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Research about Pennsylvania

Source-linked reports with geographic coverage including Pennsylvania.

At least 19 recordsLinked to original sources

The effect of temperature, flow, density and disease on smallmouth bass (Micropterus dolomieu) early life growth rate

Individual growth rates of fish are influenced by gradually changing environmental conditions and rapid-onset events. However, there is a paucity of information on drivers of spatiotemporal variability in growth rates of fishes across large spatial extents. Using a 36-year dataset of smallmouth bass ( Micropterus dolomieu ) length-at-age data across 10 river reaches in Pennsylvania, USA ( n = 54,068 individuals), we estimated annual early life growth rates (mm year −1 ) and assessed the effects of relative abundance, summer temperature, summer streamflow rate and a disease outbreak on growth rate. We found that temperature had a positive effect on growth rate, varied spatially, and had a stronger effect in diseased reaches. There was also evidence for an impact of disease on early life growth through density-dependent mechanisms, with growth rates increasing from ∼85 to 101 mm·year −1 following disease-related decreases in abundance. This study adds to our understanding of the factors that shape early life growth, including rapid-onset events like disease which can have immediate and lasting effects on growth rate trajectory.

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3D Elevation Program—Pennsylvania

Using 3D Elevation Program data, a modern illustration of each State's landscape showcases its elevation differences in vivid detail. Each State has been consistently adjusted and colorized according to its highest and lowest elevations, accentuating differences even in States with low relief. To further emphasize landforms, shaded relief has been added on top of the elevation colors.

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Sediment response to stream restoration in an agricultural watershed

Despite widespread implementation of stream restoration there is limited research of its effectiveness in impaired agricultural settings. To quantify the effects of stream restoration on sediment load, we investigated erosion and deposition along 9 stream reach sites (4 restored, 4 unrestored, and 1 downstream) in Turtle Creek, Pennsylvania, United States - a representative agricultural subbasin of the Chesapeake Bay. These stream restoration projects had the goal of reducing sediment load primarily by installing log structures and regrading streambanks in order to stabilize streambanks and reconnect floodplains. Floodplain and streambank sediment fluxes were quantified over a 3-year period through repeated measurement of clay pads and pins and measurement of sediment bulk density. Uncrewed aerial imagery was collected at 3 of the sites, then processed using structure from motion (SfM) to create high-resolution digital elevation models. Terrain analysis enabled three-dimensional understanding of the generality of the specific field flux measurement locations to the larger stream reach. Restored sites on average had deposition on streambanks compared to streambank erosion along unrestored sites, and approximately two times greater floodplain deposition than unrestored sites. Overall, restored stream had an average seven-fold increase in the net balance (sum of streambank and floodplain) of sediment flux, a 81.2 kg/m/yr increase in retention, compared to unrestored stream. The measured larger streambed particle size distribution (D 50 ) in restored streams also suggests additional improvements to instream habitat quality. Terrain analysis confirmed that the field-based measurements were representative of the larger reach. These results indicate that stream restoration in agricultural watersheds using less-engineered designs can reduce sediment loading and improve stream habitat and health goals.

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Comparison of data handling techniques for modeling bat acoustic activity

With the proliferation of acoustic sampling to investigate bat distribution and ecology, researchers have implemented a myriad of statistical modeling approaches to interpret findings. Bats are taxa of high conservation concern; therefore, ensuring the accuracy of species-level habitat association models is critical for informing management. We sought to determine prediction differences among statistical approaches to modeling counts of acoustic detections, using generalized linear mixed models with 8 acoustic data-handling techniques. We applied each approach or combination of approaches to a rare species, the northern long-eared bat ( Myotis septentrionalis ), and a common species, the eastern red bat ( Lasiurus borealis ), from summer survey results on a landscape in south-central Pennsylvania, USA, 2024. We evaluated the accuracy of habitat association models of bat acoustic activity at the species level using cross-validation and compared resulting predictions of models using spatial correlations. We determined that filtering data by automated identification software (Kaleidoscope Pro), the maximum likelihood estimate (MLE) P -value thresholds reduced relative mean absolute error (rMAE) in cross-validation of northern long-eared bat models. Using the MLE-retained data produced the most accurate predictions over using raw data or the overly conservative match ratio data. However, for the eastern red bat, the results from the most conservative approach of only retaining data with at least a 90% match ratio from software development training sets had the lowest rMAE. We have provided evidence that the current standard of filtering data by nightly MLE can result in more accurate and informative habitat-use acoustic activity models for rare bat species.

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Incorporating data sets with multiple sources of uncertainty in integrated species distribution models

Data integration methods aim to improve species distribution estimates by incorporating multiple sources of uncertainty across datasets. Two major sources of uncertainty are: (1) variation in sampling effort across space and within datasets, and (2) variation in reliability associated with data collection protocols or timing among datasets. Our goal was to evaluate how different approaches to address these uncertainties influence predictive performance of integrated models. We modeled distributions of four bird species using three datasets that differed in sampling design. We examined three strategies to reduce uncertainty: (1) filtering data, (2) incorporating functions that account for uncertainty in observation models, and (3) varying how datasets are integrated into a single estimate. We first examine methods to account for variable effort in observations, focusing on both spatial differences in sampling intensity and effort given to a single observation record. We then examine approaches to account for data sets with differing reliability. Sampling effort was best addressed through conservative filtering, including spatial thinning and excluding observations with highly variable effort. Next, we considered how to account for potential false positive detections—due to either misidentification or changes in distributions. We found that treating less reliable data as a covariate, an approach previously suggested for data integration that can greatly speed up model fitting, performed well. Other effective approaches included directly modeling false positive rates and complete exclusion of less reliable data sets. Our results provide insights into best practices in integrated modeling for handling uncertainty in integrated models. We demonstrate the flexible options available when using integrated models to address uncertainty.

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Data-collection methods for total dissolved gases monitoring, Youghiogheny River at Dam Outlet Tunnel near Confluence, Pennsylvania

Supersaturation of total dissolved gases (TDG) can potentially occur in the tailrace water at the Youghiogheny River at dam outlet tunnel near Confluence, Pennsylvania (U.S. Geological Survey [USGS] streamgaging and monitoring station 03077100). The USGS, in cooperation with the U.S. Army Corps of Engineers, established methods to collect and report TDG saturation data in the tailrace below the Youghiogheny Dam. Monitoring and TDG data collection started in June 2008 and continues currently (2025). Data are collected from June 1 through November 30 of each year, and these data are used by the U.S. Army Corps of Engineers to guide management of the dam outflow. Methods used for data collection, processing, reporting, and quality assurance for TDG monitored at USGS station 03077100 are presented in this report. The TDG data are publicly available in the USGS National Water Information System database.

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Intraspecific contact among white-tailed deer: A literature review and chronic wasting disease case study

White-tailed deer ( Odocoileus virginianus ) are a valuable game mammal in the eastern United States necessitating detailed understanding of disease transmission. We conducted a literature review on intraspecific contact (i.e., interactions wherein disease transmission may occur) among deer. From 69 studies, we identified five themes underlying research on intraspecific deer contact: physical touch, social groups, spatial overlap, association rates, and social networks. Visual observations determined physical touch to be infrequent (< 2 touches/h) and indicated deer social groups were dependent on spatial dynamics of parturition and dispersal; most females remained with matriarchal family groups while males dispersed and formed bachelor groups. Assessed using global positioning system (GPS) monitoring, spatial overlap and association rates (i.e., instances of deer in close spatial–temporal proximity) were higher in correspondence to within-group social dynamics, and between-group scores were correspondingly low. Social network analyses indicated between-group transmission may be driven by socially dominant males, often termed super-spreaders (i.e., hosts infecting disproportionately high numbers of healthy individuals). We investigated these themes via a case study of deer infected with chronic wasting disease (CWD) in southcentral Pennsylvania, United States. We assessed spatial overlap and association rates using GPS monitoring data from 180 deer. Our results supported findings in the literature, showing strong correlations among spatial overlap, association rates, and correlated movements. Further, CWD-infected deer exhibited similar association rates to deer in which CWD was not detected. Our literature review and case study indicate direct transmission of CWD and other diseases is likely greatest within social groups following seasonal behavioral dynamics and that between-group transmission is likely driven by males via dispersal and mating interactions. Our results may be used to inform population management models with future work focused on high resolution spatial assessments of transmission in localized areas.

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Statewide agent-based model for management of chronic wasting disease in white-tailed deer: PAOvCWD

Chronic wasting disease (CWD) is an always-fatal disease infecting wild cervids globally. Ecologically and economically important, CWD presents a challenge for managing white-tailed deer ( Odocoileus virginianus ). We built an agent-based model to simulate CWD transmission and assess potential management actions that could slow disease spread: PAOvCWD . We developed PAOvCWD using contact rates and other behavioral and ecological metrics estimated from deer monitored in Pennsylvania, USA. We programmed potential management responses (e.g., culling, altered hunter harvest) for all 22 Pennsylvania wildlife management units and validated the efficacy of PAOvCWD using a deer population in south-central Pennsylvania infected with CWD for > 10 years. To support applications of our model, we developed a user-friendly R pipeline that allows implementation with relatively minor modifications. Our pipeline includes four steps: • Steps 1 and 2 generate regional percent forest cover rasters and curate population-level demographics. • Step 3 initializes landscapes and agents using our PAOvPOP model. • Step 4 assesses CWD transmission and management responses using our PAOvCWD model.

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Viral outbreak dynamics and evolution in wildlife at the interface with humans

In this study, we used a multi-faceted approach to understand patterns of severe acute respiratory syndrome coronavirus 2 (SARS-CoV-2) transmission and persistence in a wild white-tailed deer ( Odocoileus virginianus ) population. Serology data indicated transmission of SARS-CoV-2 and persistence during the seven-month sampling period. Traditional disease modelling based on deer-to-deer transmission indicated relatively low prevalence with an R 0 of 1.9 and recovery period of 7 days; however, individual-based modelling informed by GPS tracked-movement data captured a potential transmission event. Phylogenetic analyses revealed a recurring pattern of divergent groups of deer-derived sequences with human-derived sequences falling close to each deer-derived cluster. Further, human-derived sequences were frequently sampled months prior to the deer-derived sequences, indicating repeated human to deer spillover. Using multiple types of data as well as both fine and broad scale analyses, we have characterized a pattern of localized outbreaks of SARS-CoV-2 within white-tailed deer populations that are likely recurring due to frequent spillover events. Our results suggest that while deer-to-deer transmission occurs over small spatiotemporal scales, SARS-CoV-2 persistence over longer periods and across larger regions is likely driven by repeated spillover from human populations.

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Invasive predatory fish occupies highest trophic position leading to expansion of isotopic niches in a riverine food web

Invasive species are drivers of ecological change with the potential to reshape the structure and function of terrestrial and aquatic ecosystems. The invasive flathead catfish ( Pylodictis olivaris ) is an opportunistic predator that has established a rapidly growing population in the Susquehanna River, Pennsylvania, USA, since they were first detected in 2002. Although the predatory effects of invasive catfishes on native fish communities have been documented, the effects of invasion on riverine food webs are poorly understood. This study quantified the effects of invasive flathead catfish on the trophic position (TP) and isotopic niche of the river's food web by comparing invaded and non-invaded sites. In addition to flathead catfish, the food web components examined included crayfish, minnows, and two ecologically and socioeconomically important fish species: the smallmouth bass ( Micropterus dolomieu ) and channel catfish ( Ictalurus punctatus ). We found that flathead catfish occupied the highest TP, with a posterior mean TP of 3.08 (95% credible interval = [2.71, 3.42]), exceeding that of the two resident fish predators, the smallmouth bass and channel catfish. The TP of the resident channel catfish, which occupies a similar ecological niche, declined after flathead catfish invasion. In fact, there was a 0.92 posterior probability that channel catfish TP was lower in invaded sites than at non-invaded sites. Using a Bayesian bivariate ellipses analysis, we found overwhelming evidence of isotopic niche expansion and overlap for all components of the food web in the presence of flathead catfish. These findings support the “trophic disruption hypothesis,” where an introduced species prompts resident species to change diets in an attempt to avoid competition and predation following invasion. Our results indicate that flathead catfish invasion is altering food web structure and energy flow in a large riverine ecosystem and contributes to the breadth of knowledge regarding how ecosystems may respond to the introduction of a large predatory fish species.

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Atypical winter coat coloration of snowshoe hares near the southern extent of their range

Many species have a variety of adaptations to winter weather, but these adaptations could become maladaptive if winter snowfall and temperatures are more variable. Snowshoe hares ( Lepus americanus ) molt from a brown summer coat to a white winter coat, but reductions in snow cover could result in phenotypic mismatch, which in turn could reduce survival. Hare populations near the southern extent of their range might be especially sensitive to phenotypic mismatch because of variable winter weather, but variation in winter coat coloration could allow for these populations to persist in inconsistent snow cover conditions. Using capture data ( n = 59 individual hares) spanning 8 years, we document the prevalence of three atypical winter coat color phenotypes (brown bodies, brown-ringed eyes, and brown ears) in a snowshoe hare population in Pennsylvania. The majority of hares in our study (84.7%) exhibited at least one of these atypical winter phenotypes, with a high probability of hares having brown-ringed eyes or brown ears, and four hares remaining brown during the winter. The presence and high prevalence of non-white winter phenotypes could be beneficial for hares in this population if winters are mild with low snow cover. If these phenotypes have a genetic basis, there may be evolutionary potential for hares to persist near the southern extent of their range, even in the face of changing winters.

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Landscape transcriptomic analysis detects thermal stress responses and potential adaptive variation in wild brook trout (Salvelinus fontinalis) during successive heatwaves

Extreme weather events, such as heatwaves, are becoming more frequent and intense as a result of climate change. Critically, such extreme weather events can be more important drivers of extirpation and selection than changes in annual or seasonal averages and they pose a particularly large threat to poikilothermic organisms. In this study, we evaluated the transcriptomic response of a coldwater adapted fish species, the eastern brook trout ( Salvelinus fontinalis ), to two successive heatwaves during July and August 2022. We sampled brook trout at eight time points from four streams ( N = 116 fish), sequenced mRNA from gill samples using TagSeq, and quantified expression levels of 32,670 unique transcripts. Multivariate analyses found that overall expression patterns in response to water temperature change were similar among streams. These analyses further detected groups of genes involved in immune response and oxygen carrier activity that were upregulated and downregulated respectively at higher water temperatures. We also detected 43 genes that were differentially expressed at different time points and followed the same expression pattern during the two heatwaves. Of these genes, 42 covaried with water temperature and most (27, 62.8 %) exhibited responses that varied by stream. Some of the differentially expressed genes, including heat shock proteins and cold-inducible RNA binding proteins, have been widely linked to temperature responses in experimental studies, whereas other genes we identified have functions that have not been well-studied in relationship to temperature or have unknown functions. This study shows the utility of landscape transcriptomic approaches to identify important biological processes governing wild organismal responses to short-term stressors. The results of this study can guide future investigations to identify phenotypic and genetic diversity that contribute to adaptive responses to heatwaves and improve predictions of how populations will respond to future climate change.

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Hydrologic investigations and a preliminary conceptual model of the groundwater system at North Penn Area 1 Superfund Site, Souderton, Montgomery County, Pennsylvania

The U.S. Geological Survey (USGS) conducted hydrogeologic investigations, reviewed existing data, and developed a preliminary conceptual model of the groundwater system as part of technical support of the U.S. Environmental Protection Agency (EPA) at the North Penn Area 1 Superfund Site (hereafter, the NP1 Site) located within the Borough of Souderton in Montgomery County, Pennsylvania. Field work and monitoring took place during 2012–18. The area is underlain by sedimentary formations that form a fractured-rock aquifer used for drinking water and industrial supply. The EPA placed the Site on the National Priorities List in 1989, identifying tetrachloroethylene (PCE) and trichloroethylene (TCE) as contaminants of concern. During 2012–18, the USGS conducted field activities that included drilling an 82-foot (ft)-deep monitoring well (MG 2220) in 2016, reconstructing a 208-ft-deep former industrial production well (MG 668 [Granite Knitting Mill]), and collecting borehole geophysical and video logs and water levels from those and five additional wells, which ranged in depth from about 50 to 200 ft below land surface. Continuous water levels were collected during 2014–17, and a synoptic set of water levels were measured in April 2018 in the seven wells. The borehole geophysical logs (caliper, acoustic televiewer, natural gamma, single-point resistance, vertical flow, and fluid temperature and resistivity) and borehole video logs in the seven wells were evaluated to assess potential for lithologic correlation and to identify and describe water-bearing features, which included both low- and high-angle fractures and other openings oriented along dipping bedding planes, joints, or possible faults. Borehole geophysical logs collected by USGS in 1992 in a 300-ft-deep former production well near the Site were also evaluated. Few to no distinctive features were identified on geophysical logs (natural gamma and single-point resistance) that could be used for correlation, thus limiting this approach to determining local geologic structure. Extensive fracturing in the upper 62 ft of monitoring well MG 2220 indicates that the well was likely drilled through a zone of faulting, and other evidence of faulting is present in the area near the Site. Assessment of continuous water levels showed hydraulic connections among some wells as indicated by rising or falling water levels in response to changes in pumping rates at nearby wells. A map of water levels measured in April 2018 indicates potential for groundwater flow generally toward the stream to the south and southwest of the Site, but the limited water-level data are insufficient to describe vertical groundwater gradients or lateral gradients in any detail. Review of 1999–2022 volatile organic compound (VOC) monitoring data collected by the Pennsylvania Department of Environmental Protection for five monitoring wells indicates that the highest groundwater concentrations of PCE and TCE were found in samples from extraction well MG 2201 (S-1) downgradient from, and nearest to, the previously identified Site contaminant source area, and these concentrations fluctuated through time. PCE concentrations were higher than TCE concentrations in samples from all five monitoring wells and were much higher than TCE concentrations in samples from extraction well MG 2201 (S-1). Temporally variable recharge is a possible factor affecting observed fluctuations in PCE concentrations in groundwater samples from well extraction MG 2201 (S-1), as indicated by a general inverse relation between PCE concentrations and water levels in a nearby long-term observation well. The PCE concentration of 1,830 micrograms per liter (μg/L) in a May 2018 water sample from monitoring well MG 2220 was more than four times the PCE concentration of 444 μg/L in a December 2017 sample from the nearby extraction well MG 2201 (S-1), which is open to fewer fractures. Low concentrations of VOCs were measured in surface water at two stream sites downgradient from wells with the highest groundwater VOC concentrations at the Site, indicating that discharge of contaminated groundwater to the stream is likely. Development of a conceptual model of the groundwater system was constrained by limited data. In areas with no pumping, groundwater-flow directions generally are thought to be controlled by topography and geologic structure (bedding orientation) and likely to the south and southwest of the Site, with local flow directions affected by orientations of fractures, joints, and local faults. Additional investigations that could help improve the conceptual model of the groundwater system and help delineate the extent of groundwater contamination and its transport are discussed.

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Predicted potential for aquatic exposure effects of per- and polyfluorinated alkyl substances (PFAS) in Pennsylvania’s statewide network of streams

Per- and polyfluoroalkyl substances (PFAS) are contaminants that can lead to adverse health effects in aquatic organisms, including reproductive toxicity and developmental abnormalities. To assess the ecological health risk of PFAS in Pennsylvania stream surface water, we conducted a comprehensive analysis that included both measured and predicted estimates. The potential combined exposure effects of 14 individual PFAS to aquatic biota were estimated using the sum of exposure-activity ratios ( Σ EARs) in 280 streams. Additionally, machine learning techniques were utilized to predict potential PFAS exposure effects in unmonitored stream reaches, considering factors such as land use, climate, and geology. Leveraging a tailored convolutional neural network (CNN), a validation accuracy of 78% was achieved, directly outperforming traditional methods that were also used, such as logistic regression and gradient boosting (accuracies of ~65%). Feature importance analysis highlighted key variables that contributed to the CNN’s predictive power. The most influential features highlighted the complex interplay of anthropogenic and environmental factors contributing to PFAS contamination in surface waters. Industrial and urban land cover, rainfall intensity, underlying geology, agricultural factors, and their interactions emerged as key determinants. These findings may help to inform biotic sampling strategies, water quality monitoring efforts, and policy decisions aimed to mitigate the ecological impacts of PFAS in surface waters.

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Factors influencing the prevalence of hyperpigmented melanistic lesions in smallmouth bass Micropterus dolomieu in the Susquehanna River Basin, Pennsylvania

Hyperpigmented melanistic lesions (HPMLs) are a visual anomaly documented on the skin of smallmouth bass Micropterus dolomieu in the Susquehanna River Basin, Pennsylvania and in numerous other geographical locations. Currently, there is a lack of information on environmental and fish characteristics that may influence the prevalence of HPMLs associated with a recently described Adomavirus . The goal of this study was to understand potential drivers associated with HPMLs in socioeconomically and ecologically important riverine smallmouth bass populations. A total of 16,220 smallmouth bass were collected and examined for HPMLs between 2012 and 2022 in the Susquehanna River Basin. Overall, HPMLs were documented on 2.9% of fish collected. The interaction between temperature and fish size suggested differing relationships between shorter and longer fish with respect to temperature. Predicted probability of HPML prevalence ranged from 1.1% (95% CI = 0.3, 3.2) at 4°C to 0.01% (CI = 0.00, 0.04) at 26°C for an age-0 (125 mm) fish. In contrast, predicted probability of HPML prevalence ranged from 10.5% (95% CI = 5.8, 18.9) at 4°C to 0.8% (CI = 0.4, 1.5) at 26°C for an adult (322 mm) fish. Overall, HPMLs were more common in longer fish during cooler temperature periods which also corresponds to key life history periods for smallmouth bass (e.g., pre-spawn and overwintering) and could represent different exposure histories for juvenile and adult fish.

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Temporal analysis of water chemistry and smallmouth bass (Micropterus dolomieu) health at two sites with divergent land use in the Susquehanna River watershed, Pennsylvania, USA

Monitoring wild fish health and exposure effects in impacted rivers and streams with differing land use has become a valuable research tool. Smallmouth bass ( Micropterus dolomieu ) are a sensitive, indicator species that exhibit signs of immunosuppression and endocrine disruption in response to water quality changes and contaminant exposure. To determine the impact of agriculture and development on smallmouth bass health, two sites (a developed/agriculture site and a forested site) in the Susquehanna River watershed, Pennsylvania were selected where bass and water chemistry were sampled from 2015 to 2019. Smallmouth bass were sampled for histopathology to assess parasite and macrophage aggregate density in the liver and spleen, condition factor (Ktl), hepatic gene transcript abundance, hepatosomatic index (HSI), and a health assessment index (HAI). Land use at the developed/agriculture site included greater pesticide application rates and phytoestrogen crop cover and more detections and higher concentrations of pesticides, wastewater-associated contaminants, hormones, phytoestrogens, and mycotoxins than at the forested site. Additionally, at the developed/agriculture site, indicators of molecular changes, including oxidative stress, immune/inflammation, and lipid metabolism-related hepatic gene transcripts, were associated with more contaminants and land use variables. At both sites, there were multiple associations of contaminants with liver and/or spleen macrophage aggregate density, indicating that changes at the molecular level seemed to be a better indicator of exposures unique to each site. The findings illustrate the importance of timing for land management practices, the complex mixtures aquatic animals are exposed to, and the temporal changes in contaminant concentration. Agricultural practices that affect hepatic gene transcripts associated with immune function and disease resistance were demonstrated which could negatively affect smallmouth bass populations.

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Feeding habits and ecological implications of the invasive Flathead Catfish in the Susquehanna River basin, Pennsylvania

Objective Flathead Catfish Pylodictis olivaris are a widespread aquatic invasive species within the United States and a recent invader in the Susquehanna River basin, Pennsylvania. Flathead Catfish are piscivores known to consume native and recreationally important fish species. In the mid-Atlantic United States, it is unknown how this invader is impacting food webs and which species may be at greatest predation risk. To address this knowledge gap, we DNA barcoded stomach contents collected from Flathead Catfish to identify prey species and elucidate potential predatory impacts of Flathead Catfish in the Susquehanna River. Methods We used a Bayesian hierarchical multivariate probit model to investigate if the probability of prey species occurrence in the diets of Flathead Catfish varied seasonally or was a function of Flathead Catfish length. Result A total of 576 Flathead Catfish were collected during 2020–2021, with 241 individuals having recoverable stomach contents. In all, we identified 47 different prey species. The most common prey species were rusty crayfish Faxonius rusticus , Margined Madtom Noturus insignis , and shiners Notropis spp . While frequency of occurrence of prey species differed across Flathead Catfish length classes (<300 mm, 301–600 mm, 601–900 mm TL), rusty crayfish were commonly found (33.7–44.0% of diets) in stomachs of all size-classes. Conclusion We found that Flathead Catfish length and seasonality did influence occurrence probability differentially for several prey species. For example, Channel Catfish Ictalurus punctatus were more likely to appear in shorter Flathead Catfish while Smallmouth Bass Micropterus dolomieu appeared in larger Flathead Catfish. We demonstrate significant variation in Flathead Catfish predation, increasing our understanding of predator–prey dynamics, which is necessary to better manage and identify future impacts to aquatic communities in the Susquehanna River basin.

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Delayed positive responses of snowshoe hares to prescribed burning in a fire-adapted ecosystem

Wildlife populations near the periphery of a species’ range are vulnerable to changes in habitat conditions and climate. However, habitat management and maintenance can help with the persistence of these susceptible populations. Snowshoe hare ( Lepus americanus ) populations near the southern extent of their range are at risk of extirpation because of changing winter conditions, coupled with reduced early-successional habitat. Prescribed fire has been used to restore and maintain early-successional habitat in the southern range of snowshoe hares, but previous research suggests that burned areas might initially be unsuitable for hares. Therefore, more information is needed on the time required for hares to start using an area post-burning, and the long-term effects of prescribed fire on hare survival. We studied snowshoe hare habitat use and survival in northeastern Pennsylvania, where prescribed fire has been used for scrub oak barrens restoration. We used GPS locations from 71 hares and used resource selection functions to analyze hare selection for burned habitats of varying ages (0–12 years post-burn) and known-fate survival models to evaluate the effects of burning on survival. Hares started using burned areas ≥7 years post-burning but avoided areas burned 0–6 years prior. In addition, hare survival was positively associated with the amount of old burn habitat (≥7 years post-burn) used by an individual. Our results indicate that prescribed burning can be beneficial for hares, but that a time lag of ≥7 years is necessary for positive responses to occur. Planning burns within a mosaic of unburned areas could allow hares to persist during the 0–6 years post-burn when areas are not suitable for hares, which in turn could benefit the persistence of southern snowshoe hare populations. Collectively, our results highlight the importance of long-term demographic monitoring to understand wildlife population responses to management actions.

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