments in Yellowstone National Park (YNP), WY, USA. Little is known of their role in food webs that exist in the Park's geothermal habitats. Eukaryotic green algae associated with a phototrophic green/purple Zygogonium microbial mat community that inhabits low-temperature regions of acidic (pH ∼ 3.0) thermal springs were found to serve as a food source for stratiomyid (Diptera: Stratiomyidae) larvae. Mercury in spring source water was taken up and concentrated by the mat biomass. Monomethylmercury compounds (MeHg + ), while undetectable or near the detection limit (0.025 ng l −1 ) in the source water of the springs, was present at concentrations of 4–7 ng g −1 dry weight of mat biomass. Detection of MeHg + in tracheal tissue of larvae grazing the mat suggests that MeHg + enters this geothermal food web through the phototrophic microbial mat community. The concentration of MeHg + was two to five times higher in larval tissue than mat biomass indicating MeHg + biomagnification occurred between primary producer and primary consumer trophic levels. The Zygogonium mat community and stratiomyid larvae may also play a role in the transfer of MeHg + to species in the food web whose range extends beyond a particular geothermal feature of YNP.

Wyoming↗

Hydrothermal processes above the Yellowstone magma chamber: Large hydrothermal systems and large hydrothermal explosions

Hydrothermal explosions are violent and dramatic events resulting in the rapid ejection of boiling water, steam, mud, and rock fragments from source craters that range from a few meters up to more than 2 km in diameter; associated breccia can be emplaced as much as 3 to 4 km from the largest craters. Hydrothermal explosions occur where shallow interconnected reservoirs of steam- and liquid-saturated fluids with temperatures at or near the boiling curve underlie thermal fields. Sudden reduction in confining pressure causes fluids to flash to steam, resulting in significant expansion, rock fragmentation, and debris ejection. In Yellowstone, hydrothermal explosions are a potentially significant hazard for visitors and facilities and can damage or even destroy thermal features. The breccia deposits and associated craters formed from hydrothermal explosions are mapped as mostly Holocene (the Mary Bay deposit is older) units throughout Yellowstone National Park (YNP) and are spatially related to within the 0.64-Ma Yellowstone caldera and along the active Norris-Mammoth tectonic corridor. In Yellowstone, at least 20 large (>100 m in diameter) hydrothermal explosion craters have been identified; the scale of the individual associated events dwarfs similar features in geothermal areas elsewhere in the world. Large hydrothermal explosions in Yellowstone have occurred over the past 16 ka averaging ~1 every 700 yr; similar events are likely in the future. Our studies of large hydrothermal explosion events indicate: (1) none are directly associated with eruptive volcanic or shallow intrusive events; (2) several historical explosions have been triggered by seismic events; (3) lithic clasts and comingled matrix material that form hydrothermal explosion deposits are extensively altered, indicating that explosions occur in areas subjected to intense hydrothermal processes; (4) many lithic clasts contained in explosion breccia deposits preserve evidence of repeated fracturing and vein-filling; and (5) areal dimensions of many large hydrothermal explosion craters in Yellowstone are similar to those of its active geyser basins and thermal areas. For Yellowstone, our knowledge of hydrothermal craters and ejecta is generally limited to after the Yellowstone Plateau emerged from beneath a late Pleistocene icecap that was roughly a kilometer thick. Large hydrothermal explosions may have occurred earlier as indicated by multiple episodes of cementation and brecciation commonly observed in hydrothermal ejecta clasts. Critical components for large, explosive hydrothermal systems include a water-saturated system at or near boiling temperatures and an interconnected system of well-developed joints and fractures along which hydrothermal fluids flow. Active deformation of the Yellowstone caldera, active faulting and moderate local seismicity, high heat flow, rapid changes in climate, and regional stresses are factors that have strong influences on the type of hydrothermal system developed. Ascending hydrothermal fluids flow along fractures that have developed in response to active caldera deformation and along edges of low-permeability rhyolitic lava flows. Alteration of the area affected, self-sealing leading to development of a caprock for the hydrothermal system, and dissolution of silica-rich rocks are additional factors that may constrain the distribution and development of hydrothermal fields. A partial low-permeability layer that acts as a cap to the hydrothermal system may produce some over-pressurization, thought to be small in most systems. Any abrupt drop in pressure initiates steam flashing and is rapidly transmitted through interconnected fractures that result in a series of multiple large-scale explosions contributing to the excavation of a larger explosion crater. Similarities between the size and dimensions of large hydrothermal explosion craters and thermal fields in Yellowstone may indicate that catastrophic events which result in large hydrothermal explosions are an end phase in geyser basin evolution. The Mary Bay hydrothermal explosion crater complex is the largest such complex in Yellowstone, and possibly in the world, with a diameter of 2.8 km in length and 2.4 km in width. It is nested in Mary Bay in the northern basin of Yellowstone Lake, an area of high heat flow and active deformation within the Yellowstone caldera. A sedimentary sequence exposed in wave-cut cliffs between Storm Point and Mary Bay gives insight into the geologic history of the Mary Bay hydrothermal explosion event. The Mary Bay explosion breccia deposits overlie sand above varved lake sediments and are separated locally into an upper and lower unit. The sand unit contains numerous small normal faults and is coextensive with the Mary Bay breccia in its northern extent. This sand may represent deposits of an earthquake-generated wave. Seismicity associated with the earthquake may have triggered the hydrothermal explosion responsible for development of the Mary Bay crater complex. Large hydrothermal explosions are rare events on a human time scale; however, the potential for additional future events of the sort in Yellowstone National Park is not insignificant. Based on the occurrence of large hydrothermal explosion events over the past 16,000 yr, an explosion large enough to create a 100-m-wide crater might be expected every 200 yr.

Wyoming↗

Elk calf survival and mortality following wolf restoration to Yellowstone National Park

We conducted a 3‐year study (May 2003–Apr 2006) of mortality of northern Yellowstone elk ( Cervus elaphus ) calves to determine the cause for the recruitment decline (i.e., 33 calves to 13 calves/100 adult F) following the restoration of wolves ( Canis lupus ). We captured, fit with radiotransmitters, and evaluated blood characteristics and disease antibody seroprevalence in 151 calves ≤6 days old (68M:83F). Concentrations (x̄, SE) of potential condition indicators were as follows: thyroxine (T4; 13.8 μg/dL, 0.43), serum urea nitrogen (SUN; 17.4 mg/dL, 0.57), γ‐glutamyltransferase (GGT; 66.4 IU/L, 4.36), gamma globulins (GG; 1.5 g/dL, 0.07), and insulin‐like growth factor‐1 (IGF‐1; 253.6 ng/mL, 9.59). Seroprevalences were as follows: brucellosis ( Brucella abortus ; 3%), bovine‐respiratory syncytial virus (3%), bovine‐viral‐diarrhea virus type 1 (25%), infectious‐bovine rhinotracheitis (58%), and bovine parainfluenza‐3 (32%). Serum urea nitrogen, GGT, GG, and IGF‐1 varied with year; T4, SUN, and GG varied with age ( P ≤ 0.01); and SUN varied by capture area ( P = 0.02). Annual survival was 0.22 (SE = 0.035, n = 149) and varied by calving area but not year. Neonates captured in the Stephens Creek/Mammoth area of Yellowstone National Park, USA, had annual survival rates >3× higher (0.54) than those captured in the Lamar Valley area (0.17), likely due to the higher predator density in Lamar Valley. Summer survival (20 weeks after radiotagging) was 0.29 (SE = 0.05, n = 116), and calving area, absolute deviation from median birth date, and GG were important predictors of summer survival. Survival during winter (Nov‐Apr) was 0.90 (SE= 0.05, n = 42), and it did not vary by calving area or year. Sixty‐nine percent ( n = 104) of calves died within the first year of life, 24% ( n = 36) survived their first year, and 7% ( n = 11) had unknown fates. Grizzly bears ( Ursus arctos ) and black bears ( Ursus americanus ) accounted for 58–60% ( n = 60–62) of deaths, and wolves accounted for 14–17% ( n = 15–18). Summer predation (95% of summer deaths) increased, and winter malnutrition (0% of winter deaths) decreased, compared with a similar study during 1987–1990 (72% and 58%, respectively). Physiological factors (e.g., low levels of GG) may predispose calves to predation. Also, the increase in bear numbers since wolf restoration and spatial components finer than the northern range should be considered when trying to determine the causes of the northern Yellowstone elk decline. This is the first study to document the predation impacts from reintroduced wolves on elk calf mortality in an ecosystem already containing established populations of 4 other major predators (i.e., grizzly and black bears, cougars [ Puma concolor ], and coyotes [ Canis latrans ]). The results are relevant to resource managers of the Yellowstone ecosystem in understanding the dynamics of the elk population, in providing harvest quota recommendations for local elk hunts to the Montana Department of Fish, Wildlife and Parks, the United States Fish and Wildlife Service regarding wolf and grizzly bear recovery, and to all areas worldwide where predators are increasing, by providing managers with information about potential carnivore impacts on elk populations.

Idaho, Montana, Wyoming↗

Water-chemistry data for selected springs, geysers, and streams in Yellowstone National Park, Wyoming, 2003-2005

Water analyses are reported for 157 samples collected from numerous hot springs, their overflow drainages, and Lemonade Creek in Yellowstone National Park (YNP) during 2003–2005. Water samples were collected and analyzed for major and trace constituents from ten areas of YNP including Terrace and Beryl Springs in the Gibbon Canyon area, Norris Geyser Basin, the West Nymph Creek thermal area, the area near Nymph Lake, Hazle Lake, and Frying Pan Spring, Lower Geyser Basin, Washburn Hot Springs, Mammoth Hot Springs, Potts Hot Spring Basin, the Sulphur Caldron area, and Lemonade Creek near the Solfatara Trail. These water samples were collected and analyzed as part of research investigations in YNP on arsenic, antimony, and sulfur redox distribution in hot springs and overflow drainages, and the occurrence and distribution of dissolved mercury. Most samples were analyzed for major cations and anions, trace metals, redox species of antimony, arsenic, iron, nitrogen, and sulfur, and isotopes of hydrogen and oxygen. Analyses were performed at the sampling site, in an on-site mobile laboratory vehicle, or later in a U.S. Geological Survey laboratory, depending on stability of the constituent and whether it could be preserved effectively. Water samples were filtered and preserved onsite. Water temperature, specific conductance, pH, Eh (redox potential relative to the Standard Hydrogen Electrode), and dissolved hydrogen sulfide were measured onsite at the time of sampling. Acidity was determined by titration, usually within a few days of sample collection. Alkalinity was determined by titration within 1 to 2 weeks of sample collection. Concentrations of thiosulfate and polythionate were determined as soon as possible (generally minutes to hours after sample collection) by ion chromatography in an on-site mobile laboratory vehicle. Total dissolved-iron and ferrous-iron concentrations often were measured onsite in the mobile laboratory vehicle. Concentrations of dissolved aluminum, arsenic, boron, barium, beryllium, calcium, cadmium, cobalt, chromium, copper, iron, potassium, lithium, magnesium, manganese, molybdenum, sodium, nickel, lead, selenium, silica, strontium, vanadium, and zinc were determined by inductively-coupled plasma-optical emission spectrometry. Trace concentrations of dissolved antimony, cadmium, cobalt, chromium, copper, lead, and selenium were determined by Zeeman-corrected graphite-furnace atomic-absorption spectrometry. Dissolved concentrations of total arsenic, arsenite, total antimony, and antimonite were determined by hydride-generation atomic-absorption spectrometry using a flow-injection analysis system. Dissolved concentrations of total mercury and methyl mercury were determined by cold-vapor atomic-fluorescence spectrometry. Concentrations of dissolved chloride, fluoride, nitrate, bromide, and sulfate were determined by ion chromatography. Concentrations of dissolved ferrous and total iron were determined by the FerroZine colorimetric method. Concentrations of dissolved nitrite were determined by colorimetry or chemiluminescence. Concentrations of dissolved ammonium were determined by ion chromatography, with reanalysis by colorimetry when separation of sodium and ammonia peaks was poor. Dissolved organic carbon concentrations were determined by the wet persulfate oxidation method. Hydrogen and oxygen isotope ratios were determined using the hydrogen and CO 2 equilibration techniques, respectively.

Wyoming↗

Thioarsenates in geothermal waters of Yellowstone National Park: Determination, preservation, and geochemical importance

Mono-, di-, tri-, and tetrathioarsenate, as well as methylated arsenic oxy- and thioanions, were determined besides arsenite and arsenate in geothermal waters of Yellowstone National Park using anion-exchange chromatography inductively coupled plasma mass spectrometry. Retention time match with synthetic standards, measured S:As ratios, and molecular electrospray mass spectra support the identification. Acidification was unsuitable for arsenic species preservation in sulfidic waters, with HCl addition causing loss of total dissolved arsenic, presumably by precipitation of arsenic-sulfides. Flash-freezing is preferred for the preservation of arsenic species for several weeks. After thawing, samples must be analyzed immediately. Thioarsenates occurred over a pH range of 2.1 to 9.3 in the geothermal waters. They clearly predominated under alkaline conditions (up to 83% of total arsenic), but monothioarsenate also was detected in acidic waters (up to 34%). Kinetic studies along a drainage channel showed the importance of thioarsenates for the fate of arsenic discharged from the sulfidic hot spring. The observed arsenic speciation changes suggest three separate reactions: the transformation of trithioarsenate to arsenite (major initial reaction), the stepwise ligand exchange from tri- via di- and monothioarsenate to arsenate (minor reaction), and the oxidation of arsenite to arsenate, which only becomes quantitatively important after thioarsenates have disappeared.

Wyoming↗

Effects of supplemental feeding on gastrointestinal parasite infection in Rocky Mountain Elk (Cervus elaphus): Preliminary observations

The effects of management practices on the spread and impact of parasites and infectious diseases in wildlife and domestic animals are of increasing concern worldwide, particularly in cases where management of wild species can influence disease spill-over into domestic animals. In the Greater Yellowstone Ecosystem, USA, winter supplemental feeding of Rocky Mountain elk ( Cervus elaphus ) may enhance parasite and disease transmission by aggregating elk on feedgrounds. In this study, we tested the effect of supplemental feeding on gastrointestinal parasite infection in elk by comparing fecal egg/oocyst counts of fed and unfed elk. We collected fecal samples from fed and unfed elk at feedground and control sites from January to April 2006, and screened all samples for parasites. Six different parasite types were identified, and 48.7% of samples were infected with at least one parasite. Gastrointenstinal (GI) nematodes (Nematoda: Strongylida), Trichuris spp., and coccidia were the most common parasites observed. For all three of these parasites, fecal egg/oocyst counts increased from January to April. Supplementally fed elk had significantly higher GI nematode egg counts than unfed elk in January and February, but significantly lower counts in April. These patterns suggest that supplemental feeding may both increase exposure and decrease susceptibility of elk to GI nematodes, resulting in differences in temporal patterns of egg shedding between fed and unfed elk.

Wyoming↗

Accelerated uplift and magmatic intrusion of the Yellowstone caldera, 2004 to 2006

The Yellowstone caldera began a rapid episode of ground uplift in mid-2004, revealed by Global Positioning System and interferometric synthetic aperture radar measurements, at rates up to 7 centimeters per year, which is over three times faster than previously observed inflation rates. Source modeling of the deformation data suggests an expanding volcanic sill of ???1200 square kilometers at a 10-kilometer depth beneath the caldera, coincident with the top of a seismically imaged crustal magma chamber. The modeled rate of source volume increase is 0.1 cubic kilometer per year, similar to the amount of magma intrusion required to supply the observed high heat flow of the caldera. This evidence suggests magma recharge as the main mechanism for the accelerated uplift, although pressurization of magmatic fluids cannot be ruled out.

Wyoming↗

Evaluating estimators for numbers of females with cubs-of-the-year in the Yellowstone grizzly bear population

Current management of the grizzly bear ( Ursus arctos ) population in Yellowstone National Park and surrounding areas requires annual estimation of the number of adult female bears with cubs-of-the-year. We examined the performance of nine estimators of population size via simulation. Data were simulated using two methods for different combinations of population size, sample size, and coefficient of variation of individual sighting probabilities. We show that the coefficient of variation does not, by itself, adequately describe the effects of capture heterogeneity, because two different distributions of capture probabilities can have the same coefficient of variation. All estimators produced biased estimates of population size with bias decreasing as effort increased. Based on the simulation results we recommend the Chao estimator for model M h be used to estimate the number of female bears with cubs of the year; however, the estimator of Chao and Shen may also be useful depending on the goals of the research.

Wyoming↗

Population growth of Yellowstone grizzly bears: Uncertainty and future monitoring

Grizzly bears ( Ursus arctos ) in the Greater Yellowstone Ecosystem of the US Rocky Mountains have recently increased in numbers, but remain vulnerable due to isolation from other populations and predicted reductions in favored food resources. Harris et al. (2006) projected how this population might fare in the future under alternative survival rates, and in doing so estimated the rate of population growth, 1983&ndash;2002. We address issues that remain from that earlier work: (1) the degree of uncertainty surrounding our estimates of the rate of population change (&lambda;); (2) the effect of correlation among demographic parameters on these estimates; and (3) how a future monitoring system using counts of females accompanied by cubs might usefully differentiate between short-term, expected, and inconsequential fluctuations versus a true change in system state. We used Monte Carlo re-sampling of beta distributions derived from the demographic parameters used by Harris et al. (2006) to derive distributions of &lambda; during 1983&ndash;2002 given our sampling uncertainty. Approximate 95% confidence intervals were 0.972&ndash;1.096 (assuming females with unresolved fates died) and 1.008&ndash;1.115 (with unresolved females censored at last contact). We used well-supported models of Haroldson et al. (2006) and Schwartz et al. (2006a,b,c) to assess the strength of correlations among demographic processes and the effect of omitting them in projection models. Incorporating correlations among demographic parameters yielded point estimates of &lambda; that were nearly identical to those from the earlier model that omitted correlations, but yielded wider confidence intervals surrounding &lambda;. Finally, we suggest that fitting linear and quadratic curves to the trend suggested by the estimated number of females with cubs in the ecosystem, and using AIC c model weights to infer population sizes and &lambda; provides an objective means to monitoring approximate population trajectories in addition to demographic analysis.

Wyoming, Montana, Idaho↗

Amphibians and disease: Implications for conservation in the Greater Yellowstone Ecosystem

The decline of amphibian populations is a world-wide phenomenon that has received increasing attention since about 1990. In 2004, the World Conservation Union&rsquo;s global amphibian assessment concluded that 48% of the world&rsquo;s 5,743 described amphibian species were in decline, with 32% considered threatened (Stuart et al. 2004). Amphibian declines are a significant issue in the western United States, where all native species of frogs in the genus Rana and many toads in the genus Bufo are at risk, particularly those that inhabit mountainous areas (Corn 2003a,b; Bradford 2005).

Wyoming↗

Carnivore re-colonisation: Reality, possibility and a non-equilibrium century for grizzly bears in the southern Yellowstone ecosystem

Most large native carnivores have experienced range contractions due to conflicts with humans, although neither rates of spatial collapse nor expansion have been well characterised. In North America, the grizzly bear (Ursus arctos) once ranged from Mexico northward to Alaska, however its range in the continental USA has been reduced by 95-98%. Under the U. S. Endangered Species Act, the Yellowstone grizzly bear population has re-colonised habitats outside Yellowstone National Park. We analysed historical and current records, including data on radio-collared bears, (1) to evaluate changes in grizzly bear distribution in the southern Greater Yellowstone Ecosystem (GYE) over a 100-year period, (2) to utilise historical rates of re-colonisation to project future expansion trends and (3) to evaluate the reality of future expansion based on human limitations and land use. Analysis of distribution in 20-year increments reflects range reduction from south to north (1900-1940) and expansion to the south (1940-2000). Expansion was exponential and the area occupied by grizzly bears doubled approximately every 20 years. A complementary analysis of bear occurrence in Grand Teton National Park also suggests an unprecedented period of rapid expansion during the last 20-30 years. The grizzly bear population currently has re-occupied about 50% of the southern GYE. Based on assumptions of continued protection and ecological stasis, our model suggests total occupancy in 25 years. Alternatively, extrapolation of linear expansion rates from the period prior to protection suggests total occupancy could take > 100 years. Analyses of historical trends can be useful as a restoration tool because they enable a framework and timeline to be constructed to pre-emptively address the social challenges affecting future carnivore recovery. ?? 2004 The Zoological Society of London.

Idaho, Montana, Wyoming↗