Geology topics

Kelly Proffitt

Publications and source records attributed to Kelly Proffitt.

15 recordsLinked to original sources

USGS2023

Wherever I may roam—Human activity alters movements of red deer (Cervus elaphus) and elk (Cervus canadensis) across two continents

Human activity and associated landscape modifications alter the movements of animals with consequences for populations and ecosystems worldwide. Species performing long-distance movements are thought to be particularly sensitive to human impact. Despite the increasing anthropogenic pressure, it remains challenging to understand and predict animals' responses to human activity. Here we address this knowledge gap using 1206 Global Positioning System movement trajectories of 815 individuals from 14 red deer ( Cervus elaphus ) and 14 elk ( Cervus canadensis ) populations spanning wide environmental gradients, namely the latitudinal range from the Alps to Scandinavia in Europe, and the Greater Yellowstone Ecosystem in North America. We measured individual-level movements relative to the environmental context, or movement expression, using the standardized metric Intensity of Use, reflecting both the directionality and extent of movements. We expected movement expression to be affected by resource (Normalized Difference Vegetation Index, NDVI) predictability and topography, but those factors to be superseded by human impact. Red deer and elk movement expression varied along a continuum, from highly segmented trajectories over relatively small areas (high intensity of use), to directed transitions through restricted corridors (low intensity of use). Human activity (Human Footprint Index, HFI) was the strongest driver of movement expression, with a steep increase in Intensity of Use as HFI increased, but only until a threshold was reached. After exceeding this level of impact, the Intensity of Use remained unchanged. These results indicate the overall sensitivity of Cervus movement expression to human activity and suggest a limitation of plastic responses under high human pressure, despite the species also occurring in human-dominated landscapes. Our work represents the first comparison of metric-based movement expression across widely distributed populations of a deer genus, contributing to the understanding and prediction of animals' responses to human activity.

Idaho, Montana, Wyoming↗

USGS2023

Diverse migratory portfolios drive inter-annual switching behavior of elk across the Greater Yellowstone Ecosystem

A growing body of evidence shows that some ungulates alternate between migratory and nonmigratory behaviors over time. Yet it remains unclear whether such short-term behavioral changes can help explain reported declines in ungulate migration worldwide, as opposed to long-term demographic changes. Furthermore, advances in tracking technology reveal that a simple distinction between migration and nonmigration may not sufficiently describe all individual behaviors. To better understand the dynamics and drivers of ungulate switching behavior, we investigated 14 years of movement data from 361 elk in 20 herds across the Greater Yellowstone Ecosystem (GYE). First, we categorized yearly individual behaviors using a clustering algorithm that identified similar migratory tactics across a continuum of behaviors. Then, we tested seven hypotheses to explain why some ungulates switch behaviors, and we evaluated how behavioral changes affected the proportions of different behaviors across the system. We identified four distinct behavioral tactics: residents (4.8% of elk-years), short-distance migrants (53.7%), elevational migrants (21.9%) and long-distance migrants (19.6%). Of the 20 herds, 18 were partially migratory, and 5 had all four movement tactics present. We observed switches between migratory tactics in all sets of consecutive years during our study period, with an average of 22.5% of individual elk changing movement tactics from one year to the next. Elk in herds with higher movement tactic diversity were significantly more likely to switch tactics and often responded more effectively to adverse environmental conditions, compared to those in herds with low movement tactic diversity. During our study period, switching increased the prevalence of both short- and long-distance migrants, decreased the prevalence of elevational migrants, and had no effect on the prevalence of residents. Our findings suggest that rather than contributing to the declining migratory behavior found in the GYE, switching behavior may enable greater resiliency to continuously changing environmental and anthropogenic conditions.

Wyoming↗

USGS2023

Multi-level thresholds of residential and agricultural land use for elk avoidance across the Greater Yellowstone Ecosystem

1. Conversion of land for settlements and agriculture is increasing globally and can influence wildlife space use. However, there is limited research to identify the thresholds of land-use change that incur wildlife avoidance and how these thresh-olds might vary across levels of selection. 2. We evaluated multi-level avoidance thresholds of elk Cervus canadensis impacted by residential development and irrigated agriculture across the Greater Yellowstone Ecosystem in Idaho, Montana and Wyoming. Using GPS data from765 elk in 21 herds, we estimated habitat selection in relation to development and agriculture at three levels (home range selection, within home range selection and movement path selection). Next, using individual selection covariates and as-sociated measures of land-use availability, we used functional-response models to evaluate how selection varied based on availability, and in turn, to estimate avoidance thresholds. 3. We found individual and level-specific variation in elk responses to environmental factors. Elk exhibited stronger responses (either selection or avoidance) when selecting home range locations (i.e. second-order selection) than when selecting areas within home ranges (i.e. third-order selection) or selecting movement paths (i.e. fourth-order selection). Importantly, elk avoidance of development and agriculture changed as the amount of land in these categories changed. Across all levels of selection elk exhibited neutral selection for human development at low levels of availability (<1.1%–2.2% developed) but avoided areas that were >1.1%–2.2% developed. Conversely, elk selected positively for irrigated agriculture at low to moderate levels of availability (<52.0%–66.2% agriculture) but exhibited neutral selection in areas that were >52.0%–66.2% agriculture. 4. Synthesis and applications . Elk avoidance of low levels of human development suggests conservation efforts such as restrictions on future development or conservation easements could focus on areas that are still below 2% developed. Additionally, because elk selection was strongest at the landscape scale, conservation actions that are based on information about the overall landscape structure may be most impactful. Our results highlight the importance of under-standing variability in wildlife habitat selection at multiple levels, particularly in relation to land-use change, and highlight how functional response modelling can help inform landscape conservation.

Montana, Wyoming↗

USGS2022

Ungulate migrations of the western United States, volume 3

Ungulates (hooved mammals) have a broad distribution across the western United States and play an important role in maintaining predator-prey dynamics, affecting vegetation communities, and providing economic benefits to regional communities through tourism and hunting. Throughout the diverse landscapes they occupy, many ungulate populations undertake seasonal migrations to exploit spatially and temporally variable resources and to avoid predation or other threats. As the human footprint continues to expand across the western United States, ungulates increasingly face more obstacles on their migratory journeys. These obstacles threaten the long-term persistence of existing migrations. As a result, wildlife management agencies across the western United States have worked to identify and protect (or enhance) ungulate migration corridors and seasonal ranges identified from global positioning system (GPS) collar data. These efforts garnered additional support through the U.S. Department of the Interior Secretarial Order (SO) 3362, which was initiated in 2018 and provided Federal support for enhancing habitat quality of big-game winter ranges and migration corridors across the western states. Further, SO 3362 prompted the U.S. Geological Survey (USGS) to establish the Corridor Mapping Team (CMT): a collaboration between USGS and participating State and Federal wildlife management agencies, as well as numerous Tribal Nations. The CMT works collaboratively to map ungulate migrations and seasonal ranges throughout the western United States within the Ungulate Migrations of the Western United States report series. Volume 1 of the series was published in 2020 and contained migrations and winter ranges from 42 herds across 5 states. Volume 2 was published in 2022 and contained migrations and seasonal ranges from an additional 65 herds. This report, Volume 3 in the series, details migrations and seasonal ranges from an additional 45 herds throughout most western states. In aggregate, the report series has detailed and mapped the migrations and seasonal ranges of 152 ungulate herds and serves as a map-based inventory of the documented ungulate migrations across the western United States. The data layers for most of the herds included in the report series are also available to the public by the USGS. In addition to the included herd maps, this volume provides an overview of the many ways the mapping efforts associated with the CMT are being integrated into local conservation, management, and policy throughout the western United States.

Arizona, California, Idaho, Nevada, New Mexico, Or↗

USGS2022

Wildlife migrations highlight importance of both private lands and protected areas in the Greater Yellowstone Ecosystem

Formally protected areas are an important component of wildlife conservation, but face limitations in their effectiveness for migratory species. Improved stewardship of working lands around protected areas is one solution for conservation planning, but private working lands are vulnerable to development. In the Greater Yellowstone Ecosystem (GYE), ungulates such as elk ( Cervus canadensis ) use both protected areas and private lands throughout their annual migrations. We studied patterns of landownership, protection, and conservation challenges within the ranges of migratory elk in the GYE. We used GPS data from 1088 elk in 26 herds to define herd-level seasonal ranges, and extracted covariates related to landownership and protection, land use, and human infrastructure and development. All elk herds used land encompassing >1 ownership type. Most elk herds (92.3 % of herds, n = 24) used the highest proportion of private land in the winter (mean = 36.2 % private land). Most elk herds' winter ranges contained the highest building densities (mean = 1.24 buildings/km 2 ), fence densities (mean = 1.02 km fence/km 2 ), and cattle grazing (mean = 1.9 cows/km 2 ), compared to migratory and summer ranges. Out of all ranges, 36.5 % of ranges did not have any zoning regulations, indicating the potential for future development. Our results show that elk in the GYE rely on habitat outside of protected areas, and face landscape-scale conservation challenges such as habitat fragmentation from human development, particularly in winter ranges. Future conservation strategies for wildlife in this system need to encompass coordination across both public and private land to ensure migratory connectivity.

Idaho, Montana, Wyoming↗

USGS2021

Elk migration influences the risk of disease spillover in the Greater Yellowstone Ecosystem

Wildlife migrations provide important ecosystem services, but they are declining. Within the Greater Yellowstone Ecosystem (GYE) some elk ( Cervus canadensis ) herds are losing migratory tendencies, which may increase spatiotemporal overlap between elk and livestock (domestic bison [ Bison bison ] and cattle [ Bos taurus ]), potentially exacerbating pathogen transmission risk. We combined disease, movement, demographic, and environmental data from eight elk herds in the GYE to examine the differential risk of brucellosis transmission (through aborted fetuses) from migrant and resident elk to livestock. For both migrants and residents, we found that transmission risk from elk to livestock occurred almost exclusively on private ranchlands as opposed to state or federal grazing allotments. Weather variability affected the estimated distribution of spillover risk from migrant elk to livestock, with a 7‐12% increase in migrant abortions on private ranchlands during years with heavier snowfall. In contrast, weather variability did not affect spillover risk from resident elk. Migrant elk were responsible for the majority (68%) of disease spillover risk to livestock because they occurred in greater numbers than resident elk. On a per‐capita basis, however, our analyses suggested that resident elk disproportionately contributed to spillover risk. In five of seven herds, we estimated that the per‐capita spillover risk was greater from residents than from migrants. Averaged across herds, an individual resident elk was 23% more likely than an individual migrant elk to abort on private ranchlands. Our results demonstrate links between migration behavior, spillover risk, and environmental variability, and highlight the utility of integrating models of pathogen transmission and host movement to generate new insights about the role of migration in disease spillover risk. Further, they add to the accumulating body of evidence across taxa that suggests that migrants and residents should be considered separately during investigations of wildlife disease ecology. Finally, our findings have applied implications for elk and brucellosis in the GYE, and suggest that managers should prioritize actions that maintain spatial separation of elk and livestock on private ranchlands during years when snowpack persists into the risk period.

Wyoming↗

USGS2020

Evaluating wildlife translocations using genomics: A bighorn sheep case study

Wildlife restoration often involves translocation efforts to reintroduce species and supplement small, fragmented populations. We examined the genomic consequences of bighorn sheep ( Ovis canadensis ) translocations and population isolation to enhance understanding of evolutionary processes that affect population genetics and inform future restoration strategies. We conducted a population genomic analysis of 511 bighorn sheep from 17 areas, including native and reintroduced populations that received 0–10 translocations. Using the Illumina High Density Ovine array, we generated datasets of 6,155 to 33,289 single nucleotide polymorphisms and completed clustering, population tree, and kinship analyses. Our analyses determined that natural gene flow did not occur between most populations, including two pairs of native herds that had past connectivity. We synthesized genomic evidence across analyses to evaluate 24 different translocation events and detected eight successful reintroductions (i.e., lack of signal for recolonization from nearby populations) and five successful augmentations (i.e., reproductive success of translocated individuals) based on genetic similarity with the source populations. A single native population founded six of the reintroduced herds, suggesting that environmental conditions did not need to match for populations to persist following reintroduction. Augmentations consisting of 18–57 animals including males and females succeeded, whereas augmentations of two males did not result in a detectable genetic signature. Our results provide insight on genomic distinctiveness of native and reintroduced herds, information on the relative success of reintroduction and augmentation efforts and their associated attributes, and guidance to enhance genetic contribution of augmentations and reintroductions to aid in bighorn sheep restoration.

Montana, Idaho, Wyoming↗

USGS2020

Wave-like patterns of plant phenology determine ungulate movement tactics

Animals exhibit a diversity of movement tactics [1]. Tracking resources that change across space and time is predicted to be a fundamental driver of animal movement [2]. For example, some migratory ungulates (i.e., hooved mammals) closely track the progression of highly nutritious plant green-up, a phenomenon called “green-wave surfing” [3-5]. Yet, general principles describing how the dynamic nature of resources determine movement tactics are lacking [6]. We tested emerging theory that predicts surfing and the existence of migratory behavior will be favored in environments where green-up is fleeting and moves sequentially across large landscapes (i.e., wave-like green-up) [7]. Landscapes exhibiting wave-like patterns of green-up facilitated surfing and explained the existence of migratory behavior across 61 populations of four ungulate species on two continents (n=1,696 individuals). At the species level, foraging benefits were equivalent between tactics, suggesting that each movement tactic is fine tuned to local patterns of plant phenology. For decades, ecologists have sought to understand how animals move to select habitat, commonly defining habitat as a set of static patches [8, 9]. Our findings indicate that animal movement tactics emerge as a function of the flux of resources across space and time, underscoring the need to redefine habitat to include its dynamic attributes. As global habitats continue to be modified by anthropogenic disturbance and climate change [10], our synthesis provides a generalizable framework to understand how animal movement will be influenced by altered patterns of resource phenology.

Alberta, British Columbia, Idaho, Wyoming↗

USGS2019

Conserving transboundary wildlife migrations: Recent insights from the Greater Yellowstone Ecosystem

Animal migrations are ecologically, culturally, and economically important. Ungulate populations in many parts of Africa, Asia, Europe, and the Americas migrate long distances to access seasonally available resources, traversing vast landscapes in large numbers. Yet some migrations are declining, raising concerns among scientists and natural resource managers. We synthesize recent advances in ungulate migration ecology with relevance to management and policy. Using case studies from the Greater Yellowstone Ecosystem (GYE), we show how new tools can be applied to map ungulate migrations and assess threats across multiple seasonal habitats, serving as a conservation roadmap. To help conserve ungulate migrations, we also propose a transboundary science, policy, and management framework that could be adapted beyond the GYE and that encompasses the needs of multiple species. The key elements of this framework consist of more widespread mapping and assessment of migrations, improved federal and state coordination across jurisdictional lines, increased investment in private land conservation, and strong engagement of local stakeholders positioned to sustain conservation activities over the long term.

Idaho, Montana, Wyoming↗

USGS2019

Modeling elk‐to‐livestock transmission risk to predict hotspots of brucellosis spillover

Wildlife reservoirs of infectious disease are a major source of human‐wildlife conflict because of the risk of potential spillover associated with commingling of wildlife and livestock. In the Greater Yellowstone Ecosystem, the presence of brucellosis ( Brucella abortus ) in free‐ranging elk ( Cervus canadensis ) populations is of significant management concern because of the risk of disease transmission from elk to livestock. We identified how spillover risk changes through space and time by developing resource selection functions using telemetry data from 223 female elk to predict the relative probability of female elk occurrence daily during the transmission risk period. We combined these spatiotemporal predictions with elk seroprevalence, demography, and transmission timing data to identify when and where abortions (the primary transmission route of brucellosis) were most likely to occur. Additionally, we integrated our predictions of transmission risk with spatiotemporal data on areas of potential livestock use to estimate the daily risk to livestock. We predicted that approximately half of the transmission risk occurred on areas where livestock may be present (i.e., private property or grazing allotments). Of the transmission risk that occurred in livestock areas, 98% of it was on private ranchlands as opposed to state or federal grazing allotments. Disease prevalence, transmission timing, host abundance, and host distribution were all important factors in determining the potential for spillover risk. Our fine‐resolution (250‐m spatial, 1‐day temporal), large‐scale (17,732 km 2 ) predictions of potential elk‐to‐livestock transmission risk provide wildlife and livestock managers with a useful tool to identify higher risk areas in space and time and proactively focus actions in these areas to separate elk and livestock to reduce spillover risk.

Montana↗

USGS2019

Elk forage and risk tradeoffs during the fall archery season

During late summer and fall, elk ( Cervus canadensis ) need access to adequate nutrition to support physiological requirements for reproduction and overwinter survival. The archery hunting season often occurs during this period and can affect distributions of elk as they seek areas that minimize perceived harvest risk. Areas that confer lower harvest risk may provide relatively low‐value nutrition, resulting in a potential tradeoff between minimizing risk and accessing adequate forage. We used radio‐collar data collected from female elk during late summer and fall (Aug–Oct) and estimated resource selection models to evaluate the extent of this potential risk‐nutrition tradeoff. To evaluate if elk exposed to a greater hunting risk altered selection for forage resources, we assessed the relationship between individuals’ selection coefficients for forage and the proportion of their late‐summer‐fall home range accessible to hunters (our metric of hunting risk). Our results indicate that during the archery season, elk with higher‐risk home ranges selected more strongly for areas farther from motorized routes than elk with lower‐risk home ranges. Regardless of the level of risk, however, elk maintained or increased selection for areas with higher forage quality, suggesting that elk did not compromise access to nutritional resources during the archery season. Elk with higher‐risk home ranges were also exposed to the poorest nutrition and increased their selection for areas with higher forage quality more strongly than elk with lower‐risk home ranges during the hunting season. Elk with lower‐risk home ranges had access to the highest nutrition, which may be due to the availability of concentrated sources of high‐quality forage from irrigated agricultural areas on private lands that restricted hunter access. Resource agencies interested in encouraging elk to remain on public lands during the hunting seasons might consider closing motorized travel during the archery season to increase security on public lands, limiting hunter pressure on public lands, improving forage quality on public lands, and working with private land owners to enhance hunter accessibility and restrict elk access to high‐quality forage resources.

Montana↗

USGS2019

Plasticity in elk migration timing is a response to changing environmental conditions

Migration is an effective behavioral strategy for prolonging access to seasonal resources and may be a resilient strategy for ungulates experiencing changing climatic conditions. In the Greater Yellowstone Ecosystem (GYE), elk are the primary ungulate, with approximately 20,000 individuals migrating to exploit seasonal gradients in forage while also avoiding energetically costly snow conditions. How climate-induced changes in plant phenology and snow accumulation are influencing elk migration timing is unknown. We present the most complete record of elk migration across the GYE, spanning 9 herds and 414 individuals from 2001 to 2017, to evaluate the drivers of migration timing and test for temporal shifts. The timing of elk departure from winter range involved a trade-off between current and anticipated forage conditions, while snow melt governed summer range arrival date. Timing of elk departure from summer range and arrival on winter range were both influenced by snow accumulation and exposure to hunting. At the GYE scale, spring and fall migration timing changed through time, most notably with winter range arrival dates becoming almost 50 days later since 2001. Predicted herd-level changes in migration timing largely agreed with observed GYE-wide changes—except for predicted winter range arrival dates which did not reflect the magnitude of change detected in the elk telemetry data. Snow melt, snow accumulation, and spring green-up dates all changed through time, with different herds experiencing different rates and directions of change. We conclude that elk migration is plastic, is a direct response to environmental cues, and that these environmental cues are not changing in a consistent manner across the GYE. The impacts of changing elk migration timing on predator–prey dynamics, carnivore–livestock conflict, disease ecology, and harvest management across the GYE are likely to be significant and complex.

Idaho, Montana, Wyoming↗

USGS2018

Land management alters traditional nutritional benefits of migration for elk

Ungulates typically migrate to maximize nutritional intake when forage varies seasonally. In western North America, however, increasing numbers of ungulates reside on low-elevation winter range year-round rather than migrating. These residents often occupy irrigated agricultural areas, but it is not known whether the nutrition provided by agricultural land exceeds that gained by migration. We evaluated the nutrition available to a partially migratory population of elk ( Cervus canadensis ) in west-central Montana where some individuals summered on low-elevation agricultural lands, whereas others summered on traditional higher-elevation ranges. We sampled elk forage plants from ground plots and adult female elk movements from global positioning system (GPS) collar locations during late summer for 2 years. We tested for differences in the nutrition provided by 11 vegetative communities commonly available to elk in the Rocky Mountains and the nutrition available in areas used by individuals whose behaviors ranged from residency to migration. We found the nutrition available to elk decreased along the continuum from resident to migratory behavior, contrary to our hypothesis that all behaviors would provide access to equivalent forage quality. The relatively small summer home ranges of migrants suggest other factors (e.g., decreased competition) may compensate for their lower nutritional value, because the size of a home range is often inversely related to its fitness benefits. We found irrigated agriculture provided the highest forage quality in low elevations, but recently burned (1–6 yr prior) dry forests at higher elevations provided forage quality approximately equivalent to that of irrigated agriculture. Excluding elk from irrigated agricultural areas should therefore reduce nutritional incentives for elk to reside at low elevations year-round. Additionally, fire in higher-elevation dry forests may temporarily increase forage quality for elk and improve nutritional benefits of migratory behavior. Our results indicate land management practices can affect nutritionally mediated fitness benefits of differing behaviors in partially migratory populations of ungulates

Montana↗

USGS2018

Estimating the risk of elk-to-livestock brucellosis transmission in Montana

Wildlife reservoirs of infectious disease are a major source of human-wildlife conflict because of the risk of potential spillover associated with commingling of wildlife and livestock. In Montana, the presence of brucellosis (Brucella abortus) in free-ranging elk (Cervus canadensis) populations is of significant management concern because of the risk of disease transmission from elk to livestock. To help mitigate potential conflict, we identified how spillover risk changes through space and time using a combination of elk population, disease, and movement data. We developed resource selection functions using telemetry data from 223 female elk to predict the relative probability of female elk occurrence on a daily basis during the 15 February-30 June transmission risk period. We combined these spatiotemporal predictions with elk seroprevalence, demography, and abortion timing data to identify when and where abortions (the primary transmission route of brucellosis) were most likely to occur. Additionally, we integrated these predictions with spatiotemporal data on livestock distribution to estimate the daily risk of livestock encountering brucellosis-induced elk abortions. We estimated that a minimum of ~17,500 adult female elk lived within our study area, which resulted in a conservative estimate of ~525 brucellosis-induced abortions each year. We predicted that approximately half of the transmission events occurred on livestock properties and 98% of those properties were private ranchlands as opposed to state or federal grazing allotments. Our fine-resolution (250-m spatial, 1-day temporal), large-scale (17,732 km2) predictions of potential elk-to-livestock transmission risk provide wildlife and livestock managers with a useful tool to identify higher risk areas in space and time and proactively focus actions in these areas to separate elk and livestock to reduce spillover risk.

Montana↗

USGS2013

Relative influence of human harvest, carnivores, and weather on adult female elk survival across western North America

Well-informed management of harvested species requires understanding how changing ecological conditions affect demography and population dynamics, information that is lacking for many species. We have limited understanding of the relative influence of carnivores, harvest, weather and forage availability on elk Cervus elaphus demography, despite the ecological and economic importance of this species. We assessed adult female survival, a key vital rate for population dynamics, from 2746 radio-collared elk in 45 populations across western North America that experience wide variation in carnivore assemblage, harvest, weather and habitat conditions. Proportional hazard analysis revealed that 'baseline' (i.e. not related to human factors) mortality was higher with very high winter precipitation, particularly in populations sympatric with wolves Canis lupus. Mortality may increase via nutritional stress and heightened vulnerability to predation in snowy winters. Baseline mortality was unrelated to puma Puma concolor presence, forest cover or summer forage productivity. Cause-specific mortality analyses showed that wolves and all carnivore species combined had additive effects on baseline elk mortality, but only reduced survival by <2%. When human factors were included, ‘total’ adult mortality was solely related to harvest; the influence of native carnivores was compensatory. Annual total mortality rates were lowest in populations sympatric with both pumas and wolves because managers reduced female harvest in areas with abundant or diverse carnivores. Mortality from native carnivores peaked in late winter and early spring, while harvest-induced mortality peaked in autumn. The strong peak in harvest-induced mortality during the autumn hunting season decreased as the number of native carnivore species increased. Synthesis and applications. Elevated baseline adult female elk mortality from wolves in years with high winter precipitation could affect elk abundance as winters across the western US become drier and wolves recolonize portions of the region. In the absence of human harvest, wolves had additive, although limited, effects on mortality. However, human harvest, and its apparent use by managers to offset predation, primarily controls overall variation in adult female mortality. Altering harvest quotas is thus a strong tool for offsetting impacts of carnivore recolonization and shifting weather patterns on elk across western North America.

Colorado;Idaho;Montana;Utah;Washington;Wyoming↗