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Research about Wyoming

Source-linked reports with geographic coverage including Wyoming.

At least 289 records · Page 16Linked to original sources

Functional attributes of ungulate migration: Landscape features facilitate movement and access to forage

Long‐distance migration by terrestrial mammals is a phenomenon critical to the persistence of populations, but such migrations are declining globally because of over‐harvest, habitat loss, and movement barriers. Increasingly, there is a need to improve existing routes, mitigate route segments affected by anthropogenic disturbance, and in some instances, determine whether alternative routes are available. Using a hypothesis‐driven approach, we identified landscape features associated with the primary functional attributes, stopovers and movement corridors, of spring migratory routes for mule deer in two study areas using resource selection functions. Patterns of selection for landscape attributes of movement corridors and stopovers mostly were similar; however, landscape features associated with movement corridors aligned better with areas that facilitated movement, whereas selection of stopovers was consistent with sites offering early access to spring forage. For movement corridors, deer selected for dry sites, low elevation, and low anthropogenic disturbance. For stopovers, deer selected for dry sites, with consistently early green‐up across years, south‐southwesterly aspects, low elevation, and low anthropogenic disturbance. Stopovers and movement corridors of a migratory route presumably promote different functions, but for a terrestrial migrant, patterns of habitat selection indicate that the same general habitat attributes may facilitate both movement and foraging in spring. Our findings emphasize the roles of topographical wetness, vegetation phenology, and anthropogenic disturbance in shaping use of the landscape during migration for this large herbivore. Avoiding human disturbance and tracking ephemeral forage resources appear to be a consistent pattern during migration, which reinforces the notion that movement during migration has a nutritional underpinning and disturbance potentially alters the net benefits of migration.

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Exploring relationships of spring green-up to moisture and temperature across Wyoming, U.S.A

Vegetation green-up signals the timing of available nutritious plants and shrubs providing high-quality forage for ungulates. In this study, we characterized spatial and temporal patterns of spring phenology and explored how they were related to preceding temperature and moisture conditions. We tested correlations between late winter weather and indicators of the onset and the length of the spring growing period with 250-m resolution time-series satellite data (2001 – 2013) for Wyoming, USA. In western Wyoming mountains, drier and warmer conditions during late winter were associated with earlier spring green-up onset of growth in forests, shrubs, and grasses. In the northeast mountains, onset of spring correlated positively with preceding warmer temperatures, but not with precipitation. In most basin and plains shrublands and grasslands, spring onset was not correlated with temperature, although earlier onset of spring was correlated with drier conditions in 25% of shrub/scrub areas. Results about the length of spring were less definitive, with warmer temperatures related to longer green-up time for 12–30% of the land cover in western mountains but to shorter green-up time periods for 10–20% of the grasses and shrubs in basins and plains. Complex phenological patterns are likely to affect ungulate foraging behaviour on a local scale.

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Testing infrared camera surveys and distance analyses to estimate feral horse abundance in a known population

We tested the use of high‐resolution infrared (IR) camera technology and distance sampling analyses to estimate abundance of feral horses ( Equus caballus ) during 2015–2016 in the McCullough Peaks Herd Management Area, Wyoming, USA. Infrared technology is becoming more common in ungulate population monitoring. The quality of IR cameras now allows ungulate species to be differentiated. Imperfect detection is a common problem in aerial surveys, so we tested the use of distance sampling analyses to account for imperfect detection probability. We conducted 2 aerial surveys in a sagebrush ecosystem with a demographically closed horse population. True abundance was known to within ±4 animals as a result of intensive, ground‐based monitoring of each animal, all of which are uniquely identifiable. After truncation of our data, the most supported detection function was a uniform function with a detection probability equal to 1.0 out to 255 m. Our analyses yielded results that were within 10% of true abundance, but the coefficient of variation (CV) was large (36–58%) assuming a small sampling fraction. However, our truncated surveys covered approximately 95% of the herd management area. By including a finite population correction factor in our calculations of variance estimates, CVs (8–13%) were dramatically reduced. We found the combination of IR surveys and distance sampling analysis to be a useful method to estimate feral horse abundance in sagebrush vegetation type, which had limited cover to obscure horses. Repeated testing in sagebrush ecosystems as well as further testing in other habitat types and under differing conditions will inform how general our approach can be.

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The interaction of exposure and warming tolerance determines fish species vulnerability to warming stream temperatures

Species vulnerability to climate change involves an interaction between the magnitude of change (exposure) and a species's tolerance to change. We evaluated fish species vulnerability to predicted stream temperature increases by examining warming tolerances across the Wyoming fish assemblage. Warming tolerance combines stream temperature with a thermal tolerance metric to estimate how much warming beyond current conditions a species can withstand. Brown trout, rainbow trout and burbot had the lowest warming tolerances and the highest proportion of currently occupied sites that will become unsuitable under predicted temperature increases. These most vulnerable species were coldwater species, but had neither the lowest thermal tolerances nor would they experience the greatest temperature increases. Our results highlight the importance of considering the interaction of exposure and warming tolerance when predicting climate change vulnerability and demonstrate an approach that can be applied broadly.

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Modeling golden eagle‐vehicle collisions to design mitigation strategies

The incidental take of golden eagles ( Aquila chrysaetos ) as a result of wind energy development requires some form of compensatory mitigation. Although several options have been proposed, only one has been formerly accepted and implemented, and the lack of options can limit the permit process for wind facilities. We developed a model to estimate numbers of golden eagles that die when struck by vehicles when eagles scavenge road kill to evaluate removal of road‐killed carcasses as an additional mitigation option. Our model estimates vehicle collision rates as a function of eagle densities, road traffic volume, and animal carcass abundance at the scale of a Wyoming, USA, county during fall‐winter, and quantifies the effects of different mitigation strategies, including estimates of uncertainty. We evaluated the plausibility of our model estimates by predicting mortality rates for each county in Wyoming and comparing overall state mortality to current estimates of mortality using derived estimates from expert judgment. We also developed a context‐dependent analysis of potential mitigation credits controlling for carcass number, traffic volume, and background carcass removals. We found that mitigation credit should be highest in areas with greatest number of carcasses. Collision mitigation is a potentially useful addition to the mitigation toolbox for wind energy development or other activities that need to offset predicted golden eagle mortality and satisfy incidental take permit requirements.

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Evaluating relationships between native fishes and habitat in streams affected by oil and natural gas development

Oil and natural gas (ONG) development can affect aquatic ecosystems through water contamination, water withdrawals and disturbance of soil and vegetation (surface disturbance) from infrastructure development. Research on how these potential sources of watershed and aquatic ecosystem impairment can affect fish assemblages is limited. Fish–habitat relationships were evaluated across stream sites experiencing differing levels of ONG development. Colorado River cutthroat trout, Oncorhynchus clarkii pleuriticus (Cope), and mottled sculpin, Cottus bairdii Girard, presence and abundance were associated with habitat conditions predominantly found in the less disturbed streams, such as higher proportion of shrub cover, greater stream depths and gravel substrate. Mountain sucker, Catostomus platyrhynchus (Cope), appeared to be a habitat generalist and was able to persist in a wide range of conditions, including degraded sites. Natural resource managers can use habitat preferences of these fish species to establish the development plans that mitigate negative effects of ONG development by protecting the aquatic habitats they rely upon.

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Combining genetic, isotopic, and field data to better describe the influence of dams and diversions on Burbot Movement in the Wind River Drainage, Wyoming

Dams and water diversions fragment habitat, entrain fish, and alter fish movement. Many Burbot Lota lota populations are declining, with dams and water diversions thought to be a major threat. We used multiple methods to identify Burbot movement patterns and assess entrainment into an irrigation system in the Wind River, Wyoming. We assessed seasonal movement of Burbot with a mark–recapture (PIT tagging) study, natal origins of entrained fish with otolith microchemistry, and historic movement with genotyping by sequencing. We found limited evidence of entrainment in irrigation waters across all approaches. The mark–recapture study indicated that out‐migration from potential source populations could be influenced by flow regime but was generally low. Otolith and genomic results suggested the presence of a self‐sustaining population within the irrigation network. We conclude that emigration from natural tributary populations is not the current source of the majority of Burbot found in irrigation waters. Instead, reservoir and irrigation canal construction has created novel habitat in which Burbot have established a population. Using a multi‐scale approach increased our inferential abilities and mechanistic understanding of movement patterns between natural and managed systems.

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Contrasting perspectives on the Lava Creek Tuff eruption, Yellowstone, from new U–Pb and 40Ar/39Ar age determinations

The youngest major caldera-forming event at Yellowstone was the ~ 630-ka eruption of the Lava Creek Tuff. The tuff as mapped consists of two major ignimbrite packages (members A and B), linked to widespread coeval fall deposits and formation of the Yellowstone Caldera. Subsequent activity included emplacement of numerous rhyolite flows and domes, and development of two structurally resurgent domes (Mallard Lake and Sour Creek) that accommodate strain due to continual uplift/subsidence cycles. Uplifted lithologies previously mapped on and adjacent to Sour Creek dome were thought to include the ~ 2.08-Ma Huckleberry Ridge Tuff, cropping out beneath Lava Creek Tuff members A and B. Mapped outcrops of this Huckleberry Ridge Tuff material were sampled as welded ignimbrite (sample YR345) on Sour Creek dome, and at nearby Bog Creek as welded ignimbrite (YR311) underlain by an indurated lithic lag breccia containing blocks of another welded ignimbrite (YR324). Zircon near-rim U–Pb analyses from these samples yield weighted mean ages of 661 ± 13 ka (YR345: 95% confidence), 655 ± 11 ka (YR311), and 664 ± 15 ka (YR324) (combined weighted mean of 658.8 ± 6.6 ka). We also studied two samples of ignimbrite previously mapped as Huckleberry Ridge Tuff on the northeastern perimeter of the Yellowstone Caldera, ~ 12 km ENE of Sour Creek dome. Sanidines from these samples yield 40 Ar/ 39 Ar age estimates of 634.5 ± 6.8 ka (8YC-358) and 630.9 ± 4.1 ka (8YC-359). These age data show that all these units represent previously unrecognized parts of the Lava Creek Tuff and do not have any relationship to the Huckleberry Ridge Tuff. Our observations and data imply that the Lava Creek eruption was more complex than is currently assumed, incorporating two tuff units additional to those currently mapped, and which themselves are separated by a time break sufficient for cooling and some reworking. The presence of a lag breccia suggests that a source vent lay nearby (< ~ 3 km) for some of the tuffs and that the Yellowstone Caldera boundary in this area could be reconsidered.

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U.S. Geological Survey science for the Wyoming Landscape Conservation Initiative—2016 annual report

This is the ninth annual report highlighting U.S. Geological Survey (USGS) science and decision-support activities conducted for the Wyoming Landscape Conservation Initiative (WLCI). The activities address specific management needs identified by WLCI partner agencies. In fiscal year (FY) 2016, there were 26 active USGS WLCI science-based projects. Of these 26 projects, one project was new for FY2016, and three were completed by the end of the fiscal year (though final products were still in preparation or review). USGS WLCI projects were grouped under five categories: (1) Baseline Synthesis, (2) Long-Term Monitoring, (3) Effectiveness Monitoring, (4) Mechanistic Studies of Wildlife, and (5) Data and Information Management. Each of these topic areas is designed to address WLCI management needs: identifying key drivers of change, identifying the condition and distribution of key wildlife species and habitats and of species’ habitat requirements, development of an integrated inventory and monitoring strategy, use of emerging technologies and development and testing of innovative methods for maximizing the efficiency and efficacy of monitoring efforts, evaluating the effectiveness of habitat treatment projects, evaluating the responses of wildlife to development, and developing a data clearinghouse and information management framework to support and provide access to results of most USGS WLCI projects. In FY2016, we assisted with updating the WLCI Conservation Action Plan and associated databases as part of the Comprehensive Assessment, and we also assisted with the Bureau of Land Management 2015 WLCI annual report. By the end of FY2016, we completed or had nearly completed assessments of WLCI energy and mineral resources and had submitted a manuscript on modeled effects of oil and gas development on wildlife to a peer-reviewed journal. We also initiated a study on the effects of wind energy on wildlife in the WLCI region. A USGS circular on WLCI long-term monitoring was in review at the end of the fiscal year, and seven projects monitoring water and vegetation (including changes in sagebrush cover and patterns of sagebrush mortality) continued through the year. USGS scientists continued many projects in FY2016 that evaluate the effectiveness of habitat conservation actions (including sagebrush, cheatgrass, and aspen habitat treatments) and provide tools in support of mechanistic studies of wildlife. In FY2016, USGS scientists, along with university and State partners, continued work on five focal wildlife species/communities (pygmy rabbits [ Brachylagus idahoensis ], greater sage grouse , mule deer, sagebrush songbirds, and native fish). In FY2016, the USGS Information Management Team presented information to WLCI scientists on how USGS tools and resources can be used to fulfill the requirements of new USGS policies regarding data release, data management, and data visualization.

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A tale of two wildfires; testing detection and prediction of invasive species distributions using models fit with topographic and spectral indices

Context Developing species distribution models (SDMs) to detect invasive species cover and evaluate habitat suitability are high priorities for land managers. Objectives We tested SDMs fit with different variable combinations to provide guidelines for future invasive species model development based on transferability between landscapes. Methods Generalized linear model, boosted regression trees, multivariate adaptive regression splines, and Random Forests were fit with location data for high cheatgrass ( Bromus tectorum ) cover in situ for two post-burn sites independently using topographic indices, spectral indices derived from multiple dates of Landsat 8 satellite imagery, or both. Models developed for one site were applied to the other, using independent cheatgrass cover data from the respective ex situ site to test model transferability. Results Fitted models were statistically robust and comparable when fit with at least 200 cover plots in situ and transferred to the ex situ site. Only the Random Forests models were robust when fit with a small number of cover plots in situ. Conclusions Our study indicated spectral indices can be used in SDMs to estimate species cover across landscapes (e.g., both within the same Landsat scene and in an adjacent Landsat scene). Important considerations for transferability include the model employed, quantity of cover data used to train/test the models, and phenology of the species coupled with the timing of imagery. The results also suggest that when cover data are limited, SDMs fit with topographic indices are sufficient for evaluating cheatgrass habitat suitability in new post-disturbance landscapes; however, spectral indices can provide a more robust estimate for detection based on local phenology.

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Occupancy modeling of Parnassius clodius butterfly populations in Grand Teton National Park, Wyoming

Estimating occupancy patterns and identifying vegetation characteristics that influence the presence of butterfly species are essential approaches needed for determining how habitat changes may affect butterfly populations in the future. The montane butterfly species, Parnassius clodius , was investigated to identify patterns of occupancy relating to habitat variables in Grand Teton National Park and Bridger-Teton National Forest, Wyoming, United States. A series of presence–absence surveys were conducted in 2013 in 41 mesic to xeric montane meadows that were considered suitable habitat for P. clodius during their flight season (June–July) to estimate occupancy ( ψ ) and detection probability ( p ). According to the null constant parameter model, P. clodius had high occupancy of ψ = 0.78 ± 0.07 SE and detection probability of p = 0.75 ± 0.04 SE. In models testing covariates, the most important habitat indicator for the occupancy of P. clodius was a strong negative association with big sagebrush ( Artemisia tridentata ; β = − 21.39 ± 21.10 SE) and lupine ( Lupinus spp.; β = − 20.03 ± 21.24 SE). While P. clodius was found at a high proportion of meadows surveyed, the presence of A. tridentata may limit their distribution within montane meadows at a landscape scale because A. tridentata dominates a large percentage of the montane meadows in our study area. Future climate scenarios predicted for high elevations globally could cause habitat shifts and put populations of P. clodius and similar non-migratory butterfly populations at risk.

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Efficacy of using data from angler-caught Burbot to estimate population rate functions

The effective management of a fish population depends on the collection of accurate demographic data from that population. Since demographic data are often expensive and difficult to obtain, developing cost‐effective and efficient collection methods is a high priority. This research evaluates the efficacy of using angler‐supplied data to monitor a nonnative population of Burbot Lota lota . Age and growth estimates were compared between Burbot collected by anglers and those collected in trammel nets from two Wyoming reservoirs. Collection methods produced different length‐frequency distributions, but no difference was observed in age‐frequency distributions. Mean back‐calculated lengths at age revealed that netted Burbot grew faster than angled Burbot in Fontenelle Reservoir. In contrast, angled Burbot grew slightly faster than netted Burbot in Flaming Gorge Reservoir. Von Bertalanffy growth models differed between collection methods, but differences in parameter estimates were minor. Estimates of total annual mortality ( A ) of Burbot in Fontenelle Reservoir were comparable between angled ( A = 35.4%) and netted fish (33.9%); similar results were observed in Flaming Gorge Reservoir for angled (29.3%) and netted fish (30.5%). Beverton–Holt yield‐per‐recruit models were fit using data from both collection methods. Estimated yield differed by less than 15% between data sources and reservoir. Spawning potential ratios indicated that an exploitation rate of 20% would be required to induce recruitment overfishing in either reservoir, regardless of data source. Results of this study suggest that angler‐supplied data are useful for monitoring Burbot population dynamics in Wyoming and may be an option to efficiently monitor other fish populations in North America.

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Identifying holes in the greater sage-grouse conservation umbrella

The umbrella species concept, wherein multiple species are indirectly protected under the umbrella of a reserve created for one, is intended to enhance conservation efficiency. Although appealing in theory and common in practice, empirical tests of the concept have been scarce. We used a real-world, semi-protected reserve established to protect a high-profile umbrella species (greater sage-grouse [ Centrocercus urophasianus ]) to investigate 2 potential mechanisms underlying the concept's successful application: reserve size and species similarity. We estimated how much habitat protection the established reserve provided to 52 species of conservation concern associated with vegetation communities where greater sage-grouse occur. To illustrate the importance of reserve size, we compared the effectiveness of the established reserve to alternative greater sage-grouse reserves of various sizes and to simulated reserves of equal size but sited with no regard for greater sage-grouse. We further assessed whether key species’ traits were associated with different levels of protection under the umbrella reserve. The established umbrella reserve protected 82% of the state's greater sage-grouse population and 0–63% of the habitat of the background species examined. The reserve outperformed equally sized, simulated reserves for only 12 of 52 background species. As expected, larger alternative reserves served as better umbrellas, but regardless of reserve size, not all species received equal protection. The established reserve was most effective at protecting the habitat of species that were most similar to the umbrella species (i.e., avian species, those highly associated with sagebrush plant communities, and those with widespread habitat). In contrast, the habitat of species with restricted distributions, particularly when combined with vegetation associations not closely matching the umbrella species, was not protected as well by the umbrella reserve. Such species require additional, targeted attention to achieve conservation objectives. Successful application of the umbrella species concept requires careful consideration of the characteristics of the umbrella species, the reserve delineated on its behalf, and the similarity of the umbrella species to its purported background species.

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Winter feeding of elk in the Greater Yellowstone Ecosystem and its effects on disease dynamics

Providing food to wildlife during periods when natural food is limited results in aggregations that may facilitate disease transmission. This is exemplified in western Wyoming where institutional feeding over the past century has aimed to mitigate wildlife–livestock conflict and minimize winter mortality of elk ( Cervus canadensis ). Here we review research across 23 winter feedgrounds where the most studied disease is brucellosis, caused by the bacterium Brucella abortus . Traditional veterinary practices (vaccination, test-and-slaughter) have thus far been unable to control this disease in elk, which can spill over to cattle. Current disease-reduction efforts are being guided by ecological research on elk movement and density, reproduction, stress, co-infections and scavengers. Given the right tools, feedgrounds could provide opportunities for adaptive management of brucellosis through regular animal testing and population-level manipulations. Our analyses of several such manipulations highlight the value of a research–management partnership guided by hypothesis testing, despite the constraints of the sociopolitical environment. However, brucellosis is now spreading in unfed elk herds, while other diseases (e.g. chronic wasting disease) are of increasing concern at feedgrounds. Therefore experimental closures of feedgrounds, reduced feeding and lower elk populations merit consideration.

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Green‐wave surfing increases fat gain in a migratory ungulate

Each spring, migratory herbivores around the world track or ‘surf’ green waves of newly emergent vegetation to distant summer or wet‐season ranges. This foraging tactic may help explain the great abundance of migratory herbivores on many seasonal landscapes. However, the underlying fitness benefits of this life‐history strategy remain poorly understood. A fundamental prediction of the green‐wave hypothesis is that migratory herbivores obtain fitness benefits from surfing waves of newly emergent vegetation more closely than their resident counterparts. Here we evaluate whether this behavior increases body‐fat levels – a critically important correlate of reproduction and survival for most ungulates – in elk Cervus elaphus of the Greater Yellowstone Ecosystem. Using satellite imagery and GPS tracking data, we found evidence that migrants (n = 23) indeed surfed the green wave, occupying sites 12.7 days closer to peak green‐up than residents (n = 16). Importantly, individual variation in surfing may help account for up to 6 kg of variation in autumn body‐fat levels. Our findings point to a pathway for anthropogenic changes to the green wave (e.g. climate change) or migrants’ ability to surf it (e.g. development) to impact migratory populations. To explore this possibility, we evaluated potential population‐level consequences of constrained surfing with a heuristic model. If green‐wave surfing deteriorates by 5–15 days from observed, our model predicts up to a 20% decrease in pregnancy rates, a 2.5% decrease in population growth, and a 30% decrease in abundance over 50 years. By linking green‐wave surfing to fitness and illustrating potential effects on population growth, our study provides new insights into the evolution of migratory behavior and the prospects for the persistence of migratory ungulate populations in a changing world.

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Pleistocene glaciation of the Jackson Hole area, Wyoming

Pleistocene glaciations and late Cenozoic offset on the Teton fault have played central roles in shaping the scenic landscapes of the Teton Range and Jackson Hole area in Wyoming. The Teton Range harbored a system of mountain-valley glaciers that produced the striking geomorphic features in these mountains. However, the comparatively much larger southern sector of the Greater Yellowstone glacial system (GYGS) is responsible for creating the more expansive glacial landforms and deposits that dominate Jackson Hole. The glacial history is also inextricably associated with the Yellowstone hotspot, which caused two conditions that have fostered extensive glaciation: (1) uplift and consequent cold temperatures in greater Yellowstone; and (2) the lowland track of the hotspot (eastern Snake River Plain) that funneled moisture to the Yellowstone Plateau and the Yellowstone Crescent of High Terrain (YCHT). The penultimate (Bull Lake) glaciation filled all of Jackson Hole with glacial ice. Granitic boulders on moraines beyond the south end of Jackson Hole have cosmogenic 10Be exposure ages of ~150 thousand years ago (ka) and correlate with Marine Isotope Stage 6. A thick loess mantle subdues the topography of Bull Lake moraines and caps Bull Lake outwash terraces with a reddish buried soil near the base of the loess having a Bk horizon that extends down into the outwash gravel. The Bull Lake glaciation of Jackson Hole extended 48 kilometers (km) farther south than the Pinedale, representing the largest separation of these two glacial positions in the Western United States. The Bull Lake is also more extensive than the Pinedale on the west (22 km) and southwest (23 km) margins of the GYGS but not on the north and east. This pattern is explained by uplift and subsidence on the leading and trailing “bow-wave” of the YCHT, respectively. During the last (Pinedale) glaciation, mountain-valley glaciers of the Teton Range extended to the western edge of Jackson Hole and built bouldery moraines that commonly enclose lakes. On the southern margin of the GYGS, prominent glacial outwash terraces define three phases of the Pinedale glaciation in Jackson Hole: Pinedale-1 (Pd-1) by Antelope Flats with subdued channel patterns on the east side of Jackson Hole; Pinedale-2 (Pd-2) by a large outwash fan that includes Baseline Flat on the west side of Jackson Hole with well-defined channel patterns; and Pinedale-3 (Pd-3) by The Potholes and other outwash fans farther up the Snake River in central Jackson Hole. During Pinedale glaciation, three glacial lobes of the GYGS fed into Jackson Hole, and the relative importance of these lobes changed dramatically through time. During the Pd-1 glaciation, the eastern Buffalo Fork lobe dominated whereas in Pd-2 and Pd-3 time the northern Snake River lobe dominated. This is consistent with migration of the GYGS center of ice mass westward and southward as glaciers built up towards the moisture source provided by storms moving northeastward up the eastern Snake River Plain. The recession of the eastern Buffalo Fork lobe in Pd-2 and Pd-3 times is consistent with an enlarged ice mass on the Yellowstone Plateau that placed the eastern part of the GYGS in a precipitation or snow shadow. In Pd-1 time, the Buffalo Fork lobe reached its maximum extent and was joined by the Pacific Creek lobe. This culmination may correlate with the ~21–18 ka ages of moraines in the Teton Range and nearby ranges. Three subdivisions of Pd-1 glaciation built moraines that are nearly or entirely covered by outwash almost 100 meters thick. In Pd-2 time, the Snake River lobe joined with the Pacific Creek lobe and built a large outwash fan south of the present-day Jackson Lake. Boulders on a moraine at the head of this fan are dated to 15.5 ± 0.5 ka. The relation between Teton glaciers and those of the GYGS is indicated by outwash from these Pd-2 moraines that partly buries outer Jenny Lake moraines dated to 15.2 ± 0.7 ka. East of the large outwash fan, Pd-2 ice advanced across the glacial-age Triangle X-2 lake sediments, perhaps in a surge. The Buffalo Fork lobe retreated more than 20 km up valley from its Pd-1 position and Pd-2 ice of the Snake River and Pacific Creek lobes advanced into the area previously occupied by the Buffalo Fork lobe. The Pd-3 position flanks the margin of Jackson Lake and represents a retreat to a stable position after the Pd-2 7-km advance that may have been a surge across the Triangle X-2 lake sediments. The Potholes and South Landing outwash fans were built in the area deglaciated by the retreat from Pd-2 to Pd-3 time. The Spalding Bay outwash fan continued to incise and a meltwater stream flowed just outside the Teton glacier that filled the present Jenny Lake and deposited the 14.4 ± 0.8 ka inner Jenny Lake moraines. Glacial outwash terraces increase in slope toward their respective moraines of the GYGS and are complex in both north-south and east-west directions. The Pd-1 terrace slopes to the west where it is buried by the Pd-2 outwash. The post-depositional tilting of the Pd-1 outwash terrace is an order of magnitude smaller than the original westward depositional slope. The Pd-1, 2, and 3 terraces have a shingle-like geometry such that the highest terrace decreases in age down valley, and in southern Jackson Hole, the Pd-3 terrace is only 3–5 m above the Snake River. In Pd-1 time the combined Buffalo Fork and Pacific Creek lobes scoured out four basins: (1) Emma Matilda Lake; (2) Two Ocean Lake; (3) a deep basin from lower Pacific Creek to beneath the Oxbows and Jackson Lake Dam; and (4) the largest basin from the lower Buffalo Fork to Deadmans Bar of the Snake River. These basins are largely filled with fine-grained sediment and are now marked by moist lowlands or lakes. In Pd-2 and Pd-3 time the Snake River lobe scoured the present 120-m deep Jackson Lake and possibly the 120-m deeper sediment-filled basin. Subglacial erosion of the Jackson Lake basin by confined water jets is supported by eskers that climb up to the head of the South Landing outwash fan.

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Paleolimnological records of nitrogen deposition in shallow, high-elevation lakes of Grand Teton National Park, Wyoming, USA

Reactive nitrogen (Nr) from anthropogenic sources has been altering ecosystem function in lakes of the Rocky Mountains, other regions of western North America, and the Arctic over recent decades. The response of biota in shallow lakes to atmospheric deposition of Nr, however, has not been considered. Benthic algae are dominant in shallow, high-elevation lakes and are less sensitive to nutrient inputs than planktonic algae. Because the benthos is typically more nutrient rich than the water column, shallow lakes are not expected to show evidence of anthropogenic Nr. In this study, we assessed sedimentary evidence for regional Nr deposition, sediment chronology, and the nature of algal community response in five shallow, high-elevation lakes in Grand Teton National Park (GRTE). Over 140 diatom taxa were identified from the sediments, with a relatively high species richness of taxa characteristic of oligotrophic conditions. The diatom assemblages were dominated by benthic taxa, especially motile taxa. The GRTE lakes demonstrate assemblage-wide shifts in diatoms, including 1) synchronous and significant assemblage changes centered on ~1960 AD; 2) pre-1960 assemblages differed significantly from post-1960 assemblages; 3) pre-1960 diatom assemblages fluctuated randomly, whereas post- 1960 assemblages showed directional change; 4) changes in &delta;15N signatures were correlated with diatom community composition. These results demonstrate recent changes in shallow high18 elevation lakes that are most correlated with anthropogenic Nr. It is also possible, however, that the combined effect of Nr deposition and warming is accelerating species shifts in benthic diatoms. While uncertainties remain about the potential synergy of Nr deposition and warming, this study adds shallow lakes to the growing list of impacted high-elevation localities in western North America.

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Using a Bayesian framework to account for advection in seven years of snowpack CO2 fluxes in a mortality-impacted subalpine forest

In subalpine forests where snowpacks can persist for 50% of the year or longer, wintertime subnivean soil respiration constitutes a significant, but poorly understood, portion of the carbon (C) cycle. The temporal and seasonal dynamics of this flux are difficult to measure compared to snow-free respiration. This makes it challenging to fully address key knowledge gaps in high-elevation forests, including effects of tree mortality on ecosystem C fluxes. Our paper attempts to overcome these challenges by developing a new method for estimating wintertime CO 2 flux from soil underneath winter snowpack based on CO 2 profiles in the snow and using this to examine potential effects of a tree mortality event on wintertime respiration. The chief problem for estimates based on snow CO 2 measurements is that during strong winds, vertical CO 2 profiles can depart significantly from linearity, resulting in CO 2 fluxes that are not steady-state diffusive fluxes. Consequently, these fluxes include an advective component induced by wind and atmospheric pressure, resulting in above-snowpack fluxes that may be temporally decoupled from biological flux production in the soil. These advective fluxes lead to errors in estimates of the total amount of CO 2 produced from the soil surface during the winter, and may complicate the detection of significant seasonal and interannual trends in snowpack CO 2 fluxes. Our goal was twofold: first, to improve methods for estimating wintertime subnivean soil respiration; and second, to use this new method to examine potential effects of forest mortality on respiration. We develop a Bayesian-based model of CO 2 snowpack profiles and accounts for CO 2 storage flux within the snowpack to calculate diffusive and advective fluxes from both the bottom and the top of the snowpack. We apply this model to hourly snowpack CO 2 profile data collected over 7 years at an AmeriFlux eddy covariance (EC) site in a Wyoming subalpine forest that had recently experienced a mass tree mortality event. Hourly advective fluxes from the snowpack surface are less variable over time than hourly EC fluxes. Nonetheless, on a daily time scale, the two methods agree well with each other. We found that soil biological CO 2 production entering the bottom of snowpack peaked 3 years after the onset of the bark beetle outbreak and 1-year after dead trees dropped their needles, presenting the first evidence of a short-term increase in respiration following bark beetle mortality. Our findings emphasize the importance of accounting for the role of wind-driven advection in wintertime respiration and suggest that tree mortality may nearly double wintertime respiration in the first few years.

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