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R. J. Greenwood

Publications and source records attributed to R. J. Greenwood.

At least 19 recordsLinked to original sources
USGS2004

Analysis of predator movement in prairie landscapes with contrasting grassland composition

Mammalian predation influences waterfowl breeding success in the U.S. northern Great Plains, yet little is known about the influence of the landscape on the ability of predators to find waterfowl nests. We used radiotelemetry to record nightly movements of red foxes ( Vulpes vulpes ) and striped skunks ( Mephitis mephitis ) in two 41.4-km 2 study areas in North Dakota. Study areas contained either 15–20% grassland (low grassland composition) or 45–55% grassland (high grassland composition). Grasslands included planted cover, pastureland, and hayland. We predicted that the type and composition of cover types in the landscape would influence both predator movement across the landscape (as measured by the fractal dimension and displacement ratio) as well as localized movement (as measured by the rate of movement and turning angle between locations) within patches of different cover types. Red fox movements were straighter (lower fractal dimensions and higher displacements) across landscapes with a low grassland composition, indicating directed movement between the more isolated patches of planted cover. Striped skunk movements did not differ between landscape types, illustrating their movement along wetland edges, which had similar compositions in both landscape types. The high variability in turning angles by red fox in planted cover and pastureland in both landscape types is consistent with restricted-area foraging. The high rate of movement by red foxes in planted cover and by striped skunks in wetland edges suggests that spatial memory may influence movement patterns. Understanding the behavior of predators in fragmented prairie landscape is essential for managing breeding habitat for grassland birds and for predicting the spatial and temporal dynamics of predators and their prey.

North Dakota↗
USGS2000

Relationships of habitat patch size to predator community and survival of duck nests

We studied duck nest success and predator community composition in relation to size of discrete patches of nesting cover in the Prairie Pothole Region (PPR) of the United States in 1993-95. We focused on nests in uplands that were seeded to perennial grasses and forbs and enrolled in the Conservation Reserve Program (CRP) in Minnesota, North Dakota, and South Dakota. We estimated daily survival rates (DSRs) of upland duck nests and indices of activity for red foxes (Vulpes vulpes), coyotes (Canis latrans), American badgers (Taxidea taxus), striped skunks (Mephitis mephitis), and Franklin's ground squirrels (Spermophilus franklinii), and related these variables to habitat patch size. The effect of patch size (small vs. large) on estimated annual mean DSR was dependent on date of nest initiation (early vs. late) and year. Examination of within-year comparisons for early and late nests suggested that DSR was generally greater in larger habitat patches. Activity indices for the 5 mammalian nest predators were influenced differently by year, location, and patch size. Activity indices of the red fox were greatest in small patches. Coyote indices were the most inconsistent, demonstrating a year X location X patch size interaction. Activity indices of the striped skunk and American badger varied only among years. Franklin's ground squirrel indices were affected by study area location, with higher indices in the southeast than the northwest. Red fox activity was weakly correlated with that of the striped skunk and coyote. Although a positive relationship between habitat patch size and nest success probably exists, we believe the experiment to fully test this hypothesis will continue to be elusive.

Journal of Wildlife Management↗
USGS1999

Foods and foraging of prairie striped skunks during the avian nesting season

Food habits of prairie skunks are not well understood, yet such knowledge might provide insight into factors influencing nest depredation. We studied food habits of radiocollared adult striped skunks (Mephitis mephitis) during 1976-78 in North Dakota, where skunks are regarded as important predators of ground-nesting birds. Plant foods, primarily grain and sunflower seeds, occurred in a larger percentage of scats in spring (15 Apr 31 May) than summer (1 Jun 15 July, P=0.04), but overall, plant foods were a minor part of skunk diets. Animal foods, primarily birds (including eggs), small rodents, and insects occurred annually in a large percentage of scats of all skunks. These foods were acquired nearly exclusively in grasslands. Percentage of scats containing animal foods was similar, irrespective of sex, season, or year (P>0.45). In spring, vertebrates occurred in a smaller percentage of scats of females than males (P<0.01). Among females, vertebrates also occurred in a smaller percentage of scats in spring than summer (P<0.01). Among all skunks, birds occurred in a smaller percentage of scats in spring than in summer (P<0.01). Both birds (P<0.02) and mammals (P<0.01) occurred in a smaller proportion of scats when wetland conditions were poor. Percentage of scats containing insects, the primary invertebrate food of skunks, did not vary by sex, season, or year (P>0.15). Insects were mostly adult and larval Coleoptera, larval Lepidoptera, and adult and nymph Orthoptera.

Wildlife Society Bulletin↗
USGS1998

Effects of food supplementation on depredation of duck nests in prairies by striped skunks

We examined provision of supplemental food as a method for reducing depredation of upland-duck nests, especially by striped skunks (Mephitis mephitis). Predators negatively influence duck recruitment in grassland ecosystems. Managers are in search of methods, particularly nonlethal methods, for reducing nest depredation. We conducted this study on 24 areas managed for wildlife production in the Prairie Pothole Region of central North Dakota during 1993-1994. We provided a mix of fish offal and sunflower seeds on 12 areas; no food was provided on the remaining 12 control areas. Although we observed a tendency during both years for higher nest success rates on provisioned areas ( = 46%, 1993; 36%, 1994) than on control areas ( = 27%, 1993; 31%, 1994), mean nest success rates (Mayfield 1961) overall did not differ significantly between food-provisioned areas ( = 41%) and control areas ( = 29%). Striped skunk depredation rate was lower on food-provisioned areas (11%) than on control areas (24%), suggesting that skunks reduced their consumption of eggs when provided with a food supplement. In 1994, habitat conditions were optimal, and ducks nested persistently into the summer when nest success rates of food-provisioned areas and control areas differed by only 5 percentage points. That year American badgers (Taxidea taxus) and Franklin's ground squirrels (Spermophilus franklinii) apparently compensated for reduced depredation by skunks. Thus, although skunks and other mammalian predators seem to have responded positively to food provisioning, nest depredations overall did not change. Provision of supplemental food apparently has limited value for managing depredation of upland duck nests in the Prairie Pothole Region where predator communities are complex.

Wildlife Society Bulletin↗
USGS1993

Distribution and abundance of predators that affect duck production--prairie pothole region

During 1983-88, the relative abundance of 18 species and species-groups of mammalian and avian predators affecting duck production in the prairie pothole region was determined in 33 widely scattered study areas ranging in size from 23-26 km2. Accounts of each studied species and species-group include habitat and history, population structure and reported densities, and information on distribution and abundance from the present study. Index values of undetected, scarce, uncommon, common, or numerous were used to rate abundance of nearly all species in each study area. Principal survey methods were livetrapping of striped skunks (Mephitis mephitis) and Franklin's ground squirrels (Spermophilus franklinii), systematic searches for carnivore tracks in quarter sections (0.65 km2), daily records of sightings of individual predator species, and systematic searches for occupied nests of tree-nesting avian predators. Abundances of predators in individual areas were studied 1-3 years.The distribution and abundance of predator species throughout the prairie pothole region have undergone continual change since settlement of the region by Europeans in the late 1800's. Predator populations in areas we studied differed markedly from those of pristine times. The changes occurred from habitat alterations, human-inflicted mortality of predators, and interspecific relations among predator species. Indices from surveys of tracks revealed a decline in the abundance of red foxes (Vulpes vulpes) and an albeit less consistent decline in the abundance of raccoons (Procyon lotor) with an increase in the abundance of coyotes (Canis latrans). Records of locations of occupied nests revealed great horned owls (Bubo virginianus) and red-tailed hawks (Buteo jamaicensis) tended to nest 0.5 km apart, and American crows (Corvus brachyrhynchos) tended to avoid nesting 0.5 km of nests of red-tailed hawks. Excluding large gulls, for which no measurements of abundance were obtained, the number of predator species averaged 12.2 (SD = 1.60) per study area; common or numerous predator species averaged 6.0 (SD = 1.54) per study area (minimal because the abundance of weasels [Mustela erminea; M. frenata] in all areas and of minks [Mustela vison] and raptors in some areas was not rated). Major changes in relative abundance of individual predator species studied >1 year were few. Predator species most restricted to the aspen parkland were the Franklin's ground squirrel, black-billed magpie (Pica pica), American crow (Corvus brachyrlus), and red-tailed hawk; species most restricted to the prairie were the badger (Taxidea taxus), Swainson's hawk (Buteo swainsoni), and ferruginous hawk (B. regalis). The coyote, black-billed magpie, and American crow were most numerous in Canada, whereas the red fox, raccoon, mink, ferruginous hawk, and great horned owl were most numerous in the United States. The number of common or numerous egg-eating predator species (excludes large gulls and weasels, which were not rated) averaged 4.6 (SD = 0.90) per study area. The average numbers of common or numerous egg-eating species per study area did not differ among provinces and states, but birds gradually replaced mammals from southeast to northwest across the region. Investigators are urged to assess composition of predator populations and relative abundance of predator species for evaluations of waterfowl recruitment.

Resource Publication↗
USGS1987

Responses of three prairie ground squirrel species, Spermophilus franklinii, S. richardsonii, and S. tridecemlineatus, to duck eggs

Franklin's Ground Squirrels (Spermophilus franklinii) preyed on eggs in 29 of 30 tests in which they were exposed to two Mallard (Anas platyrhynchos) or two Blue-winged Teal (A. discors) eggs and their discovery of eggs was confirmed. No Richardson's (S. richardsonii) or Thirteen-lined (S. tridecemlineatus) ground squirrels preyed on eggs in 104 and 54 similar tests, respectively. Franklin's Ground Squirrels exposed to clutches of six duck eggs placed in simulated nests preyed on all such clutches. Predation on simulated nests began soon after eggs were discovered, but predation of all eggs often took more than one day to complete.

Canadian Field-Naturalist↗
USGS1982

Recurrence, mortality, and dispersal of prairie striped skunks, Mephitis mephitis, and implications to rabies epizootiology

Detailed study of radio-equipped individuals of the Striped Skunk (Mephitis mephitis) in a North Dakota population provided insight into possible mechanisms for spread of rabies during spring and summer. Annual recurrence rates of 138 skunks marked on a study area averaged 11% for adult males, 43% for adult females and 9% for kits. Population changes were from mortality (including rabies) and dispersal. Five instances of adult dispersal (four by males) were recorded; maximum straight-line distance was 119 km. Some males initiated dispersal in spring. Communal denning by adults occurred rarely after whelping began but resulted in intraspecific conflict. Evidence of intraspecific and interspecific strife leading to kit mortality and some adult mortality was found at dens of 9 of 40 litters studied.

Canadian Field-Naturalist↗
USGS1981

Observations on black-crowned night heron breeding success in a North Dakota marsh

Storms and mammalian predation severely reduced production in a Black-crowned Night Heron (Nycticorax nycticorax) colony in two years and drought prevented nesting in two other years from 1976 to 1980. Maximum estimated production was 2.2 young per pair in 1978, the only year production was sufficient to maintain the population.

Canadian Field-Naturalist↗
USGS1974

Ice on waterfowl markers

Wild and captive giant Canada geese (Branta canadensis maxima) and captive mallards (Anas platyrhynchos) accumulated ice on neck collars and/or nasal saddles during winter storm periods in 1971 and 1972. Weather conditions associated with icing were documented, and characteristics of icing are discussed. Severe marker icing occurred during subfreezing weather when the windchill reached approximately -37 deg.C. Birds appeared able to de-ice nasal saddles in most instances.

Wildlife Society Bulletin↗
USGS1973

Influence of radio packs on captive mallards and blue-winged teal

Back-mounted radio packs of three weights were attached to 30 captive mallards (Anas platyrhynchos) and 30 blue-winged teal (A. discors), and physical condition of the birds was evaluated for 12 weeks. All groups with radio packs lost more weight than did control groups. Feather wear often followed by skin irritation occurred in the axillary region of the wings on 90 percent of the treated birds within 1 week, and on the backs of 55 percent of the treated birds within 4 weeks after the radio packs were attached. The treated birds appeared preoccupied with the radio packs and exhibited a partial aversion to swimming.

Journal of Wildlife Management↗
USGS1972

Effect of certain anesthetic agents on mallard ducks

Four anesthetic agents used in human or veterinary medicine and 3 experimental anesthetic preparations were evaluated for effectiveness in inducing narcosis when administered orally to game-farm mallard ducks (Anas platyrhynchos).Tribromoethanol was the only compound to satisfy criteria of initial tests. Mean duration of the induction, immobilization, and recovery periods was 2.4 minutes, 8.7 minutes, and 1.3 hours, respectively, at the median effective dosage for immobilization (ED50; 100 mg./kg. of body weight). The median lethal dosage (LD50) was 400 mg./kg. of body weight.Tribromoethanol was also tested on mallards during the reproductive season. Effects on the hatchability of eggs or the survival of young were not detected.

Journal of the American Veterinary Medical Associa↗