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Molly Van Appledorn

Publications and source records attributed to Molly Van Appledorn.

At least 19 recordsLinked to original sources

Patterns of floodplain forest mortality and recruitment along the Upper Mississippi and Illinois Rivers: Associations with forest fragmentation and flood inundation

Context Different rates of floodplain forest recruitment and mortality can reveal important changes in ecosystem processes that drive forest dynamics, resulting in net changes in forest cover, thereby influencing a wide range of river habitat and morphological characteristics. Objectives We evaluated characteristics of forest change areas in the Upper Mississippi River System. Methods An overlay technique was used to map patches of forest loss, gain, and persistence between 2010 and 2020 in relation to a series of explanatory variables. Results We quantified a net decline in forest cover ranging from 3.2 to 16.8% in the uppermost five study reaches, and a net increase in forest cover ranging from 0.5 to 4.6% in the southernmost three reaches. Patches of forest loss and persistence were similarly tall (> 15 m), dense (> 90% cover), silver maple ( Acer saccharinum ) dominated forests, whereas forest gain patches were short (< 15 m), less dense (< 66% cover) and more likely to be dominated by willow ( Salix) species. Both forest loss and gain patches were smaller than forest persistence patches and were typically found in areas with low neighborhood forest density (< 50% forested 10 ha neighborhood). Areas that experienced more than three flood events per growing season, more than 100 consecutive days of inundation during a single flood event, and more than 60 mean total days of inundation per growing season from 2011 to 2020 showed a net loss of forest cover in all study reaches. In contrast, net increases in forest cover were restricted to areas that experienced less than a single flood event per growing season, less than 40 consecutive days of inundation during a single flood event and less than 30 mean total days of inundation per growing season from 2011 to 2020. Conclusions Forest mortality along these river reaches is associated with forest fragmentation and an increasingly wetter hydrological regime.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

Proceedings of the Floodplain Vegetation Monitoring Workshop for the Long Term Resource Monitoring Element of the Upper Mississippi River Restoration Program, January 7–8, 2025, Moline, Illinois

Preface In anticipation for increased funding made possible by the Water Resources Development Act of 2020, the Upper Mississippi River Restoration (UMRR) Program identified a need to conduct river-wide assessments of floodplain vegetation. In January 2025, we assembled a group of subject matter experts to perform the following tasks: Review Upper Mississippi River Restoration’s current floodplain vegetation research portfolio, Identify important features and goals for long-term floodplain vegetation monitoring, Evaluate the suitability of existing datasets for system-wide vegetation assessments, and Discuss emerging opportunities to learn about floodplain vegetation dynamics from local-scale restoration and management projects. This document is a summarization of what occurred at the meeting and provides suggested next steps toward developing the capacity to conduct routine long-term monitoring and assessment of floodplain vegetation as part of the Upper Mississippi River Restoration Program.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

More water, more of the time: Spatial changes in flooding over 83 years in the upper Mississippi River floodplain and relationships with streamgage-derived proxies

The hydrologic regime of the upper Mississippi River (UMR) has become wetter, with greater discharges, longer-lasting high-flow conditions, and seasonal shifts in these patterns over the past several decades. How these changes are expressed spatially as floodplain inundation area, frequency, depth, duration, and timing is not well understood. It is also unclear to what degree spatial patterns of submergence are represented by examining discharge data alone. We assessed changes in floodplain inundation characteristics from 1940 to 2022 in navigation pools 3–10 of the UMR using a geospatial model to simulate daily inundation depths. Inundation characteristics shifted significantly across pools, but the direction and magnitude of change varied by pool and metric. Characteristics summarized at the pool scale correlated with streamgage-derived proxies but the strength of the relationship varied. Within pools, variability in inundation trends highlighted the importance of spatially explicit modeling. Our study demonstrates that changes in discharge over 83 years have manifested across the UMR floodplain in ways that may have consequences for ecological patterns and processes. By mapping hydrologically sensitive areas, we can anticipate which areas may be susceptible to additional shifts in river discharge in a climatically uncertain future.

Iowa, Minnesota, Wiscosnin↗

Evaluating hydrologic data products for scientific and management applications related to potential future streamflow conditions in the Upper Mississippi and Illinois Rivers

The hydrology of the Upper Mississippi and Illinois Rivers is a fundamental driver of ecosystem patterns and processes across a large portion of the United States. Quantitative hydrologic data for the main stems of these rivers underlie numerous scientific investigations, statistical models, and decision-making processes for local, State, and Federal agencies involved in the Upper Mississippi River Restoration program. Although historical hydrologic data exist, data representing potential future conditions of the Upper Mississippi and Illinois Rivers lack the resolution necessary to anticipate biotic and abiotic responses to altered hydrology and to determine resilient management actions. A source of future hydrologic scenarios is the readily available LOCA–VIC–mizuRoute hydrologic data products (named for the chain of models the data are produced from—localized constructed analogs, Variable Infiltration Capacity macroscale hydrological model, and the mizuRoute hydrologic routing model—that we shorten further to LVM in this report) that include simulated discharges for historic and future timeframes. The objective of this study is to assess the reliability of the hydrologic data products for their use in Upper Mississippi River Restoration program applications. Key study questions are (1) do the hydrologic data products reproduce characteristics of hydrology necessary to support ecological modeling and restoration decision-making applications within the Upper Mississippi River Restoration program? and (2) are there geographic differences in the reliability of the hydrologic data products? Seven characteristics of river hydrology were selected related to flow magnitude, seasonality, and regime for evaluation. The seven characteristics were calculated using observed and historical simulated hydrologic data at 19 U.S. Geological Survey streamgages throughout the basins of the Upper Mississippi and Illinois Rivers; two streamgages are located on the main stem of the Mississippi River and two streamgages are located on the main stem of the Illinois River. Statistical comparisons between observed and historical simulated characteristics indicated that the hydrologic data products did not reliably represent historical hydrologic conditions in the basin or main stem. The hydrologic data products we evaluated could not reliably capture the overall hydrologic regime or flow magnitudes; the latter is evidenced by substantial underestimates of discharge at most streamgages. Seasonal hydrologic characteristics were captured more reliably than flow magnitude, but overall correspondence was low for most streamgages. A weak latitudinal pattern in seasonal characteristics indicated the hydrologic data products poorly represent streamflow timing in snow-affected regions of the basin. Discrepancies in magnitude, seasonality, and regime indicate the potential for multiple sources of error. Because poor correspondence was present across all 19 streamgages, it was not possible to identify specific drivers of poor performance (that is, drainage area or geography). The modeling chain should be evaluated for biases associated with meteorologic forcing data, as well as hydrologic model formulation and calibration. We conclude that the hydrologic data products we evaluated appear unsuitable for applications tied to habitat and ecosystem restoration and management in the Upper Mississippi and Illinois Rivers. Plans to develop a future hydrology dataset for the Upper Mississippi River Restoration program would benefit from ongoing work to improve global climate model output downscaling methods, to improve hydrologic models, to make use of innovations in machine-learning approaches for projecting hydrology, and other efforts. The framework developed herein to evaluate hydrometeorological outputs generated using global climate models for a specific water resources application is a transferrable approach that could be applied to other data products and river systems.

Illinois, Indiana, Iowa, Minnesota, Missouri, Sout↗

Quantifying floodplain forest community change following large-scale flood events in the Upper Mississippi River System

Effects of large-scale flooding on forest composition and structure are a function of flood duration, depth, timing, and frequency. Throughout the Upper Mississippi River System (UMRS), floods in 1993 and 2019 were record-setting events followed by high rates of tree mortality. These events generated interest in species adaptations to flood event characteristics and how forest communities have changed in response to large-scale floods. We investigated associated tree mortality, how the floods differed spatially, and how floodplain forest communities have changed since 1993. Eight UMRS reaches were surveyed in a 1995 study, documenting vegetation species composition, size, and abundance. In 2021, a selection of plots (63%) were revisited and surveyed to quantify 2019 flood effects. For each site, we extracted daily inundation data for flood years and preceding decades from a surface water inundation model. We found post-flood mortality varied spatially and generally reflected inundation duration patterns. Lower latitude reaches experienced longer inundation durations and greater tree mortality in 1993 than in 2019, while higher latitude reaches experienced similar inundation duration and depth and similar mortality between events. Decadal inundation attributes also differed. During 2009–2018, inundation duration was greater and events occurred later than during 1983–1992 in all reaches. Most forest trajectories were Acer saccharinum -dominated and changed relatively little in species composition and structure. The greatest change in composition occurred at plots with high mortality from the 1993 flood, particularly in more flood-prone locations or where there were many small-diameter individuals. In plots dominated by either Quercus spp. or Populus deltoides , species importance shifted toward more shade and flood-tolerant species after 1995 surveys. Self-replacement of these species may be limited by a change in regeneration conditions resulting from an ongoing inundation regime shift in the case of Quercus spp., or succession to more shade-tolerant species in the case of Populus communities. Overall, effects on floodplain forests from the two flood events were heterogeneous. In some cases, forest change was likely just as influenced by shifts in flood regime as it was from singular flood events.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

Future forest conditions under alternative management and hydrological scenarios in the Upper Mississippi River floodplain

Context Floodplain forests are being transformed by multiple pressures, prompting widespread management and restoration efforts. It is uncertain how disturbances, including hydrologic change, and management actions will interact to influence the ecology of these threatened forests. Objectives This study examined the effects of alternative management and hydrologic regimes on forest succession at an Upper Mississippi River floodplain site with a restoration project in planning. Methods We used the spatially explicit forest landscape model, LANDIS-II, to simulate forest succession for 100 years under four hydrogeomorphic management scenarios, three forest management scenarios, and two scenarios of future hydrologic conditions. We evaluated changes in forest biomass and composition over time and assessed the relative importance of management actions and hydrologic change on succession. Results Forest aboveground biomass decreased in all management-hydrology scenarios, especially in the wetter hydrological scenario. Intensified hydrogeomorphic and forest management scenarios reduced the magnitude and extent of biomass declines; however, they were unable to prevent overall declines in biomass or cause large shifts in tree species composition. Silver maple ( Acer saccharinum ) was projected to decrease in biomass, while increases in biomass were projected for several late-successional species including swamp white oak ( Quercus bicolor ). Among the factors influencing variation in biomass, forest management had the largest influence in the first 50 years of our simulations, but hydrological regime became the most important factor by the end of the century. Conclusions Our simulations indicate that management actions could play an important role in the conservation of floodplain forests, but their effectiveness will likely be limited if recent upward trends in flooding conditions in this system continue in the future. Thus, our results highlight both the potential benefits and limitations of management actions in the face of hydrologic change.

Iowa, Minnesota, Wisconsin↗

Upper Mississippi River Restoration future hydrology meeting series

The Upper Mississippi River Restoration (UMRR) program, a broad partnership of State and Federal agencies administered by the U.S. Army Corps of Engineers, integrates ecosystem monitoring, research, and modeling to rehabilitate habitat and evaluate ecosystem trends over time in the Upper Mississippi River System. Hydrologic data are integral to the UMRR program because they are used in scientific research, decision-making, and restoration project planning. However, a lack of quantitative hydrologic data representing potential future conditions limits the ability to complete informative research on how future conditions may affect river ecology, achieve management goals, and design restoration projects for 50-year horizons. The U.S. Geological Survey and the U.S. Army Corps of Engineers led a series of workshops with UMRR partners to (1) prioritize needs for understanding future hydrology, (2) discuss appropriate datasets that could address these needs, and (3) develop a plan for acquiring and distributing a hydrologic dataset of potential future conditions. Agency priorities for understanding future hydrology were broad, spanning ecologic, geomorphic, resource management, and engineering disciplines, and were identified for a range of spatial (project site, navigation pool, reach, system) and temporal (daily, seasonal, annual) scales. The LOcalized Constructed Analogs-Variable Infiltration Capacity-mizuRoute hydrologic data products were identified as a potential source of off-the-shelf data to meet UMRR priority needs but warranted a robust quantitative evaluation. The final meeting in the series scoped a proposal to evaluate the LOcalized Constructed Analogs-Variable Infiltration Capacity-mizuRoute hydrologic data products for use in UMRR applications, including contingencies if the data were determined to be unreliable.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

Characterizing the niche of Phalaris arundinacea (reed canarygrass) in floodplain forests of the Upper Mississippi River

Information on the favorable conditions for invasive species as well as potential constraints to their distribution can be valuable for management efforts. We used a niche modeling approach to analyze the patterns of species distributions along gradients of hypothesized influential environmental variables. Many ecological datasets may have incomplete coverage across the environmental gradients, infrequent sampling under some conditions, insufficient time for an invasive species to occupy all sites, and complex interactions among environmental variables (measured or unmeasured) that may result in species response curves that are difficult to interpret and may be ecologically misleading. To ensure the model and species response curves aligned with ecological niche theory, shape constraints were imposed to guarantee relationships follow a unimodal distribution to reflect the fundamental niche (where a species could occur). We compared a shape-constrained model to an unconstrained model and interpreted the species response curves from the constrained model to better characterize the ecological niche of reed canarygrass in floodplain forests of the Upper Mississippi River, USA. We found the probability of reed canarygrass occurrence decreases with increasing tree canopy cover, tree species richness, distance from forest edge, distance from invaded wet meadows, and island isolation. Probability of reed canarygrass presence exhibited bell-shaped curves in response to hydrology (inundation depth, frequency, and duration) and forest stress metrics indicating an optimum with less favorable conditions on either end of the ecological gradients. This information could be used to prioritize restoration efforts and enhance landcover change research in forested floodplains.

upper Mississippi River↗

Workshop: Nature-based solutions to ecological and economic problems caused by extreme precipitation events in the Upper Mississippi River Basin

Future climate conditions in the Upper Mississippi River Basin are projected to include many more extreme precipitation events. These intense periods of rain can lead to flooding of the Mississippi River itself, as well the small streams and rivers that feed it. This flooding presents a challenge for local communities, farmers, small businesses, river users, and the ecosystems and wildlife in the area. To reduce the damage done by these extreme rainfall events, ‘natural solutions’ are often helpful. This might include preserving forests and grasslands to absorb rainwater before it arrives at streams or restoring wetlands to slow and clean runoff water. For river and natural resource managers to adapt to future climate by using these natural solutions, however, they require accurate information about the projected increase in extreme rainfall events and the effectiveness and costs of a variety of solutions. To provide this information, researchers must understand the decisions that managers are making and the challenges they face. In this project, researchers will host a workshop to facilitate knowledge sharing between natural resource managers, climate modelers, and scientists studying natural solutions. Participants will discuss climate projections related to extreme precipitation events in the region, as well as the management implications and decision-making challenges associated with these events. These conversations will cultivate collaborative relationships between managers and researchers to address critical knowledge gaps and improve future management efforts in the Upper Mississippi River Basin.

Upper Mississippi River basin↗

Mapping potential sensitivity to hydrogeomorphic change in the UMRS riverscape

In 2020 the U.S. Geological Survey (USGS), as part of the Upper Mississippi River Restoration (UMRR) Program, began a new project to characterize potential hydrogeomorphic change associated with hydrogeomorphic units (HGUs) and their catenae (units linked by their association with sediment sources and flow origins). The goal of the project was to develop a geographic information system (GIS) database of HGUs for the Upper Mississippi River System (UMRS) available to both scientists and river managers working on UMRR studies and HREP planning and design studies. The characterization was based on a hydrogeomorphic change hierarchical classification developed previously for the UMRS. The products were generated with automated techniques in a GIS using systemic datasets. Landforms were mapped from the 2015 UMRS topobathymetric dataset with geomorphon (shape-based) tools tailored for the large riverscape, valley bottom environments in the UMRS. A clustering analysis was applied to the resulting landforms to identify HGUs associated with zones of perennial low flows, bankfull flows, and overbank floodplains. Catenae were assembled based on the proximity of the units to the main channel, tributary mouths, and side channels from previously published aquatic areas (USACE, 2018) coupled with least-cost flowpath linkages between potential sediment origins and planform change units developed by Rogala, Fitzpatrick, and Henderson (2020). These GIS-based analyses were successful at identifying a range of HGUs using an automated technique with available data across the entire riverscape, with emphasis on those that have the potential for hydrogeomorphic change. Most of the resulting features are depositional, as expected in a large river system. However, this is the first attempt of linking tributary inputs, side channel erosion and levee breaches with their depositional counterparts. The approach was successfully piloted in Pools 8 and 10 in the Upper Impounded Reach and Pool 14 in the Lower Impounded Reach, with next steps for application in reaches of the unimpounded section and Illinois River. This report emphasizes results from Pool 10, which was the focus of most of our attention during the pilot phase.

Illinois, Indiana, Iowa, Minnesota, Missouri, Sout↗

Spatial differences in predicted Phalaris arundinacea (reed canarygrass) occurrence in floodplain forest understories

Reed canarygrass ( Phalaris arundinacea L.) is one of the most common invaders of floodplains and wetlands in North America. In the Upper Mississippi River floodplain, invasion by reed canarygrass in forest understories can inhibit forest regeneration when gaps form in the overstory. Understanding the distribution of reed canarygrass in forest understories is essential for effective management and control. We used an ensemble of species distribution models including Bayesian additive regression trees, boosted trees, and random forest algorithms to predict habitat suitability for reed canarygrass in forest understories across the Upper Mississippi River floodplain (~41,000 ha). Data from forest inventory study plots with reed canarygrass presence and absence were combined with 10 hypothesized environmental predictors of reed canarygrass invasion. We applied three approaches to better understand and incorporate the influence of spatial autocorrelation among our predictor variables, including random cross-validation, spatial cross-validation, and spatial cross-validation with Euclidean distance fields. Flood frequency, distance to contiguous floodplain, distance to forest edge, and distance to invaded wet meadow were among the most important environmental predictors across the three algorithms. Generally, the mean probability of reed canarygrass presence decreased with increasing flood depth, distance to contiguous floodplain, distance to invaded wet meadow, forest cover, and forest height, while relationships with other predictors were more variable. The ensemble of the three models (i.e., the average prediction) was used to map and summarize potential reed canary grass habitat suitability across the landscape. The maps generated quantified the habitat suitability for reed canarygrass and areas of agreement among the models in forest understories across the floodplain. This information can be used to better understand the extent of invasion, prioritize restoration efforts, and develop further research.

Illinois, Indiana, Iowa, Minnesota, Missouri, Wisc↗

The where and why of large wood occurrence in the Upper Mississippi and Illinois Rivers

Large wood (LW) plays important geomorphic and ecological roles in rivers and is widely used as a restoration tool. Changes to floodplain land use and historical removal have altered wood dynamics in fluvial systems globally. We know little about the distribution and dynamics of LW in great rivers (approximately >10 5 km 2 ) like the Upper Mississippi and Illinois Rivers despite its ecosystem importance and use in restoration projects. We assessed LW occurrence data collected by the fisheries component of the Upper Mississippi River Restoration Program's Long Term Resource Monitoring element. We analysed 25 years of data collected across six reaches of the Upper Mississippi and Illinois Rivers that represented contrasting physiographic settings, and across four aquatic area types comprising gradients of hydrology, connectivity and geomorphology. We tested hypotheses on drivers of LW occurrence using generalised linear mixed effects models, where occurrence was predicted by reach- and local-scale environmental variables. Occurrence varied significantly across reaches and aquatic area types. In general, wood occurred more frequently upriver and in side channels compared to other aquatic areas. Large wood was most strongly predicted systemically by reach identity but not local-scale variables, underscoring the importance of broad-scale physiographic gradients in defining hydrogeomorphic processes. Floodplain forests and shoreline revetment were consistently important predictors across reaches. Our findings show that the spatial variability of LW occurrence reflects the physical variability of the Upper Mississippi and Illinois Rivers. They also reveal the value in using geomorphic classifications as frameworks for understanding physical processes like LW dynamics because of their ability to contextualise site-scale conditions. The baseline understanding of LW abundance across different hydrogeomorphic gradients and scales presented here can give insight into how to more effectively target restoration efforts in great rivers and contribute to a broader understanding of LW dynamics where such studies have been lacking.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

Identifying conditions where reed canarygrass (Phalaris arundinacea) functions as a driver of forest loss in the Upper Mississippi River floodplain under different hydrological scenarios

Most of the world’s river-floodplain ecosystems are simultaneously undergoing modifications to their hydrological regimes and experiencing species invasions, making it unclear whether invasive species are the main drivers of ecosystem change or simply responding to changes in the hydrological regime. We simulated patterns of forest recruitment and succession in a 2500-ha portion of the Upper Mississippi River floodplain with and without removal of invasive Phalaris arundinacea and under two different future 100-year hydrological scenarios: a future maintaining the average flooding conditions of the past 40 years (random) and a future that projects an observed upward 40-year trend in flooding conditions forward (trending). By comparing scenarios that included Phalaris removal and ones that did not, we were able to identify the conditions where Phalaris was the main driver of forest loss vs. the conditions where hydrology was the main driver of forest loss. Areas where Phalaris was the main driver of forest loss had mean annual flood inundation durations that were similar to areas that did not lose forest cover (60–90 growing season days), while areas where flooding was the main driver of forest loss had longer mean inundation durations (102–124 growing season days). In comparison to the random hydrology scenario, the trending scenario produced a decrease in the area over which Phalaris was identified as the main driver of forest loss and an increase in the area over which flood inundation was identified as the main driver of forest loss. Thus, if the observed trends in flooding continue, our model projects an increase in the area over which eradicating Phalaris is unlikely to result in the maintenance of forest cover. We used the Resist-Accept-Direct (RAD) framework to discuss potential management options to resist changes and maintain forest cover where Phalaris is likely to be the main driver of forest loss and to accept or direct changes in areas where forest loss is likely driven by hydrological change.

Iowa, Minnesota, Wisconsin↗

Low-complexity floodplain inundation model performs well for ecological and management applications in a large river ecosystem

Flooding is a dominant physical process that drives the form and function of river-floodplain ecosystems. Efficiently characterizing flooding dynamics can be challenging, especially over geographically broad areas or at spatial and temporal scales relevant for ecosystem management activities. Here, we empirically evaluated a low-complexity geospatial model of floodplain inundation in six study segments of the Upper Mississippi River System (UMRS) by pairing spatially extensive, temporally limited and spatially limited, temporally extensive sampling designs. We found little evidence of systematic bias in model performance although discrepancies between model predictions and empirical data did occur locally. Assessments of model predictions revealed low segment-wide discrepancies of wetted extent under contrasting flow conditions and agreement for inundation event detection and duration. Model performance for predicting event frequency and duration was similar among sites expected to exhibit contrasting patterns of hydrologic connectivity with the main channel. Our results suggest that low-complexity models can efficiently characterize a critical physical process across geographically broad, complex river-floodplain ecosystems. Such tools have the potential for advancing scientific understanding of landscape-scale ecological patterns and for prioritizing management actions in large, complex river-floodplain ecosystems like the UMRS.

Illinois, Iowa, Minnesota, Missouri, Wisconsin↗

River geomorphology affects biogeochemical responses to hydrologic events in a large river ecosystem

Shifts in the frequency and intensity of high discharge events due to climate change may have important consequences for the hydrology and biogeochemistry of rivers. However, our understanding of event-scale biogeochemical dynamics in large rivers lags that of small streams. To fill this gap, we used high-frequency sensor data collected during four consecutive summers from a main channel and backwater site of the Upper Mississippi River. We identified high discharge events and calculated event concentration-discharge responses for both physical-chemical (nitrate, turbidity, and fluorescent dissolved organic matter) and biological (chlorophyll-a and cyanobacteria) constituents using metrics of hysteresis and slope. We found a range of responses across events, particularly for nitrate. Although fluorescent dissolved organic matter (FDOM) and turbidity exhibited more consistent responses across events, contrasting hysteresis metrics indicated that FDOM was flushed to the river from more distant sources than turbidity. Biological responses (chlorophyll a and cyanobacteria) differed more between sites than physical and chemical constituents. Lastly, we found that the event characteristics best explaining concentration responses differed between sites, with event magnitude more frequently related to responses in the main channel, and antecedent wetness conditions associated with response variation in the backwater. Our results indicate that event responses in large rivers are distinct across the diverse habitats and biogeochemical components of a large floodplain river, which has implications for local and downstream ecosystems as the climate shifts.

Minnesota, Wisconsin↗

Bridging the gap between mathematical biology and undergraduate education using applicable natural resource modeling

Mathematical biology is a wide field of study with many venues that undergraduate students can access through research. However, the topics of study for these students can be overwhelming, and many topics of study yield either only trivial results or abstract outcomes that are nonintuitive and diffcult to understand. We have used natural resource modeling, and more specifically, a partnership between academic researchers and federal scientists, as a bridge between undergraduate research and mathematical biology. Our collaboration is an interdisciplinary team that combines biology, mathematics, and statistics professors with government research scientists. As a team, we have mentored students through opportunities such as a Research Experiences for Undergraduates and other projects. In this article, we provide an overview of how we develop questions for undergraduates and outline two case studies, both of which resulted in peer reviewed journal articles. Last, we describe how we also transfer the results from these undergraduate projects to resource managers so the results may be applied to real world problems.

Letters in Biomathematics↗

Flood regimes alter the role of landform and topographic constraint on functional diversity of floodplain forests

Understanding patterns of species coexistence is a fundamental challenge in ecology. The physical environment is believed to play an important role, influencing patterns of dispersal and biotic interactions across space and time. Floodplain forest species are presumed to interact strongly with their environment, as evidenced by pronounced spatial variation in forest composition associated with flood-driven abiotic constraints. Questions of how, and to what degree, fine-scale heterogeneity interacts with broad-scale hydrogeomorphology to influence patterns of functional composition remain unresolved. We examined how functional diversity (i.e. richness and range of functional traits) and ecological strategies (i.e. functional trait combinations) of floodplain forest communities varied across regional and local gradients of flooding in a northern temperate region of the United States of America. We found functional diversity of woody overstory species varied across hydrogeomorphic settings and that different settings altered associations between functional diversity and both the relative elevation above and proximity to rivers. Ecological strategies shifted over local gradients of relative elevation and distance to channel with different magnitudes and directions depending on the broader hydrogeomorphic context. We found evidence that interactions among flood regimes and landform position impose different levels of functional constraint. These results indicate patterns of community assembly are not easily discerned from landform type or position alone, but rather from filtering that operates and interacts with biota over multiple spatial scales. Our results imply that it is important to characterize flood dynamics in ways that can be clearly linked to ecological processes and that treatment of floodplain landforms as transferable units across river-valley segments is problematic, even within a single basin.

Ecography↗

What is a stand? Assessing the variability of composition and structure in floodplain forest ecosystems across spatial scales in the Upper Mississippi River

The forest stand typically represents relatively homogenous forest conditions; the forest stand is generally the unit at which forest attributes are assessed, summarized, and subsequently managed. However, some ecosystems, such as the floodplain forests of the Upper Mississippi River (UMR), can exhibit high variability at fine spatial scales that can confound prescription development, implementation, and ultimate success of stand-level management actions. Here we assess how forest composition and structure vary within and across stand management units on the UMR and test at what spatial scale environmental variables relate to forest characteristics. We found that plot-level measures of composition, structure, and diversity were not well represented by site-level averages of these values. When basal area of all overstory species was combined, this variable was more closely related to “site” than to any of the environmental variables, but when analyzed by species, within-plot topographic variation (“microtopography”) was a significant positive predictor for both importance values of an individual species (swamp white oak ( Quercus bicolor Willd.)) as well as importance value of a specific group of species (oaks (Quercus spp.), bitternut hickory ( Carya cordiformis (Wangenh.) K. Koch), hackberry ( Celtis occidentalis L.), and American basswood ( Tilia americana L.)). This work highlights the challenges of using average stand conditions to summarize complex or heterogeneous systems and the need for flexibility and relaxed assumptions in defining management units in these forests.

Iowa, Minnesota, Wisconsin↗