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N.B. Kotliar

Publications and source records attributed to N.B. Kotliar.

9 recordsLinked to original sources
USGS2008

Application of the new keystone-species concept to prairie dogs: How well does it work?

It has been suggested that the keystone-species concept should be dropped from ecology and conservation, primarily because the concept is poorly defined. This prompted Power et al. (1996) to refine the definition: keystone species have large effects on community structure or ecosystem function (i.e., high overall importance), and this effect should be large relative to abundance (i.e., high community importance). Using prairie dogs ( Cynomys spp.) as an example, I review operational and conceptual difficulties encountered in applying this definition. As applied to prairie dogs, the implicit assumption that overall importance is a linear function of abundance is invalid. In addition, community importance is sensitive to abundance levels, the definition of community, and sampling scale. These problems arise largely from the equation for community importance, as used in conjunction with removal experiments at single abundance levels. I suggest that we shift from the current emphasis on the dualism between keystone and nonkeystone species and instead examine how overall and community importance vary (1) with abundance, (2) across spatial and temporal scales, and (3) under diverse ecological conditions. In addition, I propose that a third criterion be incorporated into the definition: keystone species perform roles not performed by other species or processes. Examination of how these factors vary among populations of keystone species should help identify the factors contributing to, or limiting, keystone-level functions, thereby increasing the usefulness of the keystone-species concept in ecology and conservation. Although the quantitative framework of Power et al. falls short of being fully operational, my conceptual guidelines may improve the usefulness of the keystone-species concept. Careful attention to the factors that limit keystone function will help avoid misplaced emphasis on keystone species at the expense of other species.

Conservation Biology↗
USGS2008

Opposing environmental gradients govern vegetation zonation in an intermountain playa

Vegetation zonation was investigated at an intermountain playa wetland (Mishak Lakes) in the San Luis Valley (SLV) of southern Colorado. Plant composition and abiotic conditions were quantified in six vegetation zones. Reciprocal transplants were performed to test the importance of abiotic factors in governing zonation. Abiotic conditions differed among several vegetation zones. Prolonged inundation led to anaerobic soils in the Eleocharis palustris and the submerged aquatics zones, on the low end of the site's 1.25 m elevation gradient. On the high end of the gradient, soil salinity and sodicity (a measure of exchangeable sodium) were high in the Distichlis spicata zone (electrical conductivity, EC = 5.3 dS/m, sodium absorption ratio, SAR = 44.0) and extreme in the Sarcobatus vermiculatus zone (EC = 21 dS/m, SAR = 274). Transplanted species produced maximum biomass in the zone where they originated, not in any other higher or lower vegetation zone. The greatest overall transplant effect occurred for E. palustris, which experienced a ??? 77% decline in productivity when transplanted to other zones. This study provides evidence that physical factors are a major determinant of vegetation zone composition and distribution across the entire elevation gradient at Mishak Lakes. Patterns at Mishak Lakes arise from counter-directional stress gradients: a gradient from anaerobic to well-oxygenated from basin bottom to upland and a gradient from extremely high salinity to low salinity in the opposing direction. Because abiotic conditions dominate vegetation zonation, restoration of the altered hydrologic regime of this wetland to a natural hydrologic regime may be sufficient to re-establish many of the natural biodiversity functions provided by these wetlands. ?? 2008 The Society of Wetland Scientists.

Wetlands↗
USGS2008

Simulated natural hydrologic regime of an intermountain playa conservation site

An intermountain playa wetland preserve in Colorado's San Luis Valley was studied to assess how its current hydrologic function compares to its natural hydrologic regime. Current hydrologic conditions were quantified, and on-site effects of off-site water use were assessed. A water-budget model was developed to simulate an unaltered (i.e., natural) hydrologic regime, and simulated natural conditions were compared to observed conditions. From 1998-2002, observed stream inflows accounted for ??? 80% of total annual water inputs. No ground water discharged to the wetland. Evapotranspiration (ET) accounted for ??? 69% of total annual water loss. Simulated natural conditions differed substantially from current altered conditions with respect to depth, variability, and frequency of flooding. During 1998-2002, observed monthly mean surface-water depth was 65% lower than under simulated natural conditions. Observed monthly variability in water depth range from 129% greater (May) to 100% less (September and October) than simulated. As observed, the wetland dried completely (i.e., was ephemeral) in all years; as simulated, the wetland was ephemeral in two of five years. For the period 1915-2002, the simulated wetland was inundated continuously for as long as 16 years and nine months. The large differences in observed and simulated surface-water dynamics resulted from differences between altered and simulated unaltered stream inflows. The maximum and minimum annual total stream inflows observed from 1998-2005 were 3.1 ?? 106 m3 and 0 m3, respectively, versus 15.5 ?? 106 m3 and 3.2 ?? 106 m3 under simulated natural conditions from 1915-2002. The maximum simulated inflow was 484% greater than observed. These data indicate that the current hydrologic regime of this intermountain playa differs significantly from its natural hydrologic regime, which has important implications for planning and assessing conservation success. ?? 2008, The Society of Wetland Scientists.

Wetlands↗
USGS2007

Avifaunal responses to fire in southwestern montane forests along a burn severity gradient

The effects of burn severity on avian communities are poorly understood, yet this information is crucial to fire management programs. To quantify avian response patterns along a burn severity gradient, we sampled 49 random plots (2001-2002) at the 17 351-ha Cerro Grande Fire (2000) in New Mexico, USA. Additionally, pre-fire avian surveys (1986-1988, 1990) created a unique opportunity to quantify avifaunal changes in 13 pre-fire transects (resampled in 2002) and to compare two designs for analyzing the effects of unplanned disturbances: after-only analysis and before-after comparisons. Distance analysis was used to calculate densities. We analyzed after-only densities for 21 species using gradient analysis, which detected a broad range of responses to increasing burn severity: (I) large significant declines, (II) weak, but significant declines, (III) no significant density changes, (IV) peak densities in low- or moderate-severity patches, (V) weak, but significant increases, and (VI) large significant increases. Overall, 71% of the species included in the after-only gradient analysis exhibited either positive or neutral density responses to fire effects across all or portions of the severity gradient (responses III-VI). We used pre/post pairs analysis to quantify density changes for 15 species using before-after comparisons; spatiotemporal variation in densities was large and confounded fire effects for most species. Only four species demonstrated significant effects of burn severity, and their densities were all higher in burned compared to unburned forests. Pre- and post-fire community similarity was high except in high-severity areas. Species richness was similar pre- and post-fire across all burn severities. Thus, ecosystem restoration programs based on the assumption that recent severe fires in Southwestern ponderosa pine forests have overriding negative ecological effects are not supported by our study of post-fire avian communities. This study illustrates the importance of quantifying burn severity and controlling confounding sources of spatiotemporal variation in studies of fire effects. After-only gradient analysis can be an efficient tool for quantifying fire effects. This analysis can also augment historical data sets that have small samples sizes coupled with high non-process variation, which limits the power of before-after comparisons. ?? 2007 by the Ecological Society of America.

Ecological Applications↗
USGS2006

Green-tailed Towhee response to prescribed fire in montane shrubland

Fire alters the structure and composition of shrublands and affects habitat quality for the associated avifauna. Because shrubland ecosystems have been greatly reduced from their original extent in western North America and fire is increasingly being used to manage these landscapes, a better understanding of how fire affects the associated vegetation and wildlife is imperative. We evaluated the response of Green-tailed Towhees (Pipilo chlorurus) to prescribed fire in the montane shrublands of Rocky Mountain National Park, Colorado during 2002 and 2003. Three to five years following prescribed burning, Green-tailed Towhee density and shrub cover were generally higher in unburned areas. Nests (n = 179) were located in unburned vegetation; within burned sites, all nests were in remnant patches. Green-tailed Towhee nest survival was 57% (95% CI = 49%-65%) across the two years of the study. More than half of the nests were in common juniper (Juniperus communis) shrubs, and nest survival was higher for nests in junipers than those in other shrub species. Daily nest survival rates were lower at the site with the highest density of towhees and declined over the breeding season. With regard to shrub cover, opposite trends were observed for nest-site selection and nest survival: nest plots had greater shrub cover than non-nest plots, but nest survival decreased with increasing shrub cover. Because shrub cover affects towhee density and nest survival in conflicting ways, fire management at Rocky Mountain National Park alters both habitat availability and suitability for Green-tailed Towhees. ?? The Cooper Ornithological Society 2006.

Condor↗
USGS2005

Variation in fire regimes of the rocky mountains: Implications for avian communities and fire management

Information about avian responses to fire in the U.S. Rocky Mountains is based solely on studies of crown fires. However, fire management in this region is based primarily on studies of low-elevation ponderosa pine (Pinus ponderosa) forests maintained largely by frequent understory fires. In contrast to both of these trends, most Rocky Mountain forests are subject to mixed severity fire regimes. As a result, our knowledge of bird responses to fire in the region is incomplete and skewed toward ponderosa pine forests. Research in recent large wildfires across the Rocky Mountains indicates that large burns support diverse avifauna. In the absence of controlled studies of bird responses to fire, we compared reproductive success for six cavity-nesting species using results from studies in burned and unburned habitats. Birds in ponderosa pine forests burned by stand-replacement fire tended to have higher nest success than individuals of the same species in unburned habitats, but unburned areas are needed to serve species dependent upon live woody vegetation, especially foliage gleaners. Over the last century, fire suppression, livestock grazing, and logging altered the structure and composition of many low-elevation forests, leading to larger and more severe burns. In higher elevation forests, changes have been less marked. Traditional low-severity prescribed fire is not likely to replicate historical conditions in these mixed or high-severity fire regimes, which include many mixed coniferous forests and all lodgepole pine (Pinus contorta) and spruce-fir (Picea-Abies) forests. We suggest four research priorities: (1) the effects of fire severity and patch size on species' responses to fire, (2) the possibility that postfire forests are ephemeral sources for some bird species, (3) the effect of salvage logging prescriptions on bird communities, and (4) experiments that illustrate bird responses to prescribed fire and other forest restoration methods. This research is urgent if we are to develop fire management strategies that reduce fire risk and maintain habitat for avifauna and other wildlife of the Rocky Mountains.

Conference Paper↗
USGS2002

Effects of fire and post-fire salvage logging on avian communities in conifer-dominated forests of the western United States

Historically, fire was one of the most widespread natural disturbances in the western United States. More recently, however, significant anthropogenic activities, especially fire suppression and silvicultural practices, have altered fire regimes; as a result, landscapes and associated communities have changed as well. Herein, we review current knowledge of how fire and post-fire salvaging practices affect avian communities in conifer-dominated forests of the western United States. Specifically, we contrast avian communities in (1) burned vs. unburned forest, and (2) unsalvaged vs. salvage-logged burns. We also examine how variation in burn characteristics (e.g., severity, age, size) and salvage logging can alter avian communities in burns. Of the 41 avian species observed in three or more studies comparing early post-fire and adjacent unburned forests, 22% are consistently more abundant in burned forests, 34% are usually more abundant in unburned forests, and 44% are equally abundant in burned and unburned forests or have varied responses. In general, woodpeckers and aerial foragers are more abundant in burned forest, whereas most foliage-gleaning species are more abundant in unburned forests. Bird species that are frequently observed in stand-replacement burns are less common in understory burns; similarly, species commonly observed in unburned forests often decrease in abundance with increasing burn severity. Granivores and species common in open-canopy forests exhibit less consistency among studies. For all species, responses to fire may be influenced by a number of factors including burn severity, fire size and shape, proximity to unburned forests, pre- and post-fire cover types, and time since fire. In addition, post-fire management can alter species' responses to burns. Most cavity-nesting species do not use severely salvaged burns, whereas some cavity-nesters persist in partially salvaged burns. Early post-fire specialists, in particular, appear to prefer unsalvaged burns. We discuss several alternatives to severe salvage-logging that will help provide habitat for cavity nesters. We provide an overview of critical research questions and design considerations crucial for evaluating the effects of prescribed fire and other anthropogenic disturbances, such as forest fragmentation. Management of native avifaunas may be most successful if natural disturbance regimes, including fire, are permitted to occur when possible. Natural fires could be augmented with practices, such as prescribed fire (including high-severity fire), that mimic inherent disturbance regimes.

Conference Paper↗
USGS2002

Effects of fire and post-fire salvage logging on avian communities in conifer-dominated forests of the western United States

Historically, fire was one of the most widespread natural disturbances in the western United States. More recently, however, significant anthropogenic activities, especially fire suppression and silvicultural practices, have altered fire regimes; as a result, landscapes and associated communities have changed as well. Herein, we review current knowledge of how fire and postfire salvaging practices affect avian communities in conifer-dominated forests of the western United States. Specifically, we contrast avian communities in (1) burned vs. unburned forest, and (2) unsalvaged vs. salvage-logged burns. We also examine how variation in burn characteristics (e.g., severity, age, size) and salvage logging can alter avian communities in burns. Of the 41 avian species observed in three or more studies comparing early postfire and adjacent unburned forests, 22% are consistently more abundant in burned forests, 34% are usually more abundant in unburned forests, and 44% are equally abundant in burned and unburned forests or have varied responses. In general, woodpeckers and aerial foragers are more abundant in burned forest, whereas most foliage-gleaning species are more abundant in unburned forests. Bird species that are frequently observed in stand-replacement burns are less common in understory burns; similarly, species commonly observed in unburned forests often decrease in abundance with increasing burn severity. Granivores and species common in open-canopy forests exhibit less consistency among studies. For all species, responses to tire may be influenced by a number of factors including burn severity, fire size and shape, proximity to unburned forests, pre-and post-fire cover types, and time since fire. In addition, postfire management can alter species’ responses to burns. Most cavity-nesting species do not use severely salvaged burns, whereas some cavity-nesters persist in partially salvaged burns. Early post fire specialists, in particular, appear to prefer unsalvaged burns. We discuss several alternatives to severe salvage-logging that will help provide habitat for cavity nesters. We provide an overview of critical research questions and design considerations crucial for evaluating the effects of prescribed fire and other anthropogenic disturbances, such as forest fragmentation. Management of native avifaunas may be most successful if natural disturbance regimes, including fire, are permitted to occur when possible. Natural fires could be augmented with practices, such as prescribed fire (including high-severity fire), that mimic inherent disturbance regimes.

Studies in Avian Biology↗