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Michael L. Scott

Publications and source records attributed to Michael L. Scott.

At least 19 recordsLinked to original sources

Long-term geomorphic response of a southwestern USA river following establishment and removal of an invasive riparian tree

Invasion of non-native riparian vegetation along southwestern USA rivers is associated with channel narrowing and simplification, prompting numerous and varied removal efforts. Channel width and migration rate often, but not always, increase following treatment. The cause of this variability and the duration of response is poorly understood. Using flow records and aerial imagery we quantified measurement uncertainty, change in channel width and rates of floodplain formation and erosion relative to annual peak flows before and during the invasion of Russian olive ( Elaeagnus angustifolia L.), and following removal, along the Escalante River, Utah, over a fifty-year period. Prior to the invasion, the Escalante River was undergoing a decades-long narrowing process following large, turn-of-the-20th-century floods. Russian olive created a unique geomorphic shift in the observed pattern of channel change. Dense, channel-edge establishment and morphological traits including dense, inflexible branches, resulted in enhanced channel narrowing. Because the initial spread of Russian olive was from upstream to downstream, the Russian olive forest was wider and older upstream than downstream. Consequently, channel narrowing was greater and floodplain erosion rates had already decreased in upstream reaches compared to downstream. Russian olive removal increased channel width and floodplain erosion rates in upstream reaches, where Russian olive was most abundant. In contrast, downstream reaches continued to narrow. Small but detectable increases in rates of floodplain erosion across all reaches, and increased sinuosity in some, suggest the channel is becoming more mobile in the absence of Russian olive. Results indicate channel adjustment to Russian olive removal is spatially variable and may take a decade or more. With continued expansion of native riparian vegetation, future narrowing is likely during sustained low peak flows and large-scale widening is unlikely in the absence of extreme floods or physical removal of existing riparian vegetation.

Arizona, Utah

Downstream decreases in water availability, tree height, canopy volume and growth rate in cottonwood forests along the Green River, southwestern USA

Hydrologic stress is increasing in Fremont cottonwood ( Populus fremontii ) forests across the southwestern United States because of increased temperature and streamflow diversion. The spatial variability of this stress is large yet poorly understood. Along the Yampa and Green Rivers in Colorado and Utah, vapour pressure deficit and flow diversions increase downstream. To investigate effects of this gradient on cottonwoods, we measured the percent live canopy and height of randomly selected trees at three sites: Deerlodge Park on the Yampa River (DLP), Island Park on the upper Green (ILP) and Canyonlands National Park on the lower Green (CAN). From these same trees, we took increment cores to understand differences in tree growth in each forest over time. We then related tree metrics to local water availability, streamflow and climatic data. Cottonwoods at CAN were shorter and had lower percent live canopy and growth rate than similarly aged trees upstream. CAN trees that grew higher above the water surface also tended to have lower tree growth, height and live canopy percentage. Furthermore, the correlation between tree growth and maximum vapour pressure deficit showed a much stronger negative shift since 1990 at CAN than at the other sites. All of these differences suggest higher hydrologic stress at CAN, which we attribute to the combined effects of peak flow declines from Flaming Gorge Reservoir, flow diversion and the higher and increasing vapour pressure deficit at CAN. Further research on the variability of hydrologic stress on cottonwoods could help managers anticipate and mitigate the effects of drought stress in these iconic forests.

Ecohydrology

Response of riparian vegetation to short- and long-term hydrologic variation

Increasing demand for river water now conflicts with an increasing desire to maintain riparian ecosystems. Efficiently managing river flows for riparian vegetation requires an understanding of the time scale of flow effects, but this information is limited by the absence of long-term studies of vegetation change in response to flow variation. To investigate the influence of short- and long-term flow variability and dam operation on riparian vegetation, we determined the occurrence of 107 plant species in 133 permanent plots of known inundating discharge along the Gunnison River in Colorado on five different occasions between 1990 and 2013. Individual species moved up and down the gradient of inundating discharge coincident with increases and decreases in mean annual flow, and the correlations between flow and species occurrence were strongest when flows were weighted by time before vegetation sampling with a median half-life of 1.5 years. Some tall, rhizomatous, perennial species, however, responded to flows on a longer time scale. Logistic regression of species occurrence showed a significant relation with inundation duration for 70 out of 107 species. Plot species richness and total vegetative cover decreased in association with desiccation at low inundation durations and with fluvial disturbance at high inundation durations. Within-plot similarity in species occurrence between years decreased strongly with increasing inundation duration. Moderate inundation durations were dominated by tall, rhizomatous, perennial herbs, including invasive Phalaris arundinacea (reed canary grass). Over the 23-year study period, species richness declined, and the proportion of rhizomatous perennials increased, consistent with the hypothesis that decreases in flow peaks and increases in low flows caused by flow regulation have decreased establishment opportunities for disturbance-dependent species. In summary, annual-scale changes in vegetation were strongly influenced by flow variation, and decadal-scale changes were influenced by decreases in fluvial disturbance from upstream flow regulation beginning decades prior to the onset of this study.

Colorado

Sediment-ecological connectivity in a large river network

Sediment eroded from the headwaters of a large basin strongly influences channels and ecosystems far downstream, but the connection is often difficult to trace. Disturbance-dependent riparian trees are thought to rely primarily on floods for formation of the sand bars necessary for seedling establishment, but pulses of sediment should also promote formation of such features. In order to expand understanding of the role of sediment connectivity in governing ecological processes, here we explore the hypothesis that cottonwood forest along the Green and Yampa Rivers in Utah and Colorado are dominated by trees established a century ago during a period of extensive channel migration caused by significant headwater erosion. Analysis of historical documents and aerial photographs suggests that three key tributaries of the Yampa River underwent significant historical erosion from roughly 1880 to 1940. Average width and depth of tributaries with defined arroyos increased two to six times from historical surveys, resulting in the export of ~30 million metric tons of sediment, sizably increasing the sediment load and channel migration rate of the Yampa and Green Rivers. Establishment of major portions of several downstream cottonwood forests occurred during this period of historical erosion, increased sediment loads, and heightened channel migration rates, and the area of forest dating to that time is much greater than can be explained by high flows alone. Viewed collectively, our findings suggest tributary erosion played a vital role in successful downstream forest establishment, a link we contend is best illustrated through a sediment-ecological connectivity framework. Broadly, this framework facilitates consideration of linkages between morphological and ecological processes at the watershed-scale. Development and utilization of a watershed-scale sediment-ecological connectivity perspective highlights the value of sediment as a critical ecological resource to be managed jointly with flow to ensure the maintenance of vital riverine ecosystems.

Colorado, Utah

Ecological interfaces between land and flowing water: Themes and trends in riparian research and management

This paper provides an overview of past, present and future themes for research and management of riparian zones, often relating to papers within this Wetlands Special Feature. Riparian research expanded in the United States around 1980 with themes that recognized (1) damage from excessive livestock, or (2) damage from river damming and diversion, and (3) the beneficial capacity of riparian buffers to intercept and assimilate nutrients and other water contaminants. Research expanded globally in the 1990s, with themes including (4) plant life history requirements and (5) reliance on fluvial geomorphic dynamics that enable riparian rejuvenation. Resource managers recognized that riparian areas provide (6) rich wildlife habitats (7) along with valued ecosystem services, (8) which encouraged conservation and restoration initiatives, (9) including environmental flow regimes. Floodplains are (10) vulnerable to invasive plants and management has included biocontrol such as for Tamarix in the American Southwest. Into the twenty-first century, (11) climate change is advancing, and riparian ecosystems may be especially impacted due to the compound challenges from increasing water demand and declining summer flows. As an emerging opportunity, (12) while reservoirs submerge floodplain vegetation, reservoir deltas may support compensatory riparian wetlands. (13) Studies increasingly utilize remote sensing tools including satellite imagery, LiDAR and unmanned aircraft systems, and (14) the coordination of large data sets invites digital ecology, including artificial intelligence and machine learning. Since riparian zones are centres for human activities, (15) there are opportunities for citizen science, social media and internet applications, which will increasingly democratize riparian research and management.

Wetlands

The role of a non-native tree in riparian vegetation expansion and channel narrowing along a dryland river

Along rivers, native and invasive species may establish and persist on active channel bedforms as part of channel narrowing. Using historical aerial photography and dendrochronology, we quantified spatial and temporal patterns of narrowing and vegetation expansion, including native Fremont cottonwood (Populus fremontii) and non‐native Russian olive (Elaeagnus angustifolia), along the largely unregulated Escalante River in south‐western United States. Russian olive establishment was examined with respect to hydrologic and climate variables. Narrowing along the Escalante River was initiated during a mid‐20th century drought. Cottonwood rapidly colonized higher, bar surfaces between the 1950s and 1981. Small numbers of Russian olive established in moist sites during this period as the channel narrowed by nearly 80%. After 1981, there was no obvious cottonwood establishment but low channel bars and banks were rapidly colonized by Russian olive. Hydroclimate predictors were equivocal but exponential growth of this large‐seeded, shade‐tolerant species lagged its introduction by 30 years, apparently because of delayed reproductive maturity, limited seed availability, and widespread availability of favourable establishment sites following initial channel narrowing. Sediment trapping, levee formation, and modification of channel form by dense, channel‐edge bands of Russian olive progressively limited new establishment sites and by 2000, recruitment declined sharply. Our results have implications for management of non‐native tree invasions along arid‐region rivers, including identification of low, moist, active channel bars where the establishment and physical impacts of Russian olive appear to be most pronounced and where focused management efforts are likely to be most effective.

Utah

Evaluating riparian vegetation change in canyon-bound reaches of the Colorado River using spatially extensive matched photo sets

Much of what we know about the functional ecology of aquatic and riparian ecosystems comes from work on regulated rivers (Johnson et al. 2012). What little we know about unregulated conditions on many of our larger rivers is often inferred from recollections of individuals, personal diaries, notes, maps, and collections from early scientific surveys (Webb et al. 2007) and from repeat photography (Turner and Karpiscak 1980; Webb 1996). Later surveys often included photographs typically taken at specific and relatively few locations. Because rivers and their related riparian ecosystems vary in both longitudinal and lateral dimensions, such records provide relatively general and incomplete information about historical resource conditions, especially at large spatial scales and over long spans of time.

Arizona, Colorado

River flow and riparian vegetation dynamics - implications for management of the Yampa River through Dinosaur National Monument

This report addresses the relation between flow of the Yampa River and occurrence of herbaceous and woody riparian vegetation in Dinosaur National Monument (DINO) with the goal of informing management decisions related to potential future water development. The Yampa River in DINO flows through diverse valley settings, from the relatively broad restricted meanders of Deerlodge Park to narrower canyons, including debris fan-affected reaches in the upper Yampa Canyon and entrenched meanders in Harding Hole and Laddie Park. Analysis of occurrence of all plant species measured in 1470 quadrats by multiple authors over the last 24 years shows that riparian vegetation along the Yampa River is strongly related to valley setting and geomorphic surfaces, defined here as active channel, active floodplain, inactive floodplain, and upland. Principal Coordinates Ordination arrayed quadrats and species along gradients of overall cover and moisture availability, from upland and inactive floodplain quadrats and associated xeric species like western wheat grass (Pascopyrum smithii), cheatgrass (Bromus tectorum), and saltgrass (Distichlis spicata) to active channel and active floodplain quadrats supporting more mesic species including sandbar willow (Salix exigua), wild licorice (Glycyrrhiza lepidota), and cordgrass (Spartina spp.). Indicator species analysis identified plants strongly correlated with geomorphic surfaces. These species indicate state changes in geomorphic surfaces, such as the conversion of active channel to floodplain during channel narrowing. The dominant woody riparian species along the Yampa River are invasive tamarisk (Tamarix ramosissima), and native Fremont cottonwood (Populus deltoides ssp. wislizenii), box elder (Acer negundo L. var. interius), and sandbar willow (Salix exigua). These species differ in tolerance of drought, salinity, inundation, flood disturbance and shade, and in seed size, timing of seed dispersal and ability to form root sprouts. These physiological and ecological differences interact with flow variation and geomorphic setting, resulting in differential patterns of occurrence. For example, in park settings cottonwood is far more abundant than box elder, while the reverse is true in canyons. Synthesis of existing knowledge from the Yampa and Green rivers and elsewhere suggests that the following flow-vegetation relations can be used to assess effects of future flow alterations in the Yampa River. High variability in flow within and between years removes vegetation through erosion, extended inundation and desiccation, creating the broad, open surfaces in and near the channel that are characteristic of lightly regulated rivers in western North America. This flow variability provides opportunities for establishment of disturbance-dependent riparian species. Flow regulation that results in lower peak flows and higher low flows allows proliferation of woody riparian vegetation, mostly tamarisk in canyon reaches, but both tamarisk and cottonwood in parks. Denser near-channel vegetation promotes sediment deposition leading to channel narrowing. Decreasing flow variability also increases area of species associated with extremely high and low inundation durations relative to species associated with moderate inundation duration. In addition, such flow regulation decreases occurrence of species tolerant of fluvial disturbance, while increasing occurrence of species tolerant of extended inundation. Over the long term, establishment of cottonwood and tamarisk requires disturbance by large floods, which provides openings for new individuals. At the annual time scale, establishment can occur in any year or location that provides a moist, open surface free from frequent future disturbance. In canyons, where channel movement is limited, low surfaces are too frequently disturbed for long-term survival of cottonwood, and establishment requirements are generally met only in years of moderate to high peak flows. In park settings cottonwood establishment may also occur in years of low peak flows where survival is promoted by movement of the channel away from the seedling. Peak flows early in the growing season promote establishment of cottonwood and sandbar willow seedlings relative to those of tamarisk. This is because cottonwood and willow seed release occurs early in the summer, while that of tamarisk occurs later. Late season seed release of tamarisk allows it to establish lower on the floodplain than cottonwood. Because of its shade tolerance and the energy stored in its large seeds, box elder can become established beneath existing vegetation, an ability not shared by cottonwood, tamarisk or willow. Although box elder does not require flood disturbance, it does take advantage of soil moisture from floods, which allow this species to become established high above the channel. Decreases in flow peaks, volumes or base flows decrease growth and survival of cottonwood relative to drought-tolerant tamarisk. Storing water from the spring peak in a reservoir for release after the April-July cottonwood growth window may also decrease growth and survival of cottonwood relative to tamarisk. Decreases in peak flows decrease floodplain inundation, which can reduce growth of floodplain species by preventing recharge of the floodplain aquifer. Two or more years in a row with similar flows promote establishment of woody vegetation. Subsequent sediment deposition around this vegetation, especially if the vegetation is tamarisk, results in channel narrowing and simplification. Rapid declines in the descending limb of the hydrograph kill riparian woody seedlings by desiccation. Fluctuations in the descending limb can kill seedlings by desiccation and inundation. Thus rapid declines and fluctuations would be counterproductive following early-season peaks prescribed to promote cottonwood, but would be consistent with the goal of preventing tamarisk establishment following a late-season peak. The tendency of regulated flows to keep returning to a small number of fixed discharge values (such as power plant capacity or a fixed minimum flow) can cause unnaturally sharp banding of geomorphic surfaces, topography and vegetation that is not necessarily erased by large flood peaks. Changes in sediment load relative to transport capacity may promote channel change especially in alluvial settings. For example, decreases in sediment input from the Little Snake River Basin since 1960 (or earlier) could be associated with channel narrowing and temporary increases in establishment of both cottonwood and tamarisk along the Yampa River. Increases in salinity of water or soil promote tamarisk over the native woody species. Even if water salinity does not increase, floodplain soil salinity can be increased by decreasing the flushing caused by overbank flooding.

Dinosaur National Monument, Yampa River

Distribution of invasive and native riparian woody plants across the western USA in relation to climate, river flow, floodplain geometry and patterns of introduction

Management of riparian plant invasions across the landscape requires understanding the combined influence of climate, hydrology, geologic constraints and patterns of introduction. We measured abundance of nine riparian woody taxa at 456 stream gages across the western USA. We constructed conditional inference recursive binary partitioning models to discriminate the influence of eleven environmental variables on plant occurrence and abundance, focusing on the two most abundant non-native taxa, Tamarix spp. and Elaeagnus angustifolia , and their native competitor Populus deltoides . River reaches in this study were distributed along a composite gradient from cooler, wetter higher-elevation reaches with higher stream power and earlier snowmelt flood peaks to warmer, drier lower-elevation reaches with lower power and later peaks. Plant distributions were strongly related to climate, hydrologic and geomorphic factors, and introduction history. The strongest associations were with temperature and then precipitation. Among hydrologic and geomorphic variables, stream power, peak flow timing and 10-yr flood magnitude had stronger associations than did peak flow predictability, low-flow magnitude, mean annual flow and channel confinement. Nearby intentional planting of Elaeagnus was the best predictor of its occurrence, but planting of Tamarix was rare. Higher temperatures were associated with greater abundance of Tamarix relative to P. deltoides , and greater abundance of P. deltoides relative to Elaeagnus. Populus deltoides abundance was more strongly related to peak flow timing than was that of Elaeagnus or Tamarix . Higher stream power and larger 10-yr floods were associated with greater abundance of P. deltoides and Tamarix relative to Elaeagnus . Therefore, increases in temperature could increase abundance of Tamarix and decrease that of Elaeagnus relative to P. deltoides , changes in peak flow timing caused by climate change or dam operations could increase abundance of both invasive taxa, and dam-induced reductions in flood peaks could increase abundance of Elaeagnus relative to Tamarix and P. deltoides .

Arizona, California, Colorado, Idaho, Kansas, Mont

Processes of arroyo filling in northern New Mexico, USA

We documented arroyo evolution at the tree, trench, and arroyo scales along the lower Rio Puerco and Chaco Wash in northern New Mexico, USA. We excavated 29 buried living woody plants and used burial signatures in their annual rings to date stratigraphy in four trenches across the arroyos. Then, we reconstructed the history of arroyo evolution by combining trench data with arroyo-scale information from aerial imagery, light detection and ranging (LiDAR), longitudinal profiles, and repeat surveys of cross sections. Burial signatures in annual rings of salt cedar and willow dated sedimentary beds greater than 30 cm thick with annual precision. Along both arroyos, incision occurred until the 1930s in association with extreme high flows, and subsequent filling involved vegetation development, channel narrowing, increased sinuosity, and finally vertical aggradation. A strongly depositional sediment transport regime interacted with floodplain shrubs to produce a characteristic narrow, trapezoidal channel. The 55 km study reach along the Rio Puerco demonstrated upstream progression of arroyo widening and filling, but not of arroyo incision, channel narrowing, or floodplain vegetation development. We conclude that the occurrence of upstream progression within large basins like the Rio Puerco makes precise synchrony across basins impossible. Arroyo wall retreat is now mostly limited to locations where meanders impinge on the arroyo wall, forming hairpin bends, for which entry to and exit from the wall are stationary. Average annual sediment storage within the Rio Puerco study reach between 1955 and 2005 was 4.8 × 10 5 t/yr, 16% of the average annual suspended sediment yield, and 24% of the long-term bedrock denudation rate. At this rate, the arroyo would fill in 310 yr.

New Mexico

Riparian restoration in the context of Tamarix control in the western United States: Chapter 23

This chapter focuses on the restoration of riparian systems in the context of Tamarix control—that is, Tamarix -dominated sites are converted to a replacement vegetation type that achieves specific management goals and helps return parts of the system to a desired and more natural state or dynamic. It reviews research related to restoring native riparian vegetation following tamarix control or removal. The chapter begins with an overview of objective setting and the planning of tamarix control and proceeds by emphasizing the importance of considering site-specific factors and of context in selecting and prioritizing sites for restoration. In particular, it considers valley and bottomland geomorphology, along with river flow regime and associated fluvial disturbance, surface water and groundwater availability, and soil salinity and texture. The chapter concludes with a discussion of costs and benefits associated with active, passive, and combined ecological restoration approaches, as well as the key issues to consider in carrying out restoration projects at a range of scales.

Book chapter

Woody riparian vegetation near selected streamgages in the western United States

Areal cover and occupancy of woody riparian species near 456 streamgages in the western United States were obtained from site visits during the growing seasons of 1996-2002. We made concomitant estimates of grazing intensity, channel stabilization and incision, gradient, sediment particle size, and nearby planting of Russian olive. The purpose of this publication is to describe the data set and make it available to other investigators in an electronic format.

Arizona;California;Colorado;Idaho;Kansas;Montana;N

Germination and establishment of the native plains cottonwood ( Populus deltoides Marshall subsp. monilifera ) and the exotic Russian-olive ( Elaeagnus angustifolia L.)

Russian-olive ( Elaeagnus angustifolia ) is a small Eurasian tree that has escaped from cultivation and become naturalized, primarily along watercourses throughout the western United States. We examined germination and establishment of Russian-olive and plains cottonwood ( Populus deltoides ), the principal native riparian tree of the Great Plains, under a range of experimental moisture and light conditions. The fewest seedings established under the driest conditions; seedling biomass was predictably lower in the shade; root-to-shoot ratios were higher for cottonwood, higher in the sun, and higher under drier conditions. Several interactions were also significant. The timing of germination and mortality varied between plains cottonwood and Russian-olive: cottonwood germinated in mid-June in all treatments in a single pulse with subsequent mortality; the timing and amount of Russian-olive germination differed substantially across treatments with little net mortality. Differences in life-history traits of these species, including seed size, viability, and dispersal, help explain treatment differences. Russian-olive will likely remain an important component of riparian communities along both unregulated and regulated western rivers because it succeeds under conditions optimal for cottonwood establishment and under many conditions unfavorable for cottonwood. Furthermore, many western states still encourage planting of Russian-olive, and control techniques tend to be labor-intensive and expensive .

Conservation Biology

Field-based evaluations of sampling techniques to support long-term monitoring of riparian ecosystems along wadeable streams on the Colorado Plateau

To better plan for and implement long-term ecological monitoring, we measured riparian vegetation and fluvial geomorphic features at pilot study sites on four wadeable perennial stream reaches, representative of drainages across the Colorado Plateau. Our primary objectives were to (1) collect field data, (2) evaluate the efficiency and effectiveness of various ecological measures and measurement techniques for riparian ecosystems, and (3) use field-based sampling to inform and refine the development of standard operating procedures for use in implementing integrated, long-term monitoring of riparian ecosystems. Ultimately, this work was aimed at providing NPS staff with some of the information and methods needed to design and implement long-term monitoring of NPS riparian resources, which is both relevant to management, and fully operational within institutional resource constraints. Our results suggest that selecting sampling reaches and establishing a sampling frame of 11 transects, across a range of stream types, is feasible given a limited set of decision rules. A distinctive feature of richness across all sites was the high percentage of rare species, defined here as species having a single occurrence at a site. Rare species represented from 33 percent to 47 percent of the species total across the four pilot sites. Our data show that the two smallest quadrat sizes, 0.01 m 2 and 0.1 m 2 , rarely had any species that occurred in the desired frequency range and can be omitted from the monitoring protocol. Few species fell within the 30&ndash;70 percent range in the 1-m 2 quadrats, but this quadrat size appears to be useful at the Tsaile Creek (CACH) site. We recommend continuing to collect information at the 1-m 2 scale and reevaluating its usefulness after more data are available from different types of sites. The 10-m 2 quadrat is adequate for monitoring changes in frequencies of very common species at all sites. Based on pilot study results, we conclude that at sites with low total species numbers (< 60 species), 40&ndash;60, 10-m 2 quadrats, would be sufficient to characterize overall species diversity for relatively common species. At sites with higher total numbers of species (> 100), 60&ndash;80, 10-m 2 quadrats would be required to characterize overall species diversity. Rare species of interest should be monitored using alternative approaches, such as a site inventory and/or mapping (see Elzinga and others, 1998). A large number of the systematically placed 10-m 2 quadrats span two or more geomorphic surfaces, especially adjacent to the channel. This makes resolution of species affinities with distinct geomorphic landforms difficult. Thus, we provide an amendment to improve characterization of herbaceous and shrub species on narrow, near-channel surfaces by sampling additional 0.5-m by 1-m quadrats on those surfaces. It appears that for sites in narrow valley settings where riparian zones average less than approximately 40 m, the number of 10-m 2 quadrats systematically placed on 11 transects will not provide shrub cover estimates at 20 percent precision. In such cases, additional sample reaches should be added in order to attain a minimum of 130 to 140 10-m 2 shrub quadrats. The line-intercept technique can provide a relatively rapid, reach-scale quantification of proportional cover for woody vegetation and geomorphic surface types and that variance in these measures stabilizes by the eighth or ninth transect sampled. An overlay of the distribution of geomorphic surface data derived from line-intercept sampling on topographic survey information indicates that delineation of geomorphic surfaces could be done in conjunction with the topographic survey of each transect, obviating the need to record surface breaks using the line intercept. To include geomorphic surface identifications with the topographic survey, surface breaks and transitional surfaces should be included and identified in the survey, in addition to systematically placed survey points. Compared to 5-m by 20-m tree quadrats, belt transects were shown to provide similar estimates of stand structure (stem density and stand basal area) in less than 30 percent of the time. Further, for the streams sampled, there were no statistically significant differences in stem density and basal area estimates between 10-m and 20-m belt transects and the smaller belts took approximately half the time to sample. There was, however, high variance associated with estimates of stand structure for infrequently occurring stems, such as large, relict or legacy riparian trees. Legacy riparian trees occurred in limited numbers at all sites sampled. A reachscale population census of these trees indicated that the 10-m belt transects tended to underestimate both stem density and basal area for these riparian forest elements and that a complete reach-scale census of legacy trees averaged less than one hour per site.

Colorado Plateau

Cottonwood in the Missouri Breaks National Monument

Concerns about cottonwood along the Wild and Scenic reach of the upper Missouri River include declining forests of sparse old trees with little recruitment of new individuals, impacts of cattle crazing and recreational use, and effects of flow alterations from operation of upstream dams and changes in tributary inflows.

Missouri

Flood dependency of cottonwood establishment along the Missouri River, Montana, USA

Flow variability plays a central role in structuring the physical environment of riverine ecosystems. However, natural variability in flows along many rivers has been modified by water management activities. We quantified the relationship between flow and establishment of the dominant tree (plains cottonwood, Populus deltoides subsp. monilifera ) along one of the least hydrologically altered alluvial reaches of the Missouri River: Coal Banks Landing to Landusky, Montana. Our purpose was to refine our understanding of how local fluvial geomorphic processes condition the relationship between flow regime and cottonwood recruitment. We determined date and elevation of tree establishment and related this information to historical peak stage and discharge over a 112-yr hydrologic record. Of the excavated trees, 72% were established in the year of a flow >1400 m 3 /s (recurrence interval of 9.3 yr) or in the following 2 yr. Flows of this magnitude or greater create the necessary bare, moist establishment sites at an elevation high enough to allow cottonwoods to survive subsequent floods and ice jams. Almost all cottonwoods that have survived the most recent flood (1978) were established >1.2 m above the lower limit of perennial vegetation (active channel shelf). Most younger individuals were established between 0 and 1.2 m, and are unlikely to survive over the long term. Protection of riparian cottonwood forest along this National Wild and Scenic section of the Missouri River depends upon maintaining the historical magnitude, frequency, and duration of floods >1400 m 3 /s. Here, a relatively narrow valley constrains lateral channel movement that could otherwise facilitate cottonwood recruitment at lower flows. Effective management of flows to promote or maintain cottonwood recruitment requires an understanding of locally dominant fluvial geomorphic processes.

Montana