Geology topics

Lisa D. Allen

Publications and source records attributed to Lisa D. Allen.

13 recordsLinked to original sources

USGS2026

Distribution and Abundance of Least Bell’s Vireo ( Vireo bellii pusillus ) and Southwestern Willow Flycatcher ( Empidonax traillii extimus ) at the Sepulveda Dam Basin, Los Angeles County, California—2025 Data Summary

Executive Summary We surveyed for Least Bell’s Vireos ( Vireo bellii pusillus ; vireo) and Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) along Bull Creek, Haskell Creek, and the Los Angeles River (Sepulveda Dam project area) in Los Angeles County, California, in 2025. Four vireo surveys were completed between April 16 and July 2, 2025, and three flycatcher surveys were completed between May 21 and July 2, 2025. We found 11 territorial male vireos, 4 of which were confirmed as paired, and 3 transient vireos. Of the 11 territorial vireos, 6 were detected along the Los Angeles River, 3 along Haskell Creek, and 2 along Bull Creek. Forty-three percent of vireos were detected in habitat characterized as mixed willow riparian, and most vireos were detected in habitat with greater than 50-percent native plant cover. No flycatchers were observed in the Sepulveda Dam project area in 2025.

California↗

USGS2026

Southwestern Willow Flycatcher ( Empidonax traillii extimus ) surveys at the city of Carlsbad Preserve, San Diego County, California—2025 data summary

Executive Summary We surveyed for Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) at five survey areas within the City of Carlsbad Preserve, Carlsbad, California, in 2025. Three flycatcher surveys were completed between May 16 and June 30, 2025. One transient flycatcher was observed at the Lake Calavera survey area in the City of Carlsbad Preserve in 2025.

California↗

USGS2024

Least Bell's Vireos and Southwestern Willow Flycatchers—Breeding activities and habitat use—2023 annual report

Executive Summary We completed four protocol surveys for Least Bell’s Vireos ( Vireo bellii pusillus ; hereinafter vireo) during the breeding season, supplemented by weekly territory monitoring visits between April 6 and July 20 at the San Luis Rey Flood Risk Management Project Area (hereinafter Project Area). We identified a total of 136 territorial male vireos; 121 were confirmed as paired, and 4 were confirmed as single males. For the remaining 11 territories, we were unable to confirm breeding status. In 2023, two transient vireos were detected. The vireo population in the Project Area increased by 2 percent from 2022 to 2023. Populations in southern San Diego County also increased (by 6 percent on the Otay River) or were stable (Salt Creek/Wolf Canyon). In contrast, the vireo population at Marine Corps Base Camp Pendleton (MCBCP) and at Marine Corps Air Station decreased by 2 and 10 percent, respectively. We used an index of treatment (hereinafter Treatment Index) to evaluate the effect of ongoing vegetation clearing on the Project Area vireo population. The Treatment Index measures the cumulative effect of vegetation treatment within a territory by using the percentage area treated weighted by the number of years since treatment. We determined that the Treatment Index for an unoccupied habitat was more than four times higher than that of an occupied habitat, indicating that vireos selected habitats that were less treated in which to settle. We monitored vireo nests at three general site types: (1) within the flood channel where non-native and native vegetation removal has occurred regularly (hereinafter Channel), (2) three sites near the flood channel where limited non-native and native vegetation removal has occurred (hereinafter Off-channel), and (3) three sites that have been actively restored by planting native vegetation (hereinafter Restoration). Nesting activity was monitored in 84 territories, 4 of which were occupied by single males. Overall, 46 percent of completed nests were successful, and nest success did not differ among the three sites. In 2023, we found that territories in the Channel had greater hatching success per egg compared to Off-channel, but there were no other differences with regard to clutch size, hatching, or fledging success among Channel, Off-channel, and Restoration sites. Overall breeding success and productivity were slightly higher in 2023 than in 2022, with pairs fledging an average±standard deviation of 3.1±2.1 young and 79 percent of pairs fledging at least 1 young. To investigate if the cumulative years of treatment had an effect on vireo reproductive effort, we looked at the effects of the Treatment Index on reproductive parameters. Results from generalized linear models indicated that treatment did not have an effect on vireo nesting effort (the number of nest attempts) or the number of vireo fledglings per pair produced in 2023. Similarly, we did not detect an effect of Treatment Index on the daily survival rate (DSR) of nests. Analysis of vegetation data collected at vireo nests from 2006 to 2023 did not reveal an effect of vegetation cover at the nest on DSR. We did find, however, that Channel nests were placed higher in and farther from the edge of the host plant than Off-channel nests. Within sites, we did not detect any differences in vegetation cover between successful and unsuccessful nests. Red/arroyo willow ( Salix laevigata or Salix lasiolepis ) and mule fat ( Baccharis salicifolia ) were the species most commonly selected for nesting by vireos in all three site types. Black willow ( Salix gooddingii ) and sandbar willow ( Salix exigua ) also were commonly used. Vireos used a wider variety of species for nesting in Channel and Off-channel sites (10 and 13 species, respectively) compared to Restoration sites (2 species), although there was limited nesting in Restoration sites in 2023. There were 51 vireos banded before the 2023 breeding season that were resighted and identified at the Project Area in 2023. Two of these vireos were originally banded outside of the Project Area, at the Santa Margarita River on MCBCP. Adult birds of known age ranged from 1 to 7 years old. Between 2006 and 2023, survival of males (66±11 percent) was consistently higher than that of females (60±12 percent). First-year birds from 2006 to 2022 had an average annual survival of 15±5 percent. First-year dispersal in 2023 averaged 20.2±31.3 kilometers (km), with the longest dispersal (76.3 km) by a female that was recaptured at Wolf Canyon, a tributary to Otay River. From 2007 to 2012, most returning first-year vireos returned to the Project Area, whereas from 2014 to 2016, a greater proportion of returning birds dispersed to areas outside of the Project Area. From 2018 to 2022, the trend shifted, and more first-year vireos returned to the Project Area, except for 2022 when only one out of five first-year vireos returned to the Project Area. This trend continued in 2023: 71 percent of all first-year vireos returned to the Project Area, and 29 percent dispersed to areas outside of the Project Area (San Diego River and Wolf Canyon). Most of the returning adult male vireos showed strong between-year fidelity to their previous territories. In 2023, 94 percent of males (34/36) occupied a territory that they had defended in 2022 (within 100 meters [m]). In 2023, 33 percent of females (1/3) detected returned to a territory they occupied in 2022. The average between-year movement for returning adult vireos was 0.2±0.9 km. The amount of treatment at adults’ 2022 territories did not affect the distance adults moved to their 2023 territories. We completed four protocol surveys for the endangered Southwestern Willow Flycatcher ( Empidonax traillii extimus ; hereinafter flycatcher) at the Project Area between May 15 and July 21, 2023. In 2023, four transient Willow Flycatchers were detected in the Project Area. Two transients were detected in Reach 1, one in Reach 3a, and one in Whelan Mitigation. No resident flycatchers were documented in the Project Area in 2023. A total of 46 vegetation transects (516 points) were sampled in the Project Area in 2023. There were 71 percent (368/516) of points located in the Channel, and 22 percent (113/516) were in Upper Pond. The remaining 7 percent (35/516) of points were at the Whelan Restoration site. Foliage cover below 1 m was higher at the Channel points and Upper Pond compared to Whelan Restoration. From 1 to 3 m, foliage cover was similar at all 3 sites; however, above 3 m foliage cover was higher in the Channel compared to the Upper Pond and Whelan Restoration sites. Average canopy height was higher in the Channel (5.6±3.8 m) compared to Upper Pond (4.7±2.7 m) and Whelan Restoration (4.0±2.0 m). From 2006 to 2023, total foliage cover declined from 2 to 3 m and above 6 m in the Channel, in contrast to Upper Pond and Whelan Restoration, where little directional change in vegetation cover has occurred and where vegetation cover has largely recovered to 2006 levels. Within the Channel, the steepest declines occurred between 2009 and 2013 and between 2014 and 2016. Since 2016, we observed an increase in foliage cover, largely herbaceous, between 0 and 2 m within the Channel. Although increases were observed at all height classes after 2016, percentage cover has remained below levels measured before 2009. We sampled vegetation at 45 vireo nests and 45 random plots (territory plots) within territories in the Channel and Upper Pond after the 2023 breeding season. Vireos in the Channel established territories in areas with significantly more cover from 3 to 7 m but less cover below 1 m relative to the available habitat. Within territories, Channel vireos selected nest sites largely at random, but with significantly less foliage cover from 4 to 5 m. Vireos at Upper Pond established territories in areas with significantly more foliage cover below 4 m and from 5 to 6 m relative to available habitat. Within territories, Upper Pond vireos also selected nest sites at random except for a preference for sites with significantly less foliage cover below 1 m.

Open-File Report↗

USGS2023

Least Bell's Vireos and Southwestern Willow Flycatchers at the San Luis Rey Flood Risk Management Project Area in San Diego County, California—Breeding activities and habitat use—2022 annual report

Executive Summary We completed four protocol surveys for Least Bell’s Vireos ( Vireo bellii pusillus ; vireo) during the breeding season, supplemented by weekly territory monitoring visits. We identified a total of 133 territorial male vireos; 114 were confirmed as paired, and 3 were confirmed as single males. For the remaining 16 territories, we were unable to confirm breeding status. Two transient vireos were detected in 2022. The vireo population in the Project Area increased by 9 percent from 2021 to 2022. The vireo population at Marine Corps Base Camp Pendleton also increased (4 percent), whereas the population at Marine Corps Air Station remained relatively stable (decreased from 10 pairs to 9) and the Otay River population decreased by 10 percent (2 territories). We used an index of treatment (Treatment Index) to evaluate the effect of on-going vegetation clearing on the Project Area vireo population. The Treatment Index measures the cumulative effect of vegetation treatment within a territory (since 2005) by using the percentage area treated weighted by the number of years since treatment. We determined that the Treatment Index for unoccupied habitat was more than four times that of occupied habitat, indicating that vireos selected habitat that was less treated in which to settle. We monitored vireo nests at three general site types: (1) within the flood channel where exotic and native vegetation removal has occurred regularly (Channel), (2) three sites near the flood channel where limited exotic and native vegetation removal has occurred (Off-channel), and (3) three sites that have been actively restored by planting native vegetation (Restoration). Nesting activity was monitored in 80 territories, 3 of which were occupied by single males and 1 by a male whose breeding status could not be confirmed. Overall, 38 percent of completed nests were successful and nest success did not differ among the three sites. In 2022, there were no differences with regard to clutch size, hatching, or fledging success among Channel, Off-channel and Restoration sites. Overall breeding success and productivity were slightly higher in 2022 than in 2021, with 72 percent of pairs fledgling at least one young and pairs fledging an average of 2.2±1.7 young. To investigate if the cumulative years of treatment had an effect on vireo reproductive effort, we looked at the effects of the Treatment Index on reproductive parameters. Results from generalized linear models indicated that treatment did not have an effect on vireo nesting effort or the number of vireo fledglings per pair produced in 2022. Similarly, we did not detect an effect of Treatment Index on daily survival rate (DSR) of nests. Analysis of vegetation data collected at vireo nests from 2006 to 2022 did not reveal an effect of vegetation cover at the nest on DSR. We did find, however, that Channel nests were placed higher in the host plant than Off-channel nests. In the Channel and Off-channel sites, successful nests were placed closer to the edge of the host plants than unsuccessful nests. Additionally, successful Off-channel nests were placed lower in the vegetation, in shorter host plants, and closer to the edge of the vegetation clump than unsuccessful nests. Red/arroyo willow ( Salix laevigata or Salix lasiolepis ) were the species most commonly selected for nesting by vireos in all three site types. Black willow ( Salix gooddingii ) and mule fat ( Baccharis salicifolia ) also were commonly used. Vireos used a wider variety of species for nesting in Channel and Off-channel sites (eight and six species, respectively) compared to Restoration sites (two species), although there was limited nesting in Restoration sites in 2022. There were 43 vireos banded before the 2022 breeding season that were resighted and identified at the Project Area in 2022, all of which were originally banded in the Project Area. Adult birds of known age ranged from 1 to 7 years old. A total of 146 vireos were newly banded in 2022. There were 8 adult vireos banded with a unique color combination, and 138 nestlings were banded with a single dark blue numbered federal band on the left leg. Between 2006 and 2022, survivorship of males (66±11 percent) was consistently higher than that of females (59±12 percent). First-year birds from 2006 to 2022 had an average annual survivorship of 15±6 percent. First-year dispersal in 2022 averaged 6.7±7.4 kilometers (km), with the longest dispersal (15.3 km) by a male that was recaptured at Fallbrook Creek, Fallbrook Naval Weapons Station (FNWS). From 2007 to 2011, most returning first-year vireos returned to the Project Area, whereas from 2014 to 2016, the majority of returning birds dispersed to areas outside of the Project Area. From 2018 to 2021, the trend shifted, and more first-year vireos returned to the Project area. In 2022, only one first-year vireo returned to the project area and two dispersed to sites outside the Project Area (upstream to the middle San Luis Rey River and to Fallbrook Creek, FNWS). However, the total number of identified first-year vireos was low and the trend in 2022 will likely shift as additional returning first-year vireos are identified in subsequent years. Most of the returning adult male vireos showed strong between-year site fidelity to their previous territories. Seventy-three percent of males (27/37) occupied a territory in 2022 that they had defended in 2021 (within 100 meters [m]). There were no females (0/4) detected in 2022 that returned to a territory they occupied in 2021; however, 50 percent of females (2/4) detected in 2022 returned to areas adjacent to their previous territories (within 300 m). The average between-year movement for returning adult vireos was 0.3±0.7 km. The amount of treatment at adults’ 2021 territories did not affect the distance adults moved to their 2022 territories. We completed four protocol surveys for the endangered Southwestern Willow Flycatcher ( Empidonax traillii extimus ; flycatcher) at the Project Area between May 16 and July 25, 2022. Four transient Willow Flycatchers were detected in the Project Area in 2022. Two transients were detected in Reach 1, one in Reach 3a, and one in Pilgrim Pond. There were not any resident flycatchers documented in the Project Area in 2022. A total of 46 vegetation transects (528 points) were sampled at the Project Area in 2022. Seventy-one percent (378/528) of points were located in the Channel, and 22 percent (115/528) were in Upper Pond. The remaining 7 percent (35/528) of points were at the Whelan Restoration site. Foliage cover below 2 m was higher at the Channel points compared to Upper Pond and Whelan Restoration, which can be attributed to the dense herbaceous vegetation that grows after mowing. Above 2 m, foliage cover was similar at the Channel and Whelan Restoration sites and was higher than at Upper Pond. Average canopy height was higher in the Channel (5.6±3.4 m) compared to Upper Pond (4.7±2.9 m) and Whelan Restoration (4.6±1.9 m). From 2006 to 2022, total foliage cover declined above 2 m in the Channel, in contrast to Upper Pond and Whelan Restoration, where little directional change in vegetation cover has occurred and where vegetation cover has largely recovered to 2006 levels. Within the Channel, the steepest declines occurred between 2009 and 2013 and between 2014 and 2016. Since 2016, we observed an increase in foliage cover, largely herbaceous, between 0 and 2 m within the Channel. The percent cover remained below levels detected before 2009 for other height classes. We sampled vegetation at 44 vireo nests and 44 random plots (“territory” plots) within territories in the Channel and Upper Pond after the 2022 breeding season. Vireos in the Channel established territories in areas with significantly more cover from 3 to 6 m but less cover below 2 m relative to the available habitat. Within territories, Channel vireos selected nest sites with significantly more foliage cover from 2 to 3 m. Vireos at Upper Pond established territories in areas with significantly more foliage cover from 5 to 6 m and below 1 m relative to available habitat. However, within territories, Upper Pond vireos selected nest sites with significantly less foliage cover from 5 to 6 m and below 1 m. Data either are not available or have limited availability owing to restrictions of the funding entity (U.S. Army Corps of Engineers). Please contact Christopher Chabot, Planning Division, Los Angeles District, U.S. Army Corps of Engineers, for more information.

Califoria↗

USGS2023

Southwestern Willow Flycatcher (Empidonax traillii extimus) surveys at the city of Carlsbad Preserve, San Diego County, California—2022 data summary

Executive Summary We surveyed for Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) at five survey areas within the City of Carlsbad Preserve, Carlsbad, California, in 2022. Three flycatcher surveys were completed between May 18 and June 29, 2022. Territorial or transient flycatchers were not observed at the City of Carlsbad Preserve in 2022.

California↗

USGS2022

Least Bell's Vireos and Southwestern Willow Flycatchers at the San Luis Rey flood risk management project area in San Diego County, California: Breeding activities and habitat use—2021 Annual report

California↗

USGS2022

Distribution and abundance of Least Bell’s Vireos ( Vireo bellii pusillus ) and Southwestern Willow Flycatchers ( Empidonax traillii extimus ) on the Middle San Luis Rey River, San Diego County, southern California—2021 Data summary

Executive Summary We surveyed for Least Bell’s Vireos ( Vireo bellii pusillus ; vireo) and Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2021. Surveys were conducted from April 13 to July 14 (vireo) and from May 18 to July 13 (flycatcher). We found 180 vireo territories, at least 125 of which were occupied by pairs. The vireo population size decreased by 6 percent from 2020 to 2021. In 2021, vireo territories decreased by 14 percent in the portion of the middle San Luis Rey River that burned in 2017 and decreased by 2 percent outside of the burned area. Vireos used six different habitat types in the survey area: (1) willow-cottonwood, (2) mixed willow riparian, (3) riparian scrub, (4) upland scrub, (5) willow-sycamore, and (6) non-native. Forty percent of the vireos were detected in habitat characterized as willow-cottonwood, and 97 percent of the vireos were detected in habitat with greater than 50-percent native plant cover. Of the 14 banded vireos detected in the survey area, 3 were resighted with a full color-band combination. There were 10 other vireos with single (natal) federal bands that were recaptured, identified, and color-banded in 2021. One vireo with a single dark blue federal band, indicating that it was banded as a nestling at the lower San Luis Rey River, could not be recaptured for identification. The 10 natal vireos that were recaptured on the middle San Luis Rey River dispersed from 2.0 to 11.7 kilometers from their natal territories. Banded vireos with a known age ranged from 1 to 7 years old. One resident flycatcher, of undetermined breeding status, was observed in the survey area in 2021. The resident flycatcher (male) was detected in a territory of mixed willow habitat with greater than 50-percent native plant cover. He was detected from May 20 to June 4, 2021, and no evidence of pairing or nesting was observed. The male flycatcher was resighted with a unique color-band combination and had occupied the same territory since 2018.

California↗

USGS2021

Least Bell's Vireos and Southwestern Willow Flycatchers at the San Luis Rey flood risk management project area in San Diego County, California—Breeding activities and habitat use—2020 annual report

Executive Summary Surveys and monitoring for the endangered Least Bell’s Vireo ( Vireo bellii pusillus ; vireo) were done at the San Luis Rey Flood Risk Management Project Area (Project Area) in the city of Oceanside, San Diego County, California, between March 31 and July 20, 2020. We completed four protocol surveys during the breeding season, supplemented by weekly territory monitoring visits. We identified a total of 161 territorial male vireos; 145 were confirmed as paired and 4 were confirmed as single males. For the remaining 12 territories, we were unable to confirm pair status. Three transient vireos were detected in 2020. The vireo population in the Project Area increased by 26 percent from 2019 to 2020. Vireo populations increased across San Diego County, with a 39-percent increase documented at Marine Corps Base Camp Pendleton (MCBCP); a 58-percent increase at Marine Corps Air Station; a 78-percent increase on the Otay River; and a 7-percent increase in the population on the middle San Luis Rey River. We used an index of treatment (Treatment Index) to evaluate the impact of on-going vegetation clearing on the Project Area vireo population. The Treatment Index measures the cumulative effect of vegetation treatment within a territory (since 2005) by using the percent area treated weighted by the number of years since treatment. We found that the Treatment Index for unoccupied habitat was more than five times that of occupied habitat, indicating that vireos selected less disturbed habitat in which to settle. We monitored vireo nests at three general site types: (1) within the flood channel where exotic and native vegetation removal has occurred regularly (Channel), (2) three sites next to the flood channel where limited exotic and native vegetation removal has occurred (Off-channel), and (3) three sites that have been actively restored by planting native vegetation (Restoration). Nesting activity was monitored in 100 territories, 4 of which were occupied by single males. Hatching success was higher in the Channel relative to the Off-channel. We found no other differences between Channel, Off-channel, and Restoration nests in terms of clutch size or fledging success. There also was no difference in measures of productivity per pair between Channel, Off-channel, Restoration, and Mixed territories (territories that were classified as one site type but nesting occurred in another site type, or where multiple site types were used for nesting). Overall, breeding success and productivity were lower in 2020 than in 2019, with 69 percent of pairs fledgling at least one young and pairs fledging an average of 2.1±1.7 young. To investigate whether the cumulative years of treatment had an impact on vireo reproductive effort, we looked at the effects of the Treatment Index on reproductive parameters. Results from generalized linear models indicated that treatment did not have an effect on vireo nesting effort or the number of vireo fledglings per pair produced in 2020. Similarly, our analysis of nest survival for 2020 revealed no effect of Treatment Index on daily survival rate. Analysis of vegetation data collected at vireo nests from 2006 to 2020 revealed that vegetation at 1–2 meters (m) from the ground was the most important predictor of daily survival rate. There were differences in nest-placement characteristics among site types and successful/unsuccessful nests. Channel nests were placed higher in the vegetation than Off-channel or Restoration nests. Host plant height, distance to edge of host plant, and distance to edge of vegetation clump were greater at Channel sites compared with Off-channel sites, but were not different from Restoration sites. Within sites, we found only one difference between successful and unsuccessful nests. At Off-channel sites, successful nests were placed higher in the vegetation than unsuccessful nests. Red/arroyo willow ( Salix laevigata or Salix lasiolepis ) and mule fat ( Baccharis salicifolia ) were the species most commonly selected for nesting by vireos in all 3 site types. Vireos used a wider variety of species for nesting in Channel and Off-channel sites (7 and 10 species, respectively) compared to Restoration sites (3 species). Ninety-three vireos banded before the 2020 breeding season were resighted and identified at the Project Area in 2020, all of which were originally banded at the Project Area. Adult birds of known age ranged from 1 to 9 years old. A total of 171 vireos were newly banded in 2020. Twenty-eight adult vireos were banded with a unique color combination, and 143 nestlings were banded with a single dark blue numbered federal band on the left leg. Between 2006 and 2020, survivorship of males (67±10 percent) was consistently higher than females (59±11 percent). First-year birds from 2006 to 2020 had an average over-winter survivorship of 17±5 percent. First-year dispersal in 2020 averaged 2.9±2.9 kilometers (km), with the longest dispersal (13.5 km) by a female that was recaptured at Las Flores Creek, MCBCP. From 2007 to 2012, most returning first-year vireos returned to the Project Area, whereas from 2013 to 2017, the majority of returning birds dispersed to areas outside of the Project Area. In 2018, the trend shifted, and most first-year vireos returned to the Project area. This trend continued in 2020 with most first-year vireos returning to the Project Area; 77 percent of all re-encountered first-year birds returned to the Project Area and 23 percent dispersed to areas outside of the Project Area (upstream to the middle San Luis Rey River and to drainages on MCBCP). Most of the returning adult male vireos showed strong between-year site fidelity to their previous territories. Eighty percent of males (45/56) occupied a territory in 2020 that they had defended in 2019 (within 100 m). Thirty-three percent of females (2/6) detected in 2020 returned to a territory that they occupied in 2019. The average between-year movement for returning adult vireos was 0.1±0.5 km. We completed four protocol surveys for the endangered Southwestern Willow Fycatcher ( Empidonax traillii extimus ; flycatcher) at the Project Area between May 20 and July 20, 2020. No Willow Flycatchers were detected in the Project Area in 2020. A total of 46 vegetation transects (526 points) were sampled at the San Luis Rey Flood Risk Management Project Area in 2020. Seventy-one percent (376/526) of points were in the Channel and 22 percent (115/526) were at Upper Pond. The remaining 7 percent (35/526) were at the Whelan Restoration site. Foliage cover below 1 m was higher at the Channel points compared to Upper Pond and Whelan Restoration. Higher foliage cover in the Channel was attributed to the higher herbaceous component. However, foliage cover from 1 to 3 m was higher at the Whelan Restoration site compared to both Upper Pond and the Channel. Average canopy height was similar at all three site types and was 4.4 m or less. From 2006 to 2020, total foliage cover declined above 1 m in the Channel, from 4 to 5 m at Upper Pond, and above 8 m at Whelan Restoration. Within the Channel, the steepest declines occurred between 2009 and 2013 and between 2014 and 2016. Since 2016, we observed an increase in percent foliage between 0 and 2 m within the Channel, but for other height classes, percent cover remained below levels detected before 2009. Changes in cover at Upper Pond and Whelan Restoration appeared to be driven by the loss of tall tree cover. The vegetation mowing and treatment activities, in combination with lack of precipitation (especially between 2012 and 2016), may have contributed to the decline in foliage cover observed from 2006 to 2020. We sampled vegetation at 49 vireo nests and 49 random plots (“territory” plots) within territories in the Channel and Upper Pond following the 2020 breeding season. Vireos in the Channel selected territories with significantly more foliage cover above 2 m but less cover below 1 m relative to the available habitat. In contrast, Channel vireos selected nest sites within their territories with lower foliage cover above 3 m and were non-selective with regard to cover below 2 m. Vireos at Upper Pond generally were less selective with regard to territory and nest sites but tended to select territories with more foliage cover from 1 to 2 m and above 8 m, and they selected nest sites within their territories with greater foliage cover from 0 to 1 m.

California↗

USGS2021

Distribution and abundance of Least Bell's Vireos and Southwestern Willow Flycatchers on the middle San Luis Rey River, San Diego County, southern California—2020 data summary

We surveyed for Least Bell’s Vireos ( Vireo bellii pusillus ; vireo) and Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2020. Surveys were conducted from April 13 to July 13 (vireo) and from May 14 to July 13 (flycatcher). We found 192 vireo territories, at least 150 of which were occupied by pairs. Vireo territories increased by 40 percent from 2019 to 2020 in the portion of the middle San Luis Rey River that burned as a result of a wildfire in 2017. In contrast, vireo territories decreased by 5 percent from 2019 to 2020 in the unburned portion of the middle San Luis Rey River. Vireos used six different habitat types in the survey area: (1) willow-cottonwood, (2) mixed willow riparian, (3) riparian scrub, (4) upland scrub, (5) willow-sycamore, and (6) non-native. Forty-nine percent of the vireos were detected in habitat characterized as willow-cottonwood, and 93 percent of the vireos were detected in habitat with greater than 50-percent native plant cover. Of the 17 banded vireos detected in the survey area, 6 were resighted with a full color-band combination. Two other vireos with single (natal) federal bands were recaptured, identified, and color-banded in 2020. Eight vireos with a single dark blue federal band, indicating that they were banded as nestlings on the lower San Luis Rey River (LSLR), could not be recaptured for identification. One vireo with a single gold federal band, indicating that it was banded as a nestling at Marine Corps Base Camp Pendleton (MCBCP), could not be recaptured for identification. The two natal vireos that were recaptured on the middle San Luis Rey River dispersed from 2.6 to 6.2 kilometers (km) from their natal territories. Banded vireos with a known age ranged from 1 to 8 years old. One resident flycatcher was observed in the survey area in 2020. The resident flycatcher (male) was detected in a territory of mixed willow habitat with greater than 50-percent native plant cover. He was detected as a single male from May 27 to July 2, 2020, and no evidence of pairing or nesting was observed. The male flycatcher was resighted with a unique color-band combination and had occupied the same territory since 2018.

California↗

USGS2020

Distribution and abundance of Least Bell’s Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) on the Middle San Luis Rey River, San Diego County, southern California—2019 data summary

We surveyed for Least Bell’s Vireos ( Vireo bellii pusillus ; vireo) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2019, and we surveyed and conducted nest monitoring for Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) in a survey area where breeding had historically been documented on the middle San Luis Rey River, in 2019. Surveys were conducted from April 11 to June 24 (vireo) and from May 16 to July 15 (flycatcher). We found 179 vireo territories, at least 124 of which were occupied by pairs. Vireo territories increased by 100 percent within the portion of the middle San Luis Rey River that burned as a result of a wildfire in 2017. In contrast, vireo territories increased by 5 percent within the unburned portion of the middle San Luis Rey River. Vireos used five different habitat types in the survey area: mixed willow riparian, willow-cottonwood, riparian scrub, willow-sycamore, and upland scrub. Fifty-two percent of the vireos were detected in habitat characterized as mixed willow, and 92 percent of the vireos were detected in habitat with greater than 50 percent native plant cover. Of the 12 banded vireos detected in the survey area, 5 were resighted with a full color-band combination. One adult female with a unique color-band combination immigrated to the middle San Luis Rey River from Marine Corps Base Camp Pendleton (MCBCP). Five other vireos with single (natal) federal bands were recaptured, identified, and color banded in 2019. Two vireos with a single dark blue federal band, indicating that they were banded as nestlings on the lower San Luis Rey River (LSLR), could not be recaptured for identification. The five natal vireos that were recaptured on the middle San Luis Rey River dispersed from 1.4 to 8.3 kilometers (km) from their natal territories. Banded vireos with a known age ranged from 1 to 11 years old. One resident flycatcher was observed in the survey area in 2019. The resident flycatcher (male) was detected in a territory of mixed willow habitat with greater than 50 percent native plant cover. He was detected as a single male from May 16 to July 17, 2019, and no evidence of pairing or nesting was observed. The male flycatcher with a unique color-band combination occupied the same territory in 2018 and 2019.

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USGS2019

Distribution and abundance of Least Bell’s Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) on the Middle San Luis Rey River, San Diego County, Southern California—2018 data summary

We surveyed for Least Bell’s Vireos ( Vireo bellii pusillus ; vireo) and Southwestern Willow Flycatchers ( Empidonax traillii extimus ; flycatcher) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2018. Surveys were conducted from April 17 to July 16 (vireo) and from May 16 to July 27 (flycatcher). We found 148 vireo territories, at least 90 of which were occupied by pairs. Six additional transient vireos were detected. Vireos used six different habitat types in the survey area: mixed willow, willow-cottonwood, riparian scrub, willow-sycamore, upland scrub, and non-native habitat. Forty-one percent of the vireos were detected in habitat characterized as mixed willow, and 97 percent of the vireos were detected in habitat with greater than 50 percent native plant cover. Of 10 banded vireos detected in the survey area, 5 had been given full color-band combinations prior to 2018. Four other vireos with single (natal) federal bands were recaptured, identified, and color banded in 2018. One vireo with a single dark blue federal band, indicating that it was banded as a nestling on the lower San Luis Rey River, could not be recaptured for identification. One resident flycatcher and eight transient flycatchers of unknown subspecies were observed in the survey area in 2018. The resident flycatcher (male) was detected in a territory of mixed willow habitat with greater than 95 percent native plant cover. He was detected as a single male from May 24 to July 17, 2018, and no evidence of pairing or nesting was observed. The male flycatcher, detected with a single natal band, was recaptured, identified, and given a unique color combination in 2018. The male flycatcher was originally banded as a nestling on the middle San Luis Rey River in 2016. The eight transient flycatchers were detected from May 25 to June 8, 2018, in mixed willow riparian, willow-cottonwood, and riparian scrub habitat with greater than 95 percent native plant cover.

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USGS2018

Distribution and abundance of Least Bell’s Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) on the Middle San Luis Rey River, San Diego County, southern California—2017 data summary

We surveyed for Least Bell’s Vireos (LBVI) ( Vireo bellii pusillus ) and Southwestern Willow Flycatchers (SWFL) ( Empidonax traillii extimus ) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2017. Surveys were conducted from April 13 to July 11 (LBVI) and from May 16 to July 28 (SWFL). We found 146 LBVI territories, at least 107 of which were occupied by pairs. Five additional transient LBVIs were detected. LBVIs used five different habitat types in the survey area: mixed willow, willow-cottonwood, willow-sycamore, riparian scrub, and upland scrub. Forty-four percent of the LBVIs occurred in habitat characterized as mixed willow and 89 percent of the LBVI territories occurred in areas with greater than 50 percent native plant cover. Of 16 banded LBVIs detected in the survey area, 8 had been given full color-band combinations prior to 2017. Four other LBVIs with single (natal) federal bands were recaptured and banded in 2017. Three LBVIs with single dark blue federal bands indicating that they were banded as nestlings on the lower San Luis Rey River and one LBVI with a single gold federal band indicating that it was banded as a nestling on Marine Corps Base Camp Pendleton (MCBCP) could not be recaptured for identification. One banded LBVI emigrated from the middle San Luis Rey River to the lower San Luis Rey River in 2017. One resident SWFL territory and one transient Willow Flycatcher of unknown subspecies (WIFL) were observed in the survey area in 2017. The resident SWFL territory, which was comprised of mixed willow habitat (5–50 percent native plant cover), was occupied by a single male from May 22 to June 21, 2017. No evidence of pairing or nesting activity was observed. The SWFL male was banded with a full color-combination indicating that he was originally banded as a nestling on the middle San Luis Rey River in 2014 and successfully bred in the survey area in 2016. The male SWFL left the middle San Luis Rey River after June 21, 2017 and subsequently was detected on the San Dieguito River on June 26, 2017, by USGS biologists. The transient WIFL was detected on May 30, 2017, in mixed willow habitat comprised of 50–95 percent of native plant cover.

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USGS2017

Distribution and abundance of Least Bell’s Vireos (Vireo bellii pusillus) and Southwestern Willow Flycatchers (Empidonax traillii extimus) on the Middle San Luis Rey River, San Diego, southern California—2016 data summary

Executive Summary We surveyed for Least Bell’s Vireos (LBVI) ( Vireo bellii pusillus ) and Southwestern Willow Flycatchers (SWFL) ( Empidonax traillii extimus ) along the San Luis Rey River, between College Boulevard in Oceanside and Interstate 15 in Fallbrook, California (middle San Luis Rey River), in 2016. Surveys were done from March 30 to July 11 (LBVI) and from May 18 to July 30 (SWFL). We found 142 LBVI territories, at least 106 of which were occupied by pairs. Six additional transient LBVIs were detected. Of 20 banded LBVIs detected in the survey area, 9 had been given full color-band combinations prior to 2016, although we were unable to determine the exact color combination of 1 female LBVI. Seven other LBVIs with single (natal) federal bands were recaptured and banded in 2016. Four vireos with single dark blue federal bands indicating that they were banded as nestlings on the lower San Luis Rey River could not be recaptured for identification. Three SFWL territories were observed in the survey area in 2016. Two territories were occupied by pairs and one by a male of unknown breeding status. Both pairs attempted to nest at least once, and both pairs were successful, fledging three young each. Nesting began in early June and continued into July. Brown-Headed Cowbird ( Molothrus ater ) eggs were not observed in either nest. An additional 12 transient Willow Flycatchers of unknown subspecies were detected in 2016. Two of the five resident SWFLs were originally banded as nestlings on Marine Corps Base Camp Pendleton. One male and one female were banded as nestlings on Camp Pendleton in 2009 and 2011, respectively. One natal male of unknown breeding status, originally banded as a nestling on the middle San Luis Rey River in 2015, was recaptured and given a unique color combination in 2016. This male was later detected on Marine Corps Base Camp Pendleton.

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