Geology topics

Mark K. Sogge

Publications and source records attributed to Mark K. Sogge.

At least 19 recordsLinked to original sources

USGS2018

Patch age since disturbance drives patch dynamics for flycatchers breeding in both reservoir and riverine habitat

Species dependent upon early-successional landscapes often occupy patches at different stages of recovery after disturbance. The demographic processes that drive patch dynamics in these systems have rarely been described but are important for developing effective conservation and management plans, especially when humans have modified the timing and intensity of disturbances that drive regeneration. In riparian systems, disturbance by floods historically initiated plant regeneration, but many rivers are now regulated and stream flows disrupted by dams and reservoirs. We studied the demography and patch dynamics of an endangered, neotropical migrant bird dependent on remnant riparian patches for breeding, the southwestern willow flycatcher ( Empidonax trailli extimus ), over 9 yr at both a riverine and reservoir site in central Arizona. We found that at both sites, number of territories/ha within patches increased for 2–4 yr after colonization and then declined, with several patches abandoned after 6–10 yr. Age of birds increased with patch age, with younger birds in colonizing patches and older, site-faithful birds in older patches, while mean per capita reproductive success did not differ with patch age. Natal dispersal and breeding dispersal were primarily from intermediate-aged patches into either young- or other intermediate-aged patches. At both riverine and reservoir sites, both the number of patches and the number of territorial birds increased over time, with the percentage of territories shifting into younger and younger patches. The type of disturbance driving patch regeneration differed between riverine and reservoir sites (seasonal flooding vs. falling lake levels due to drought), but the demographic patterns did not, indicating that reservoirs can generate patch dynamics similar to those on rivers. Managing stream flows and reservoir levels to maintain disturbance cycles sufficient to generate riparian patches at different stages of regeneration through time would benefit succession-dependent species like the endangered flycatcher we studied, whether those disturbances arise from natural flooding events along free-flowing rivers or through changes in reservoir levels.

Ecosphere↗

USGS2018

Extreme drought alters frequency and reproductive success of floaters in Willow Flycatchers

Changes in habitat quality, including those caused by extreme events like droughts and floods, could alter costs and benefits of territoriality and thereby the prevalence and reproductive consequences for individuals capable of breeding that do not do so (floaters). We studied floating behavior in a population of Southwestern Willow Flycatchers ( Empidonax traillii extimus ) in central Arizona during one year of extreme drought, one year of lake inundation, and three years of near average precipitation. In all years, most floaters were second year (SY) males, and most subsequently settled outside of the patch where they were detected in the floating year, suggesting that floaters did not “queue” at high-quality territories in order to achieve higher reproductive success in subsequent years. Instead, cohorts that floated in non-drought years had lower apparent survival and lower reproductive success compared to territorial birds. In the extreme drought year, however, the number of floaters was 1.5 times greater than in all other years combined, more females floated, and apparent survival and mean annual productivity in subsequent years was higher for males that floated in that year than for those that were territorial. Inundation of habitat due to rising reservoir levels did not result in an increase in floaters because many birds nested in inundated areas where trees projected above the water so that the relative amount of available habitat was not reduced to the extent habitat models predicted. Overall, our results indicate that the prevalence and reproductive and demographic consequences of floating can change under extreme climatic events like severe drought.

The Auk↗

USGS2017

Survivorship across the annual cycle of a migratory passerine, the willow flycatcher

Annual survivorship in migratory birds is a product of survival across the different periods of the annual cycle (i.e. breeding, wintering, and migration), and may vary substantially among these periods. Determining which periods have the highest mortality, and thus are potentially limiting a population, is important especially for species of conservation concern. To estimate survival probabilities of the willow flycatcher Empidonax traillii in each of the different periods, we combined demographic data from a 10-year breeding season study with that from a 5-year wintering grounds study. Estimates of annual apparent survival for breeding and wintering periods were nearly identical (65–66%), as were estimates of monthly apparent survival for both breeding and wintering stationary periods (98–99%). Because flycatchers spend at least half the year on the wintering grounds, overall apparent survivorship was lower (88%) on the wintering grounds than on the breeding grounds (97%). The migratory period had the highest mortality rate, accounting for 62% of the estimated annual mortality even though it comprises only one quarter or less of the annual cycle. The migratory period in the willow flycatcher and many other neotropical migrants is poorly understood, and further research is needed to identify sources of mortality during this crucial period.

Journal of Avian Biology↗

USGS2013

Tamarisk in riparian woodlands: A bird’s eye view

This chapter presents a “bird's eye” view of tamarisk and examines some issues surrounding the management of tamarisk in riparian woodlands. The focus on birds is based on the fact that they are a relatively well-studied group that can provide important insights into the role of tamarisk in riparian ecosystems. Because the decline of native riparian habitat occurred concurrently with the spread of tamarisk, this invasive species has been portrayed as a key factor in the reduction of riparian breeding bird numbers. The chapter begins with an overview of the early perceptions and realities of why and how birds use tamarisk before turning to a discussion of the history of tamarisk control and its effects on birds. It then considers some of the changing perspectives about the management of tamarisk and riparian habitats in western North America.

Book chapter↗

USGS2011

Winter distribution of willow flycatcher subspecies

Documenting how different regions across a species' breeding and nonbreeding range are linked via migratory movements is the first step in understanding how events in one region can influence events in others and is critical to identifying conservation threats throughout a migratory animal's annual cycle. We combined two studies that evaluated migratory connectivity in the Willow Flycatcher ( Empidonax traillii ), one using mitochondrial DNA sequences from 172 flycatchers sampled throughout their winter range, and another which examined morphological characteristics of 68 museum specimens collected in the winter range. Our results indicate that the four subspecies occupy distinct but overlapping regions of the winter range. Connectivity between specific breeding and winter grounds appears to be moderate to strong, with distributions that suggest migration patterns of both the chain and leap-frog types connecting the breeding and nonbreeding grounds. The Pacific lowlands of Costa Rica appear to be a key winter location for the endangered Southwestern Willow Flycatcher ( E. t. extimus ), although other countries in Central America may also be important for the subspecies.

The Condor↗

USGS2011

Tamarisk biocontrol using tamarisk beetles: Potential consequences for riparian birds in the southwestern United States

The tamarisk beetle ( Diorhabda spp.), a non-native biocontrol agent, has been introduced to eradicate tamarisk ( Tamarix spp.), a genus of non-native tree that has become a dominant component of riparian woodlands in the southwestern United States. Tamarisk beetles have the potential to spread widely and defoliate large expanses of tamarisk habitat, but the effects of such a widespread loss of riparian vegetation on birds remains unknown. We reviewed literature on the effects of other defoliating insects on birds to investigate the potential for tamarisk beetles to affect birds positively or negatively by changing food abundance and vegetation structure. We then combined data on the temporal patterns of tamarisk defoliation by beetles with nest productivity of a well-studied riparian obligate, the Southwestern Willow Flycatcher ( Empidonax traillii extimus ), to simulate the potential demographic consequences of beetle defoliation on breeding riparian birds in both the short and long term. Our results highlight that the effects of tamarisk biocontrol on birds will likely vary by species and population, depending upon its sensitivity to seasonal defoliation by beetles and net loss of riparian habitat due to tamarisk mortality. Species with restricted distributions that include areas dominated by tamarisk may be negatively affected both in the short and long term. The rate of regeneration and/or restoration of native cottonwoods ( Populus spp.) and willows ( Salix spp.) relative to the rate of tamarisk loss will be critical in determining the long-term effect of this large-scale ecological experiment.

Arizona, Colorado, New Mexico, Utah↗

USGS2010

A Natural History Summary and Survey Protocol for the Southwestern Willow Flycatcher

The Southwestern Willow Flycatcher (Empidonax traillii extimus) has been the subject of substantial research, monitoring, and management activity since it was listed as an endangered species in 1995. When proposed for listing in 1993, relatively little was known about the flycatcher's natural history, and there were only 30 known breeding sites supporting an estimated 111 territories rangewide (Sogge and others, 2003a). Since that time, thousands of presence/absences surveys have been conducted throughout the historical range of the flycatcher, and many studies of its natural history and ecology have been completed. As a result, the ecology of the flycatcher is much better understood than it was just over a decade ago. In addition, we have learned that the current status of the flycatcher is better than originally thought: as of 2007, the population was estimated at approximately 1,300 territories distributed among approximately 280 breeding sites (Durst and others, 2008a). Concern about the Southwestern Willow Flycatcher on a rangewide scale was brought to focus by Unitt (1987), who described declines in flycatcher abundance and distribution throughout the Southwest. E. t. extimus populations declined during the 20th century, primarily because of habitat loss and modification from activities, such as dam construction and operation, groundwater pumping, water diversions, and flood control. In 1991, the U.S. Fish and Wildlife Service (USFWS) designated the Southwestern Willow Flycatcher as a candidate category 1 species (U.S. Fish and Wildlife Service, 1991). In July 1993, the USFWS proposed to list E. t. extimus as an endangered species and to designate critical habitat under the Act (U.S. Fish and Wildlife Service, 1993). A final rule listing E. t. extimus as endangered was published in February 1995 (U.S. Fish and Wildlife Service, 1995); critical habitat was designated in 1997 (U.S. Fish and Wildlife Service, 1997). The USFWS Service released a Recovery Plan for the Southwestern Willow Flycatcher in 2002 (U.S. Fish and Wildlife Service, 2002), and re-designated critical habitat in 2005 (U.S. Fish and Wildlife Service, 2005).

Techniques and Methods↗

USGS2010

Geographic variation in the plumage coloration of willow flycatchers Empidonax traillii

The ability to identify distinct taxonomic groups of birds (species, subspecies, geographic races) can advance ecological research efforts by determining connectivity between the non-breeding and breeding grounds for migrant species, identifying the origin of migrants, and helping to refine boundaries between subspecies or geographic races. Multiple methods are available to identify taxonomic groups (e.g., morphology, genetics), and one that has played an important role for avian taxonomists over the years is plumage coloration. With the advent of electronic devices that can quickly and accurately quantify plumage coloration, the potential of using coloration as an identifier for distinct taxonomic groups, even when differences are subtle, becomes possible. In this study, we evaluated the degree to which plumage coloration differs among the four subspecies of the willow flycatcher Empidonax traillii, evaluated sources of variation, and considered the utility of plumage coloration to assign subspecies membership for individuals of unknown origin. We used a colorimeter to measure plumage coloration of 374 adult willow flycatchers from 29 locations across their breeding range in 2004 and 2005. We found strong statistical differences among the mean plumage coloration values of the four subspecies; however, while individuals tended to group around their respective subspecies' mean color value, the dispersion of individuals around such means overlapped. Mean color values for each breeding site of the three western subspecies clustered together, but the eastern subspecies' color values were dispersed among the other subspecies, rather than distinctly clustered. Additionally, sites along boundaries showed evidence of intergradation and intermediate coloration patterns. We evaluated the predictive power of colorimeter measurements on flycatchers by constructing a canonical discriminant model to predict subspecies origin of migrants passing through the southwestern U.S. Considering only western subspecies, we found that individuals can be assigned with reasonable certainty. Applying the model to migrants sampled along the Colorado River in Mexico and the U.S. suggests different migration patterns for the three western subspecies. We believe that the use of plumage coloration, as measured by electronic devices, can provide a powerful tool to look at ecological questions in a wide range of avian species.

Journal of Avian Biology↗

USGS2008

Using Molecular Genetic Markers to Resolve a Subspecies Boundary: The Northern Boundary of the Southwestern Willow Flycatcher in the Four-Corner States

*Executive Summary* The northern boundary of the endangered Southwestern Willow Flycatcher (Empidonax traillii extimus) is currently approximated as running through southern Colorado and Utah, but the exact placement is uncertain because this subspecies shares a border with the more northern and non-endangered E. t. adastus. To help resolve this issue, we evaluated the geographic distribution of mitochondrial and nuclear DNA by sampling breeding sites across the four-corner states (Arizona, Colorado, New Mexico, and Utah). We found that breeding sites clustered into two major groups generally consistent with the currently designated boundary, with the exception of three sites situated along the current boundary. However, delineating a precise boundary that would separate the two subspecies is made difficult because (1) we found evidence for a region of intergradation along the boundary area, suggesting the boundary is not discreet, and (2) the boundary region is sparsely populated, with too few extant breeding populations to precisely locate a boundary. The boundary region encompasses an area where elevation changes markedly over relatively short distances, with low elevation deserts to the south and more mesic, higher elevation habitats to the north. We hypothesized that latitudinal and elevational differences and their concomitant ecological effects could form an ecological barrier that inhibited gene flow between the subspecies, forming the basis for the subspecies boundary. We modeled changes in geographic patterns of genetic markers as a function of latitude and elevation finding significant support for this relationship. The model was brought into a GIS environment to create multiple subspecies boundaries, with the strength of each predicted boundary evaluated on the basis of how much genetic variation it explained. The candidate boundary that accounted for the most genetic variation was situated generally near the currently recognized subspecies boundary, but should be more biologically meaningful because it incorporates the landscape features that may be driving separation of the subspecies. Even so, we caution that using any narrow boundary line as an indicator of subspecies identity could be misleading because biologically the boundary is a region of intergradation rather than a discrete line. Designating, a boundary ultimately becomes a regulatory and management decision based on how much of the genetic variation unique to a subspecies should be protected. We discuss how the results of this study can help guide this decision process by wildlife policy makers.

Open-File Report↗

USGS2008

Yellow-billed Cuckoo Distribution, Abundance, and Habitat Use Along the Lower Colorado River and Its Tributaries, 2007 Annual Report

This 2007 annual report details the second season of a 2-year study documenting western yellow-billed cuckoo (Coccyzus americanus occidentalis) distribution, abundance, and habitat use throughout the Lower Colorado River Multi-Species Conservation Program boundary area. We conducted cuckoo surveys at 40 sites within 14 areas, between 11 June and 9 September 2007. The 169 surveys across all sites yielded 163 yellow-billed cuckoo detections. Cuckoos were detected at 25 of the 40 sites, primarily at the Bill Williams River National Wildlife Refuge (NWR) study area (n = 139 detections; 85 percent of all detections). Detections declined slightly through the cuckoo breeding season, with most detections occurring in the first and second survey periods (n = 92; 54 percent). We detected breeding activity only at the Bill Williams River NWR, where we confirmed 27 breeding events, including two nesting observations. However, the breeding status of most detected birds was unknown. We used playback broadcast recordings to survey for yellow-billed cuckoos. Compared to simple point counts or surveys, this method increases the number of detections of this secretive, elusive species. It has long been suspected that cuckoos have a fairly low response rate, and that the standard survey method of using broadcast recordings might fail to detect all birds present in an area. In 2007, we found that the majority (84 percent) of cuckoo detections were solicited through broadcast at all study sites. The number of solicited detections was highest during the first survey period and declined as the breeding season progressed, while the number of unsolicited detections (cuckoos heard calling before broadcast was initiated) remained fairly constant through the first, second, and third survey periods. The majority (66 percent) of cuckoo detections, solicited or unsolicited, were aural, 23 percent were both heard and seen, and 11 percent were visual detections only. We also found that 50 percent of all responses by cuckoos were evenly split between the first and second broadcasts at sites with >10 detections, while 45 percent of responses occurred after a single broadcast at the sites with <10 detections. We refined our collection of vegetation data in 2007 and found that across the entire study area the dominant tree species were tamarisk (Tamarix spp.), willow (Salix spp.), and cottonwood (Populus spp.). The smallest size class (<8 cm diameter at breast height) trees were the most common and were dominated by tamarisk, but cottonwood and willows were well represented in the larger size classes. Sites that were occupied by yellow-billed cuckoos generally had higher canopies, denser cover in the upper layers of the canopy, and sparse shrub layers compared to unoccupied sites that consistently had higher densities of woody species. As most occupied sites were within the Bill Williams River NWR and most unoccupied sites were at Grand Canyon National Park/Lake Mead National Recreation Area, vegetation characteristics at these study areas drove the cuckoo distribution patterns we observed in 2007. However, there was a range of habitat conditions in locations that were used by yellow-billed cuckoos across the entire lower Colorado River Multi-Species Conservation Program study area. We measured microclimate variables (temperature, relative humidity, soil moisture) at occupied and unoccupied sites, and found that, across the entire study area, occupied sites were consistently cooler during the day and more humid during the day and night compared to unoccupied sites, but that soil moisture did not differ between occupied and unoccupied sites. While most cuckoo detections occurred at Bill Williams River NWR, with generally cooler and more humid conditions, cuckoos were also detected at study areas that had hotter and dryer microclimate conditions. We did not find any relationship of canopy cover characteristics to temperature or soil moisture, suggesting

Open-File Report↗

USGS2008

Southwestern Willow Flycatcher Breeding Site and Territory Summary - 2007

The Southwestern willow flycatcher (Empidonax traillii extimus; hereafter references to willow flycatcher and flycatcher refer to E.t. extimus, except where specifically noted) is an endangered bird that breeds only in dense riparian habitats in parts of six Southwestern states (Arizona, New Mexico, southern California, extreme southern Nevada, southern Utah, and southwestern Colorado). Since 1993, hundreds of Southwestern willow flycatcher surveys have been conducted each year, and many new flycatcher breeding sites located. This document synthesizes the most current information available on all known Southwestern willow flycatcher breeding sites. This rangewide data synthesis was designed to meet two objectives: (1) identify all known Southwestern willow flycatcher breeding sites and (2) assemble data to estimate population size, location, habitat, and other information for all breeding sites, for as many years as possible, from 1993 through 2007. This report provides data summaries in terms of the number of flycatcher sites and the number of territories. When interpreting and using this information, it must be kept in mind that a 'site' is a geographic location where one or more willow flycatchers establishes a territory. Sites with unpaired territorial males are considered breeding sites, even if no nesting attempts were documented. A site is often a discrete patch of riparian habitat but may also be a cluster of riparian patches; there is no standardized definition for site, and its use varies within and among states. For example, five occupied habitat patches along a 10-km stretch of river might be considered five different sites in one state but only a single site in another state. This lack of standardization makes comparisons based on site numbers problematic. Researchers for this report generally deferred to statewide summary documents or to local managers and researchers when delineating a site for inclusion in the database. However, to avoid inflating the number of sites and to establish more consistent definitions of the term 'site', adjacent and nearby sites from statewide reports were sometimes considered as a single site for the purposes of the rangewide data summary. Any combining or splitting of sites at the rangewide level was done on a case-by-case basis. Because of differences in site definitions, one should not evaluate the relative importance of a geographic region (such as drainage, watershed, or state) simply according to the number of flycatcher sites. A 'territory' is an exclusive defended area within a breeding site. Although detailed monitoring studies have identified unpaired territorial males and polygynous males at some flycatcher breeding sites, for the purposes of this report a territory is equivalent to the exclusive breeding area of a pair of flycatchers. In general, the concept of territory is more similar among states and different investigators than site; thus, it is a more robust unit to use for summaries and comparisons. However, note that the definition of a polygynous territory is not consistent among states; a male polygynously paired with two females would be considered one territory in some states and two territories in other states. For each site, we referred to reports or spoke directly with researchers and managers to gather information such as management entity/agency, location (state, drainage, elevation), gross habitat type (native, exotic, or mixed; dominant tree species), and number of flycatcher territories. Synthesizing the information on more than 200 breeding sites is challenging because annual data-collection and survey-reporting requirements are not standardized rangewide, and the nature and degree of readily available information varies widely from state to state. This is particularly true for areas such as California, where there are many flycatcher sites but surveyors are not required to submit standardized flycatcher survey forms. The lack of c

Open-File Report↗

USGS2008

The Colorado Plateau III: integrating research and resources management for effective conservation

Roughly centered on the Four Corners region of the southwestern United States, the Colorado Plateau covers an area of 130,000 square miles. The relatively high semi-arid province boasts nine national parks, sixteen national monuments, many state parks, and dozens of wilderness areas. With the highest concentration of parklands in North America and unique geological and ecological features, the area is of particular interest to researchers. Derived from the Eighth Biennial Conference of Research on the Colorado Plateau, this third volume in a series of research on the Colorado Plateau expands upon the previous two books. This volume focuses on the integration of science into resource management issues, summarizes what criteria make a successful collaborative effort, outlines land management concerns about drought, provides summaries of current biological, sociological, and archaeological research, and highlights current environmental issues in the Four Corner States of Arizona, New Mexico, Colorado, and Utah. With broad coverage that touches on topics as diverse as historical aspects of pronghorn antelope movement patterns through calculating watershed prescriptions to the role of wind-blown sand in preserving archaeological sites on the Colorado River, this volume stands as a compendium of cuttingedge management-oriented research on the Colorado Plateau. The book also introduces, for the first time, tools that can be used to assist with collaboration efforts among landowners and managers who wish to work together toward preserving resources on the Colorado Plateau and offers a wealth of insights into land management questions for many readers, especially people interested in the natural history, biology, anthropology, wildlife, and cultural management issues of the region.

Utah;Arizona;Colorado;New Mexico↗

USGS2007

Yellow-Billed Cuckoo Distribution, Abundance, and Habitat Use Along the Lower Colorado and Tributaries, 2006 Annual Report

Executive Summary This 2006 annual report details the first season of a 2-year study documenting western yellow-billed cuckoo (Coccyzus americanus occidentalis) distribution, abundance, and habitat use throughout the Lower Colorado River Multi-Species Conservation Plan boundary area. We conducted cuckoo surveys at 55 sites within 17 areas, between 11 June and 13 September. The 243 visits across all sites yielded 180 yellow-billed cuckoo detections. Cuckoos were detected at 27 of the 55 sites, primarily at the Bill Williams River National Wildlife Refuge AZ sites (n = 117 detections) and the Grand Canyon National Park-Lake Mead National Recreation Area AZ delta sites (n = 29 detections). There were also cuckoos at the Gila River-Colorado River Confluence, AZ (n = 9), Overton Wildlife Management, NV area (n = 7), and Limitrophe Division North, AZ (n = 6); however, at these sites the numbers were much lower and very few of these birds were considered to be paired or breeding. The greatest number of detections (n = 79) occurred during the second survey period (3-23 July). In 2006, we confirmed five breeding events, including one nesting observation and sightings of four juveniles; all confirmed breeding was at the Bill Williams River NWR and Grand Canyon NP-Lake Mead NRA delta sites. The breeding status of most of our detections were unknown, however, we observed 17 adult cuckoos carrying nest material or food and 40 cuckoo detections were detected while counter-calling occurred in same area during repeated surveys. We used playback recordings to survey for western yellow-billed cuckoos. Compared to simple point counts or surveys, this method increases the number of detections of this secretive, elusive species. It has long been suspected that cuckoos have a fairly low response rate, and that the standard survey method of using playback recordings may fail to detect all birds present in an area. In 2006, we found that the majority (72%) of cuckoo detections were solicited through playback at all study sites. The number of solicited detections peaked during the first half of July and then declined as the breeding season progressed, while the number of unsolicited detections (cuckoos heard calling before playback was initiated) remained fairly constant. The majority (64%) of cuckoo detections, solicited or unsolicited, were aural; 27 percent were both heard and seen and nine percent were visual detections only. Cuckoos in areas with the largest populations had the highest rate of vocalizations before playback or after the first broadcast. In contrast, more than half the responses at sites with fewer cuckoos (with < 10 detections per site) first occurred after three or more playback recordings. This type of baseline information will be used to help refine the survey protocol for 2007, and to create hypotheses that can serve as the foundation for a full-scale evaluation and optimization of this survey technique. Our preliminary analysis of vegetation data from occupied and unoccupied sites in 2006 focused on general patterns in the distribution and abundance of woody species. The density and composition of woody riparian vegetation varied considerably among the study areas. Much of the variation in tree density was due to the patterns of abundance of trees in the smallest size class (< 8 cm dbh). The dominant tree species at the cuckoo survey sites were cottonwood, willow, and tamarisk. Tamarisk was the most common tree, due to the abundance of small (< 8 cm dbh) individuals. When occupied and unoccupied sites were compared, occupied sites tended to have higher average canopy cover, attributable to higher average cover of the mid and low canopy. The dominant canopy at occupied sites most often consisted of cottonwood or willow trees. In addition, occupied sites in most areas had lower than average total tree density whereas unoccupied sites were denser than average. When densities of trees in different size classes were com

Open-File Report↗

USGS2007

Using a remote sensing/GIS model to predict southwestern Willow Flycatcher breeding habitat along the Rio Grande, New Mexico

Introduction The Southwestern Willow Flycatcher (Empidonax traillii extimus; hereafter SWFL) is a federally endangered bird (USFWS 1995) that breeds in riparian areas in portions of New Mexico, Arizona, southwestern Colorado, extreme southern Utah and Nevada, and southern California (USFWS 2002). Across this range, it uses a variety of plant species as nesting/breeding habitat, but in all cases prefers sites with dense vegetation, high canopy, and proximity to surface water or saturated soils (Sogge and Marshall 2000). As of 2005, the known rangewide breeding population of SWFLs was roughly 1,214 territories, with approximately 393 territories distributed among 36 sites in New Mexico (Durst et al. 2006), primarily along the Rio Grande. One of the key challenges facing the management and conservation of the Southwestern Willow Flycatcher is that riparian areas are dynamic, with individual habitat patches subject to cycles of creation, growth, and loss due to drought, flooding, fire, and other disturbances. Former breeding patches can lose suitability, and new habitat can develop within a matter of only a few years, especially in reservoir drawdown zones. Therefore, measuring and predicting flycatcher habitat - either to discover areas that might support SWFLs, or to identify areas that may develop into appropriate habitat - requires knowledge of recent/current habitat conditions and an understanding of the factors that determine flycatcher use of riparian breeding sites. In the past, much of the determination of whether a riparian site is likely to support breeding flycatchers has been based on qualitative criteria (for example, 'dense vegetation' or 'large patches'). These determinations often require on-the-ground field evaluations by local or regional SWFL experts. While this has proven valuable in locating many of the currently known breeding sites, it is difficult or impossible to apply this approach effectively over large geographic areas (for example, the middle Rio Grande). The SWFL Recovery Plan (USFWS 2002) recognizes the importance of developing new approaches to habitat identification, and recommends the development of drainage-scale, quantitative habitat models. In particular, the plan suggests using models based on remote sensing and Geographic Information System (GIS) technology that can capture the relatively dynamic habitat changes that occur in southwestern riparian systems. In 1999, Arizona Game and Fish Department (AGFD) developed a GIS-based model (Hatten and Paradzick 2003) to identify SWFL breeding habitat from Landsat Thematic Mapper imagery and 30-m resolution digital elevation models (DEMs). The model was developed with presence/absence survey data acquired along the San Pedro and Gila rivers, and from the Salt River and Tonto Creek inlets to Roosevelt Lake in southern Arizona (collectively called the project area). The GIS-based model used a logistic regression equation to divide riparian vegetation into 5 probability classes based upon characteristics of riparian vegetation and floodplain size. This model was tested by predicting SWFL breeding habitat at Alamo Lake, Arizona, located 200 km from the project area (Hatten and Paradzick 2003). The GIS-based model performed as expected by identifying riparian areas with the highest SWFL nest densities, located in the higher probability classes. In 2002, AGFD applied the GIS-based model throughout Arizona, for riparian areas below 1,524 m (5,000 ft) elevation and within 1.6 km of perennial or intermittent waters (Dockens et al. 2004). Overall model accuracy (using probability classes 1-5, with class 5 having the greatest probability of nesting activity) for predicting the location of 2001 nest sites was 96.5 percent; accuracy decreased when fewer probability classes were defined as suitable. Map accuracy, determined from errors of commission, increased in higher probability classes in a fashion similar to errors of omission. Map accuracy, li

Colorado, New Mexico↗

USGS2007

The ecology of the Southwestern Willow Flycatcher in central Arizona - A 10-year synthesis report

BACKGROUND From 1996 to 2005, the U.S. Geological Survey (USGS) conducted a demographic study of the Southwestern Willow Flycatcher (Empidonax traillii extimus) in Arizona in collaboration with the Arizona Game and Fish Department (AGFD). The study was begun the year following the listing of the Southwestern Willow Flycatcher as an endangered species. At the time of the listing, very little was known about the biology and threats to the flycatcher, and one of the main objectives of the study was to gather detailed long-term information on the biology of the flycatcher. This report is organized into eight chapters. Following the introductory chapter, we deal with specific aspects of flycatcher ecology and habitat use in each of six separate chapters. We end with a concluding chapter that synthesizes information into broad topical themes that address key management issues. Each of the core chapters (chapters 2 through 7) conclude with a list of management considerations derived from the findings of the respective chapter.

Open-File Report↗

USGS2007

Southwestern Willow Flycatcher Breeding Site and Territory Summary - 2006

Introduction The Southwestern Willow Flycatcher (Empidonax traillii extimus) is an endangered bird that breeds only in dense riparian habitats in six southwestern states (southern California, extreme southern Nevada, southern Utah, southwestern Colorado, Arizona, and New Mexico). Since 1993, hundreds of Southwestern Willow Flycatcher surveys have been conducted each year, and many new flycatcher breeding sites located. This document synthesizes information on all known Southwestern Willow Flycatcher breeding sites. This rangewide data synthesis was designed to meet these objectives: * identify all known Southwestern Willow Flycatcher breeding sites, and * assemble data on population size, location, habitat, and other information for all breeding sites, for as many years as possible, from 1993 through 2006. This report provides data summaries in terms of the number of flycatcher sites and the number of territories.

Open-File Report↗

USGS2000

Breeding season ecology: Chapter 6 in Status, ecology and conservation of the Southwestern Willow Flycatcher (RMRS-GTR-60)

The willow flycatcher ( Empidonax traillii ) breeds across much of the conterminous United States and in portions of extreme southern Canada. As might be expected in such a wide-ranging species, willow fly- catchers in different portions of the range exhibit differences in appearance, song, and ecological characteristics. The intent of this chapter is to provide information on the breeding-season ecology of the southwestern subspecies, E.t. extimus . However, most ecological studies to date have dealt with other willow flycatcher subspecies. Relatively few studies have been published on E.t. extimus , and much of what is currently known is presented in unpublished literature (e.g., agency and consulting firm reports); these sources are relied upon heavily in this chapter. This chapter does not address habitat characteristics in depth, other than for nest sites (refer to A Survey of Current Breeding Habitats for additional details). Although southwestern willow flycatchers are frequent victims of nest parasitism by brown-headed cowbirds ( Molothrus ater ), this chapter will not address the topic of parasitism and its effect on breeding ecology (refer to The Ecology of Brown-headed Cowbirds and their Effects on Southwestern Willow Flycatchers for details). Readers interested in more details of willow flycatcher biology and ecology are encouraged to read McCabe’s (1991) treatise, which is based on over a decade of willow flycatchers research in Wisconsin and includes comparisons with other populations and subspecies.

General Technical Report↗

USGS1997

Status and breeding ecology of the southwestern willow flycatcher in the Grand Canyon

Ernpidonax trailIll extirnus is one of several recognized subspecies of the Willow Flycatcher (Unitt 1987, Browning 1993), a neotropical migrant that breeds across much of North America. This southwestern race is a riparian obligate, nesting in dense patches of willow ( Salix sp.), willow-cottonwood ( Populus sp.), or other similarly structured habitats. In some areas of the Southwest, it nests in dense stands of tamarisk ( Tamarix sp.). Willow Flycatchers were once widespread and locally common in the Southwest (Unitt 1987) but have declined to the point that E. t. extirnus was listed as an endangered subspecies in 1995 (USFWS 1995).

Western Birds↗