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Research about Wyoming

Source-linked reports with geographic coverage including Wyoming.

At least 325 records · Page 18Linked to original sources

Empirical estimation of recreational exploitation of burbot, Lota lota, in the Wind River drainage of Wyoming using a multistate capture–recapture model

Burbot, Lota lota (Linnaeus), is a regionally popular sportfish in the Wind River drainage of Wyoming, USA, at the southern boundary of the range of the species. Recent declines in burbot abundances were hypothesised to be caused by overexploitation, entrainment in irrigation canals and habitat loss. This study addressed the overexploitation hypothesis using tagging data to generate reliable exploitation, abundance and density estimates from a multistate capture–recapture model that accounted for incomplete angler reporting and tag loss. Exploitation rate μ was variable among the study lakes and inversely correlated with density. Exploitation thresholds μ 40 associated with population densities remaining above 40% of carrying capacity were generated to characterise risk of overharvest using exploitation and density estimates from tagging data and a logistic surplus-production model parameterised with data from other burbot populations. Bull Lake (μ = 0.06, 95% CI: 0.03–0.11; μ 40 = 0.18) and Torrey Lake (μ = 0.02, 95% CI: 0.00–0.11; μ 40 = 0.18) had a low risk of overfishing, Upper Dinwoody Lake had intermediate risk (μ = 0.08, 95% CI: 0.02–0.32; μ 40 = 0.18) and Lower Dinwoody Lake had high risk (μ = 0.32, 95% CI: 0.10–0.67; μ 40 = 0.08). These exploitation and density estimates can be used to guide sustainable management of the Wind River drainage recreational burbot fishery and inform management of other burbot fisheries elsewhere.

Wyoming↗

The greenscape shapes surfing of resource waves in a large migratory herbivore

The Green Wave Hypothesis posits that herbivore migration manifests in response to waves of spring green-up (i.e. green-wave surfing). Nonetheless, empirical support for the Green Wave Hypothesis is mixed, and a framework for understanding variation in surfing is lacking. In a population of migratory mule deer ( Odocoileus hemionus ), 31% surfed plant phenology in spring as well as a theoretically perfect surfer, and 98% surfed better than random. Green-wave surfing varied among individuals and was unrelated to age or energetic state. Instead, the greenscape, which we define as the order, rate and duration of green-up along migratory routes, was the primary factor influencing surfing. Our results indicate that migratory routes are more than a link between seasonal ranges, and they provide an important, but often overlooked, foraging habitat. In addition, the spatiotemporal configuration of forage resources that propagate along migratory routes shape animal movement and presumably, energy gains during migration.

Wyoming↗

Patterns in Greater Sage-grouse population dynamics correspond with public grazing records at broad scales

Human land use, such as livestock grazing, can have profound yet varied effects on wildlife interacting within common ecosystems, yet our understanding of land-use effects is often generalized from short-term, local studies that may not correspond with trends at broader scales. Here we used public land records to characterize livestock grazing across Wyoming, USA, and we used Greater Sage-grouse ( Centrocercus urophasianus ) as a model organism to evaluate responses to livestock management. With annual counts of male Sage-grouse from 743 leks (breeding display sites) during 2004–2014, we modeled population trends in response to grazing level (represented by a relative grazing index) and timing across a gradient in vegetation productivity as measured by the Normalized Vegetation Difference Index (NDVI). We found grazing can have both positive and negative effects on Sage-grouse populations depending on the timing and level of grazing. Sage-grouse populations responded positively to higher grazing levels after peak vegetation productivity, but populations declined when similar grazing levels occurred earlier, likely reflecting the sensitivity of cool-season grasses to grazing during peak growth periods. We also found support for the hypothesis that effects of grazing management vary with local vegetation productivity. These results illustrate the importance of broad-scale analyses by revealing patterns in Sage-grouse population trends that may not be inferred from studies at finer scales, and could inform sustainable grazing management in these ecosystems.

Wyoming↗

When perception reflects reality: Non-native grass invasion alters small mammal risk landscapes and survival

Modification of habitat structure due to invasive plants can alter the risk landscape for wildlife by, for example, changing the quality or availability of refuge habitat. Whether perceived risk corresponds with actual fitness outcomes, however, remains an important open question. We simultaneously measured how habitat changes due to a common invasive grass (cheatgrass, Bromus tectorum ) affected the perceived risk, habitat selection, and apparent survival of a small mammal, enabling us to assess how well perceived risk influenced important behaviors and reflected actual risk. We measured perceived risk by nocturnal rodents using a giving-up density foraging experiment with paired shrub (safe) and open (risky) foraging trays in cheatgrass and native habitats. We also evaluated microhabitat selection across a cheatgrass gradient as an additional assay of perceived risk and behavioral responses for deer mice ( Peromyscus maniculatus ) at two spatial scales of habitat availability. Finally, we used mark-recapture analysis to quantify deer mouse apparent survival across a cheatgrass gradient while accounting for detection probability and other habitat features. In the foraging experiment, shrubs were more important as protective cover in cheatgrass-dominated habitats, suggesting that cheatgrass increased perceived predation risk. Additionally, deer mice avoided cheatgrass and selected shrubs, and marginally avoided native grass, at two spatial scales. Deer mouse apparent survival varied with a cheatgrass–shrub interaction, corresponding with our foraging experiment results, and providing a rare example of a native plant mediating the effects of an invasive plant on wildlife. By synthesizing the results of three individual lines of evidence (foraging behavior, habitat selection, and apparent survival), we provide a rare example of linkage between behavioral responses of animals indicative of perceived predation risk and actual fitness outcomes. Moreover, our results suggest that exotic grass invasions can influence wildlife populations by altering risk landscapes and survival.

Wyoming↗

Stratigraphic cross sections of the Niobrara interval of the Cody Shale and associated rocks in the Wind River Basin, central Wyoming

The Wind River Basin in Wyoming is one of many structural and sedimentary basins that formed in the Rocky Mountain foreland during the Laramide orogeny. The basin is nearly 200 miles long, 70 miles wide, and encompasses about 7,400 square miles in central Wyoming. The basin is bounded by the Washakie Range, Owl Creek uplift, and southern Bighorn Mountains on the north, the Casper arch on the east, the Granite Mountains on the south, and Wind River Range on the west. Many important conventional oil and gas fields producing from reservoirs ranging in age from Mississippian through Tertiary have been discovered in this basin. In addition, an extensive unconventional overpressured basin-centered gas accumulation has been identified in Cretaceous and Tertiary strata in the deeper parts of the basin. It has long been suggested that various Upper Cretaceous marine shales, including the Cody Shale, are the principal hydrocarbon source rocks for many of these accumulations. With recent advances and success in horizontal drilling and multistage fracture stimulation, there has been an increase in exploration and completion of wells in these marine shales in other Rocky Mountain Laramide basins that were traditionally thought of only as hydrocarbon source rocks. The two stratigraphic cross sections presented in this report were constructed as part of a project carried out by the U.S. Geological Survey to characterize and evaluate the undiscovered continuous (unconventional) oil and gas resources of the Niobrara interval of the Upper Cretaceous Cody Shale in the Wind River Basin in central Wyoming. The primary purpose of the cross sections is to show the stratigraphic relationship of the Niobrara equivalent strata and associated rocks in the lower part of the Cody Shale in the Wind River Basin. These two cross sections were constructed using borehole geophysical logs from 37 wells drilled for oil and gas exploration and production, and one surface section along East Sheep Creek near Shotgun Butte in the northwestern part of the basin. Both lines originate at the East Sheep Creek surface section and end near Clarkson Hill in the extreme southeastern part of the basin. The stratigraphic interval extends from the upper part of the Frontier Formation to the middle part of the Cody Shale. The datum is the base of the “chalk kick” marker bed, a distinctive resistivity peak or zone in the lower part of the Cody Shale. A gamma ray and (or) spontaneous potential (SP) log was used in combination with a resistivity log to identify and correlate units. Marine molluscan index fossils collected from nearby outcrop sections were projected into the subsurface to help determine the relative ages of the strata and aid in correlation.

Wyoming↗

Climate changes and wildfire alter vegetation of Yellowstone National Park, but forest cover persists

We present landscape simulation results contrasting effects of changing climates on forest vegetation and fire regimes in Yellowstone National Park, USA, by mid-21st century. We simulated potential changes to fire dynamics and forest characteristics under three future climate projections representing a range of potential future conditions using the FireBGCv2 model. Under the future climate scenarios with moderate warming (>2°C) and moderate increases in precipitation (3–5%), model simulations resulted in 1.2–4.2 times more burned area, decreases in forest cover (10–44%), and reductions in basal area (14–60%). In these same scenarios, lodgepole pine ( Pinus contorta ) decreased in basal area (18–41%), while Douglas-fir ( Pseudotsuga menziesii ) basal area increased (21–58%). Conversely, mild warming (<2°C) coupled with greater increases in precipitation (12–13%) suggested an increase in forest cover and basal area by mid-century, with spruce and subalpine fir increasing in abundance. Overall, we found changes in forest tree species compositions were caused by the climate-mediated changes in fire regime (56–315% increase in annual area burned). Simulated changes in forest composition and fire regime under warming climates portray a landscape that shifts from lodgepole pine to Douglas-fir caused by the interaction between the magnitude and seasonality of future climate changes, by climate-induced changes in the frequency and intensity of wildfires, and by tree species response.

Wyoming↗

The Finley Site: Antiquity of the Finley Site

This report is based on two months reconnaissance in the summer of 1941 in the Eden Valley, Wyoming. The work is as yet far from complete and the conclusions presented here must be regarded as tentative. It is hoped that in the future more extensive geological work may be undertaken. The Finley site provides a promising opportunity for a determination of the age of the Yuma culture. The points and bones described by Dr. Howard occur in a culture layer which has been found in a large dune area, here called the Killpecker dunes. The deposition of the culture layer is one of a series of events in the dune area which can be correlated with the cutting and filling of gravel terraces in the Eden Valley to the west, and these in turn may be related to the glacial chronology in the Wind River Mountains which lie to the north.

Wyoming↗

Geology, palynology, and climatic significance of two pre-Pinedale Lake sediment sequences in and near Yellowstone National Park

Pollen analysis of a section of lake sediments at Grassy Lake Reservoir indicates a vegetational sequence changing from tundra, to spruce-fir-pine forest, to pine forest, to tundra at the top. Pollen analysis of a section of lake sediments on Beaverdam Creek indicates a tundra vegetation at the base, followed by a brief episode of spruce-fir forest and a return to a tundra vegetation at the top. The analyses of both sections suggest a cold to cool to cold climatic sequence, interpreted as interstadial in character. However, differences suggest that they represent separate interstadials. Pinedale Till disconformably overlies the lake deposits at Grassy Lake Reservoir. The upper sediments contain wood 14 C dated at >42,000 yr; the lowermost interfinger with till shown to be more than about 70,000 yr old. The deposits at Beaverdam Creek grade upward into proglacial Pinedale deposits, contain an ash that is probably about 70,000 yr old near their base, and rest comformably on gravel that grades down into lake sediments containing wood debris suggestive of an older climatic amelioration. We conclude that the warmest part of the interstadial at Grassy Lake Reservoir is probably more than 70,000 yr old, and that the warmest part of the interstadial analyzed at Beaverdam Creek is slightly younger than 70,000 yr old.

Wyoming↗

A loess–paleosol record of climate and glacial history over the past two glacial–interglacial cycles (~ 150 ka), southern Jackson Hole, Wyoming

Loess accumulated on a Bull Lake outwash terrace of Marine Oxygen Isotope Stage 6 (MIS 6) age in southern Jackson Hole, Wyoming. The 9 m section displays eight intervals of loess deposition (Loess 1 to Loess 8, oldest), each followed by soil development. Our age-depth model is constrained by thermoluminescence, meteoric 10 Be accumulation in soils, and cosmogenic 10 Be surface exposure ages. We use particle size, geochemical, mineral-magnetic, and clay mineralogical data to interpret loess sources and pedogenesis. Deposition of MIS 6 loess was followed by a tripartite soil/thin loess complex (Soils 8, 7, and 6) apparently reflecting the large climatic oscillations of MIS 5. Soil 8 (MIS 5e) shows the strongest development. Loess 5 accumulated during a glacial interval (~ 76–69 ka; MIS 4) followed by soil development under conditions wetter and probably colder than present. Deposition of thick Loess 3 (~ 43–51 ka, MIS 3) was followed by soil development comparable with that observed in Soil 1. Loess 1 (MIS 2) accumulated during the Pinedale glaciation and was followed by development of Soil 1 under a semiarid climate. This record of alternating loess deposition and soil development is compatible with the history of Yellowstone vegetation and the glacial flour record from the Sierra Nevada.

Wyoming↗

Using dynamic population simulations to extend resource selection analyses and prioritize habitats for conservation

Prioritizing habitats for conservation is a challenging task, particularly for species with fluctuating populations and seasonally dynamic habitat needs. Although the use of resource selection models to identify and prioritize habitat for conservation is increasingly common, their ability to characterize important long-term habitats for dynamic populations are variable. To examine how habitats might be prioritized differently if resource selection was directly and dynamically linked with population fluctuations and movement limitations among seasonal habitats, we constructed a spatially explicit individual-based model for a dramatically fluctuating population requiring temporally varying resources. Using greater sage-grouse ( Centrocercus urophasianus) in Wyoming as a case study, we used resource selection function maps to guide seasonal movement and habitat selection, but emergent population dynamics and simulated movement limitations modified long-term habitat occupancy. We compared priority habitats in RSF maps to long-term simulated habitat use. We examined the circumstances under which the explicit consideration of movement limitations, in combination with population fluctuations and trends, are likely to alter predictions of important habitats. In doing so, we assessed the future occupancy of protected areas under alternative population and habitat conditions. Habitat prioritizations based on resource selection models alone predicted high use in isolated parcels of habitat and in areas with low connectivity among seasonal habitats. In contrast, results based on more biologically-informed simulations emphasized central and connected areas near high-density populations, sometimes predicted to be low selection value. Dynamic models of habitat use can provide additional biological realism that can extend, and in some cases, contradict habitat use predictions generated from short-term or static resource selection analyses. The explicit inclusion of population dynamics and movement propensities via spatial simulation modeling frameworks may provide an informative means of predicting long-term habitat use, particularly for fluctuating populations with complex seasonal habitat needs. Importantly, our results indicate the possible need to consider habitat selection models as a starting point rather than the common end point for refining and prioritizing habitats for protection for cyclic and highly variable populations.

Wyoming↗

Monitoring gas and heat emissions at Norris Geyser Basin, Yellowstone National Park, USA based on a combined eddy covariance and Multi-GAS approach

We quantified gas and heat emissions in an acid-sulfate, vapor-dominated area (0.04-km 2 ) of Norris Geyser Basin, located just north of the 0.63 Ma Yellowstone Caldera and near an area of anomalous uplift. From 14 May to 3 October 2016, an eddy covariance system measured half-hourly CO 2 , H 2 O and sensible ( H ) and latent ( LE ) heat fluxes and a Multi-GAS instrument measured (1 Hz frequency) atmospheric H 2 O, CO 2 and H 2 S volumetric mixing ratios. We also measured soil CO 2 fluxes using the accumulation chamber method and temperature profiles on a grid and collected fumarole gas samples for geochemical analysis. Eddy covariance CO 2 fluxes ranged from − 56 to 885 g m − 2 d − 1 . Using wavelet analysis, average daily eddy covariance CO 2 fluxes were locally correlated with average daily environmental parameters on several-day to monthly time scales. Estimates of CO 2 emission rate from the study area ranged from 8.6 t d − 1 based on eddy covariance measurements to 9.8 t d − 1 based on accumulation chamber measurements. Eddy covariance water vapor fluxes ranged from 1178 to 24,600 g m − 2 d − 1 . Nighttime H and LE were considered representative of hydrothermal heat fluxes and ranged from 4 to 183 and 38 to 504 W m − 2 , respectively. The total hydrothermal heat emission rate ( H + LE + radiant) estimated for the study area was 11.6 MW and LE contributed 69% of the output. The mean ± standard deviation of H 2 O, CO 2 and H 2 S mixing ratios measured by the Multi-GAS system were 9.3 ± 3.1 parts per thousand, 467 ± 61 ppmv, and 0.5 ± 0.6 ppmv, respectively, and variations in the gas compositions were strongly correlated with diurnal variations in environmental parameters (wind speed and direction, atmospheric temperature). After removing ambient H 2 O and CO 2 , the observed variations in the Multi-GAS data could be explained by the mixing of relatively H 2 O-CO 2 -H 2 S-rich fumarole gases with CO 2 -rich and H 2 O-H 2 S-poor soil gases. The fumarole H 2 O/CO 2 and CO 2 /H 2 S end member ratios (101.7 and 27.1, respectively, on average) were invariant during the measurement period and fell within the range of values measured in direct fumarole gas samples. The soil gas H 2 O/CO 2 end member ratios (~ 15–30) were variable and low relative to the fumarole end member, likely resulting from water vapor loss during cooling and condensation in the shallow subsurface, whereas the CO 2 /H 2 S end member ratio was high (~ 160), presumably related to transport of CO 2 -dominated soil gas emissions mixed with trace fumarolic emissions to the Multi-GAS station. Nighttime eddy covariance ratios of H 2 O to CO 2 flux were typically between the soil gas and fumarole end member H 2 O/CO 2 ratios defined by Multi-GAS measurements. Overall, the combined eddy covariance and Multi-GAS approach provides a powerful tool for quasi-continuous measurements of gas and heat emissions for improved volcano-hydrothermal monitoring.

Wyoming↗

Comparison of burbot populations across adjacent native and introduced ranges

Introduced species are a threat to biodiversity. Burbot, Lota lota, a fish native to the Wind River Drainage, Wyoming and a species of conservation concern, have been introduced into the nearby Green River Drainage, Wyoming, where they are having negative effects on native fish species. We compared these native and introduced burbot populations to evaluate potential mechanisms that could be leading to introduction success. We examined genetic ancestry, physical habitat characteristics, community composition, and burbot abundance, relative weight, and size structure between the native and introduced range to elucidate potential differences. The origin of introduced burbot in Flaming Gorge Reservoir is most likely Boysen Reservoir and several nearby river populations in the native Wind River Drainage. Burbot populations did not show consistent differences in abundance, size structure, and relative weight between drainages, though Fontenelle Reservoir, in the introduced drainage, had the largest burbot. There were also limited environmental and community composition differences, though reservoirs in the introduced drainage had lower species richness and a higher percentage of non-native fish species than the reservoir in the native drainage. Burbot introduction in the Green River Drainage is likely an example of reservoir construction creating habitat with suitable environmental conditions to allow a southwards range expansion of this cold-water species. An understanding of the factors driving introduction success can allow better management of species, both in their introduced and native range.

Wyoming↗

The thermal regime and species composition of fish and invertebrates in Kelly Warm Spring, Grand Teton National Park, Wyoming

We evaluated the thermal regime and relative abundance of native and nonnative fish and invertebrates within Kelly Warm Spring and Savage Ditch, Grand Teton National Park, Wyoming. Water temperatures within the system remained relatively warm year-round with mean temperatures >20 °C near the spring source and >5 °C approximately 2 km downstream of the source. A total of 7 nonnative species were collected: Convict/Zebra Cichlid ( Cichlasoma nigrofasciatum ), Green Swordtail ( Xiphophorus hellerii ), Tadpole Madtom ( Noturus gyrinus ), Guppy ( Poecilia reticulata ), Goldfish ( Carassius auratus ), red-rimmed melania snail ( Melanoides tuberculata ), and American bullfrog tadpoles ( Lithobates catesbeianus ). Nonnative fish (Zebra Cichlids and Green Swordtails), red-rimmed melania snails, and bullfrog tadpoles dominated the upper 2 km of the system. Abundance estimates of the Zebra Cichlid exceeded 12,000 fish/km immediately downstream of the spring source. Relative abundance of native species increased movingdownstream as water temperatures attenuated with distance from the thermally warmed spring source; however, nonnative species were captured 4 km downstream from the spring. Fish diseases were prevalent in both native and nonnative fish from the Kelly Warm Spring pond. Clinostomum marginatum, a trematode parasite, was found in native species samples, and the tapeworm Diphyllobothrium dendriticum was present in samples from nonnative species. Diphyllobothrium dendriticum is rare in Wyoming. Salmonella spp. were also found in some samples of nonnative species. These bacteria are associated with aquarium fish and aquaculture and are generally not found in the wild.

Wyoming↗

Introduced American Bullfrog distribution and diets in Grand Teton National Park

Introduced American Bullfrogs (Lithobates catesbeianus) have been present in Grand Teton National Park since approximately the 1950s, but little is known about their distribution and potential impacts. In this study, we surveyed the current bullfrog distribution and spatial overlap with sympatric native amphibians in the park, and characterized post-metamorphic bullfrog diets from July – September 2015. Despite surveys in multiple large rivers and floodplain habitats, we only documented bullfrogs in a geothermal pond and 5 km of stream channel immediately downstream of this pond. In these waters, bullfrogs overlapped with native amphibians at the downstream end of their distribution, and we did not document native amphibians in bullfrog stomach contents. Larger bullfrogs (SVL ≥ 96 mm) primarily consumed native rodents (especially meadow voles, Microtus pennsylvanicus ), while smaller bullfrogs frequently consumed native invertebrates and less frequently consumed non-native invertebrates and fish. Taken together, these data indicate that the distribution and implications of the bullfrog invasion in Grand Teton National Park are currently localized to a small area, so these bullfrogs should therefore be vulnerable to eradication.

Wyoming↗

Oil and gas development influences big-game hunting in Wyoming

Development from extracting oil and gas resources can have unintended effects on multiple ecosystem functions, with cascading effects on wildlife, ecosystem services, and local economies. Big-game hunting opportunities may be closely related to these effects, but empirical analyses of impacts of energy development on hunting are limited. We examined the influence of oil and gas development density on harvest efficiency, or harvest per unit of hunter effort, within all hunt areas in Wyoming, USA, from 2008 to 2014 for 3 big-game species: elk ( Cervus canadensis ), mule deer ( Odocoileus hemionus ), and pronghorn ( Antilocapra americana ). Using harvest/hunter day as the response variable, we compared linear mixed-effects models for each species that included total well density (i.e., all wells constructed up to the year of record), active well density (i.e., only those wells currently producing oil or gas in that year), or neither as a predictor variable. We used well densities as indicators of development in the absence of data specifying the locations of other oil and gas infrastructure (e.g., roads, well pads). Models also accounted for the fixed effects of road density, hunter density, proportion of the area that is public land with unrestricted hunter access, proportion of the area that is forested, year of observation, and random effects of variation among hunt areas nested within associated game herd units. Presence of oil and gas wells had a positive influence on harvest efficiency for elk and mule deer. Although there was no overall effect to pronghorn, there was a negative influence of wells on juvenile pronghorn harvest efficiency. Changes in harvest efficiency due to expanding oil and gas development could alter the time spent hunting by hunters and their chances of harvesting an animal. This could have subsequent impacts on hunter satisfaction, game populations, and economic revenue generated from recreational hunters.

Wyoming↗

Estimated nitrogen and phosphorus inputs to the Fish Creek watershed, Teton County, Wyoming, 2009–15

Nutrients, such as nitrogen and phosphorus, are essential for plant and animal growth and nourishment, but the overabundance of bioavailable nitrogen and phosphorus in water can cause adverse health and ecological effects. It is generally accepted that increased primary production of surface-water bodies because of high inputs of nutrients is now the most important polluting effect in surface water in the developed world.

Teton County↗

Forecasting climate change impacts on plant populations over large spatial extents

Plant population models are powerful tools for predicting climate change impacts in one location, but are difficult to apply at landscape scales. We overcome this limitation by taking advantage of two recent advances: remotely sensed, species-specific estimates of plant cover and statistical models developed for spatiotemporal dynamics of animal populations. Using computationally efficient model reparameterizations, we fit a spatiotemporal population model to a 28-year time series of sagebrush ( Artemisia spp.) percent cover over a 2.5 × 5 km landscape in southwestern Wyoming while formally accounting for spatial autocorrelation. We include interannual variation in precipitation and temperature as covariates in the model to investigate how climate affects the cover of sagebrush. We then use the model to forecast the future abundance of sagebrush at the landscape scale under projected climate change, generating spatially explicit estimates of sagebrush population trajectories that have, until now, been impossible to produce at this scale. Our broadscale and long-term predictions are rooted in small-scale and short-term population dynamics and provide an alternative to predictions offered by species distribution models that do not include population dynamics. Our approach, which combines several existing techniques in a novel way, demonstrates the use of remote sensing data to model population responses to environmental change that play out at spatial scales far greater than the traditional field study plot.

Wyoming↗

The extra mile: Ungulate migration distance alters the use of seasonal range and exposure to anthropogenic risk

Partial migration occurs across a variety of taxa and has important ecological and evolutionary consequences. Among ungulates, studies of partially migratory populations have allowed researchers to compare and contrast performance metrics of migrants versus residents and examine how environmental factors influence the relative abundance of each. Such studies tend to characterize animals discretely as either migratory or resident, but we suggest that variable migration distances within migratory herds are an important and overlooked form of population structure, with potential consequences for animal fitness. We examined whether the variation in individual migration distances (20–264 km) within a single wintering population of mule deer ( Odocoileus hemionus ) was associated with several critical behavioral attributes of migration, including timing of migration, time allocation to seasonal ranges, and exposure to anthropogenic mortality risks. Both the timing of migration and the amount of time animals allocated to seasonal ranges varied with migration distance. Animals migrating long distances (150–250 km) initiated spring migration more than three weeks before than those migrating moderate (50–150 km) or short distances (<50 km). Across an entire year, long-distance migrants spent approximately 100 more days migrating compared to moderate- and short-distance migrants. Relatedly, winter residency of long-distance migrants was 71 d fewer than for animals migrating shorter distances. Exposure to anthropogenic mortality factors, including highways and fences, was high for long-distance migrants, whereas vulnerability to harvest was high for short- and moderate-distance migrants. By reducing the amount of time that animals spend on winter range, long-distance migration may alleviate intraspecific competition for limited forage and effectively increase carrying capacity. Clear differences in winter residency, migration duration, and risk of anthropogenic mortality among short-, moderate-, and long-distance migrants suggest fitness trade-offs may exist among migratory segments of the population. Future studies of partial migration may benefit from expanding comparisons of residents and migrants, to consider how variable migration distances of migrants may influence the costs and benefits of migration.

Wyoming↗