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Joshua D. Stafford

Publications and source records attributed to Joshua D. Stafford.

At least 19 recordsLinked to original sources

Engaging adult R3 outdoor educational program design to strengthen participation in hunting and fishing

Declining participation in hunting and fishing has been slowed by women who are learning numerous outdoor skills through recruitment, retention, and reactivation [R3] programs. R3 programs are hosted by state agencies, non-governmental organizations, and businesses that take a variety of approaches to develop participant skills and confidence. To understand how R3 program design influences hunting and fishing participation, we conducted 11 focus group interviews with 72 participants at women’s R3 programs to identify program design components that women appreciate when learning to hunt and fish. Well-designed programs offered customized educational approaches to align with participants’ needs, created a supportive environment to foster a sense of belonging, and streamlined participation through event logistics. Programs that incorporated these design components promoted skill development, boosted self-confidence, and helped participants build a supportive network to independently hunt and fish. Program leaders can apply these findings to bolster participation in hunting and fishing.

Human Dimensions of Wildlife

Control of a dominant predator influences the occurrence of a mesocarnivore of conservation concern

Context Interspecific interactions shape ecological communities, influence community dynamics, and drive co-evolution. Despite their ecological significance, predation and competition remain understudied in plains spotted skunks ( Spilogale interrupta ), a species of conservation concern. Clarifying how predator management influences their occurrence is crucial for effective conservation. Aims We investigated how coyote ( Canis latrans ) management affects the occupancy of plains spotted skunks and whether interspecific interactions with domestic cats ( Felis catus ) and striped skunks ( Mephitis mephitis ) influence plains spotted skunk occurrence. Methods We analysed live-trap data from east-central South Dakota collected in spring of 2021 and 2022. The study area encompassed portions of counties that implemented disparate predator management regimes, including one with systematic annual coyote removal and another without. We used single-species occupancy models to estimate detection and occupancy probabilities for plains spotted skunks, domestic cats, and striped skunks, incorporating environmental factors, including the site-specific predator control regime. We then applied conditional two-species occupancy models to test whether cats and striped skunks influenced plains spotted skunk occurrence. Key results Plains spotted skunks had the lowest occupancy, followed by domestic cats, and striped skunks. Our findings showed significant associations between coyote removal and occupancy probabilities for each mesocarnivore species. Plains spotted skunks had higher occupancy in areas where coyotes were annually removed. Spotted skunk occurrence was not conditional on either domestic cat or striped skunk occurrence. Conclusions In our study system, cats appear to pose less predation risk to spotted skunks than do other predators, reducing the likelihood that cats significantly influence spotted skunk occupancy. Defensive behaviours and use of spatial refugia by plains spotted skunks may further mitigate predation risk. In addition, co-evolutionary pressures may have led to trait adaptations that facilitate the independent co-occurrence of plains spotted skunks and striped skunks. Implications Our findings highlighted the ecological consequences of predator management and the importance of considering predator–prey dynamics in conservation and management planning. Strategies aimed at conserving plains spotted skunks should integrate predator control measures while considering broader mesocarnivore community interactions.

South Dakota

Fine-scale farming features drive resource selection of a small carnivore of conservation concern

Anthropogenic factors are accelerating species extinction, with small mammalian carnivores among the most affected. These species play vital ecological roles, yet their conservation needs are often overlooked. Our study focused on the plains spotted skunk ( Spilogale interrupta (Rafinesque, 1820)), a small carnivore that has experienced population declines. We hypothesized that their resource selection was influenced by factors expected to influence prey availability, protection from predators, and human activity. We tracked 14 plains spotted skunks in east-central South Dakota, USA, over 2 years during spring and summer. Using mixed-effects logistic regression, we identified seasonal habitat associations. In spring, plains spotted skunks selected areas near farming structures and human development, avoiding high wetland density and crop cover. In summer, they continued to select areas near farming structures and low human development, but also high wetland density and pasture, while avoiding hay bales and crop cover. Our first analysis of the species’ resource use in the Great Plains indicates that plains spotted skunks select habitats with permanent small-scale agricultural features and varying levels of human development across seasons. Our findings suggest that species’ persistence in the region may depend on conservation strategies that account for seasonal planning, habitat heterogeneity, and key agricultural structures.

Canadian Journal of Zoology

Module 4: Basic biology of wetland animals

An assortment of animals inhabit wetlands, with many invertebrates, amphibians and reptiles, fishes, birds, and some mammals being wetland specialists. Wetlands provide ample water, food, and cover that benefits resident animals, but periodic high floods, frequent drying, and harsh water qualities (low-oxygen, acidic, saline) can stress wetland animals. Animals that rely on wetlands have developed numerous adaptations to cope with these stresses. The animal biodiversity in wetlands is particularly valued by people, from deep historical and cultural connections, to modern ecotourists and bird watchers who observe it and hunters and anglers who appreciate the consumptive resources they provide. Animals such as alligators and beavers create or physically-modify wetland environments; this activity is called ecosystem engineering and makes these animals among the most important organisms to wetland ecology. Wetlands are high in biodiversity and support rich communities of wetland invertebrates, are primary breeding habitats for most amphibians, and nearly all migratory birds rely on wetlands during some part of their annual cycle. The interactions of animals with plants (herbivory, cutting) and each other (predation) strongly shape wetland environments and affect nutrient cycling. Human impacts on wetlands can negatively affect many animals through habitat loss and degradation, resulting in many wetland amphibians, reptiles, and birds being listed as threatened and endangered species. Thus, wetland conservation is vital to conserve many animal species.

Wetland Science & Practice

Pasture and diurnal temperature are key predictors of regional Plains Spotted Skunk (Spilogale interrupta) distribution

The Plains Spotted Skunk ( Spilogale interrupta ) is a small carnivore native to central North America that has experienced significant population reductions, and there is a lack of information about the species that could inform conservation. Our study aimed to address knowledge gaps about the distribution and habitat associations of the species in South Dakota using species distribution modeling. We used species location data collected from state natural resource managers, trappers, and members of online social media groups dedicated to hunting and wildlife conservation; environmental predictors; and 6 predictive modeling algorithms (i.e., artificial neural networks, artificial classification tree analysis, generalized boosting models, maximum entropy, multivariate adaptive regression splines, and random forests) to develop climate and landcover ensemble distribution models. The most important climate and landcover predictors were mean temperature diurnal range (i.e., average monthly differences between daily high and low temperatures) and proportion of area classified as pasture. Ensemble model concordance identified approximately 31,300 km 2 of potential Plains Spotted Skunk habitat primarily in eastern South Dakota and between the watersheds of the Missouri and James rivers. Our results offer insights that can guide conservation and inform effective management strategies for conserving Plains Spotted Skunk populations in the northern Great Plains. The promotion of low-intensity agricultural practices such as maintaining pastures, farm buildings, fences rows, and the management of woodland encroachment may improve habitat suitability and facilitate the recovery of plains spotted skunks in the region.

South Dakota

Factors influencing autumn–winter movements of midcontinent Mallards and consequences for harvest and habitat management

Annual phenology and distributions of migratory wildlife have been noticeably influenced by climate change, leading to concerns about sustainable populations. Recent studies exploring conditions influencing autumn migration departure have provided conflicting insights regarding factors influencing the movements of Mallards ( Anas platyrhynchos ), a popular game species. We determined factors affecting timing and magnitude of long-distance movements of 97 juvenile Mallards during autumn-winter across the midcontinent of North America marked with implanted transmitters in North and South Dakota, 2018–2019. Factors influencing variation in movement timing, along with direction and magnitudes, depended on type of movement (i.e., regional [25–310 km], initial migration, or subsequent migration movements [>310 km]). Photoperiod influenced probability of initiating all movements, although the effect was most influential for regional movements. Minimum temperature most influenced initial migration events (probability of movement increased 29% for each 1°C decrease); favorable winds also increased likelihood of initial migration events. Probability of subsequent migration events increased 80% for each 1 cm increase in depth of snow. Subsequent migration movements also were 2.0 times more likely to occur on weekend days, indicating disturbance from humans may influence movements. Migration distances increased 166 km for each 1°C reduction in minimum temperature. We also observed markedly different autumn-winter distributions of marked birds between years. Median locations during autumn-winter 2018–2019 were ~250 km farther north and ~300 km farther west during mid-December–January compared to the same time in 2019–2020. Concurrently, harvest rates for marked females and males were 10% and 26% during autumn-winter 2018–2019 and 26% and 31% during autumn-winter 2019–2020. Climate-related changes may result in increasingly variable autumn-winter distributions, with implications for wildlife recreationalists, conservation planners, and harvest managers.

Ecology and Evolution

Evaluation of fall-seeded cover crops for grassland nesting waterfowl in eastern South Dakota

The Prairie Pothole Region (PPR) is the primary breeding ground for many species of North American waterfowl. The PPR was historically dominated by mixed and tallgrass prairies interspersed with wetlands, but >70% of the native grassland area has been lost due to widespread conversion to croplands. Cover cropping is a reemerging farming technique that may provide suitable nesting cover for grassland nesting waterfowl in active croplands, but waterfowl nest survival in fall cover-cropped fields has not been evaluated. We studied use (nest abundance and density) and nest survival of breeding waterfowl in fall-seeded cover crops and perennial cover during 2018 and 2019. We searched 2,094 ha of cover crops and 1,604 ha of perennial cover and found 123 and 304 duck nests, respectively, in each cover type. Estimated nest success (34-day interval) was 3.7% and 16.6% in cover crops during 2018 and 2019, respectively, versus 22.1% in 2018 and 24.9% in 2019 in perennial cover, with increased success of cover-crop fields in 2019 resulting from precipitation that prevented most fields from being planted to row crops. In a model that included effects of planting, daily nest survival in perennial cover was 0.944 (SD = 0.026) in 2018 and 0.960 (SD = 0.019) in 2019. Estimated daily nest survival was 0.912 (SD = 0.040) in 2018 and 0.960 (SD = 0.019) in 2019 during intervals when planting did not occur, but was only 0.417 (SD = 0.124) in 2018 and 0.612 (SD = 0.117) in 2019 on the day that planting occurred. Estimated nest densities in 2018 and 2019, adjusted for nests that failed prior to discovery, were 5.1 (SE = 1.1) and 11.0 (SE = 3.1) nests 100-ha −1 in perennial cover, but only 2.1 (SE = 0.8) and 2.6 (SE = 0.7) in cover crops, respectively. Based on observed nest initiation and planting dates, about 70% of duck nests in cover crops would experience planting events in a typical growing season. Our results suggest that under current management techniques, fall-seeded cover crops offer poor nesting habitat for waterfowl; however, the important benefits cover crops provide to soil health, water quality, and other ecosystem services remain.

South Dakota

Landowner and biologist perceptions of game bird predators and management

Nest survival has been identified as one of the most influential vital rates causing population change in game birds, and depredation, often influenced by habitat loss and fragmentation, is the primary cause of nest failure of upland game birds. We were interested in quantifying and comparing the perspectives of landowners and biologists in South Dakota regarding complex predator-prey interactions to improve communication and management efficacy. We developed a questionnaire regarding the following: 1) general attitude statements about game bird species; 2) perceived impacts of 9 factors (e.g., development, pollution, predators) and 13 potential predators on game bird abundances; and 3) attitude statements regarding use of lethal predator control and nesting habitat management practices. A cluster analysis using landowner attitude statements about predator management identified 3 landowner segments that had strong (most supportive; 37%), moderate (moderately supportive; 35%), or weak (least supportive; 28%) attitude statements about lethal predator control. Landowner segments most supportive and moderately supportive of predator control rated predators as the primary negative factor impacting game bird abundances and agreed that predators were the primary cause of game bird abundance declines, whereas the landowner segment least supportive of predator control rated habitat loss as the top factor and disagreed that predators were the primary cause of game bird declines. Biologists rated habitat loss as the top factor negatively impacting game bird abundances and disagreed that predators were the primary cause of game bird abundance declines. Thus, when considering the effectiveness of strategies to reduce nest depredation, most landowners focused on the direct cause of nest failures (predators), whereas biologists focused on an indirect cause (habitat loss). Perception differences among these groups emphasizes the need for better communication on proximate and ultimate factors affecting game bird populations and how these differences may impact management decisions.

South Dakota

Fish functional gradients along a reservoir cascade

1.The transformations of fish assemblages caused by reservoir cascades can be severe at the reach scale, but basin-scale effects are less clear. However, prevailing river concepts provide a framework for predicting basin-scale effects. 2. To determine if predictions made by the River Continuum Concept relative to the function of fish assemblages are sustained in a temperate river transformed into a reservoir cascade, we examined longitudinal trends in the distribution of fish functional traits over 23 reservoirs of the Tennessee River, USA. 3. In all, 115 species were recorded representing 62 traits, with trait richness increasing longitudinally in a downstream direction. Trophic, reproductive, and habitat traits showed various increasing and decreasing patterns up and down the reservoir cascade. The observed gradients in trait richness and trait distributions were generally consistent with those expected in unregulated rivers, with few unexpected results. 4. The transformation of lotic systems into lentic ones has changed habitats and sources of food and encouraged the proliferation of certain feeding (e.g., detritivores, planktivores, invertivores, piscivores), reproduction (e.g., nest spawners polyphils, broadcast spawners phytolithophils), and habitat (slow current, lacustrine, large river) traits. In essence, reservoirs have expanded downstream habitats in an upstream direction, and thus allowed upstream expansion of species and traits that would have normally not been well represented in upper reaches of the Tennessee River basin. Nevertheless, the impounded Tennessee River has maintained much of its functional integrity, despite extensive alterations to the riverscape. 5. We suggest that while reservoirs have been shown to have major local-scale effects on riverine fish assemblages, with access to riverine habitats, and with proactive conservation strategies, fish functional richness can remain remarkably high at the basin scale.

Alabama, Georgia, Kentucky, North Carolina, South

The first documented interaction between a long-tailed weasel (Mustela frenata) and a plains spotted skunk (Spilogale interrupta) carcass

A novel interaction between a long-tailed weasel ( Mustela frenata ) and a plains spotted skunk ( Spilogale interrupta ) carcass is detailed. In November 2020, a farmer in Edmunds County in north-central South Dakota sent in a video recording of a long-tailed weasel with a spotted skunk carcass. Location of the event, carcass condition, and recorded behavior of the long-tailed weasel offer probable, but unconfirmed, evidence that the spotted skunk was killed by the long-tailed weasel.

South Dakota

The local responses of aquatic ecosystems to adjacent grassland conservation: Can streams of dreams exist in a degraded riverscape?

Landscape homogenization and the removal of riparian areas have altered stream ecosystems worldwide. Numerous conservation programmes attempt to improve water quality and increase instream habitat heterogeneity to elicit desired biological responses. However, the effectiveness of many conservation efforts on isolated stream fragments remains unknown, especially in grassland regions. The effects of grassland conservation practices and the re-establishment of riparian corridors in the James River basin, South Dakota (USA) on stream water quality, habitat availability and aquatic macroinvertebrate and fish assemblages were studied in an agriculturally dominated prairie landscape. Grassland conservation efforts may have repaired riparian condition, reduced turbidity and created more diverse instream habitat complexes at conservation sites based on comparisons with paired reference reaches. Reference sites were relatively homogeneous, with prevalent siltation, bank erosion and disturbances to the riparian vegetation. Owing to significant riparian vegetation development, overhanging and aquatic vegetation, benthic detritus and woody materials were significantly more common at conservation reaches. Restoration efforts that assume ‘if you (re-)build it, they will come’ (i.e. the ‘field of dreams’ hypothesis) underestimate other important barriers to biodiversity restoration in dynamic, grassland riverscapes. Although aquatic organisms in grassland ecosystems are adapted to rapidly inhabit available habitats, the development of niche space at conservation reaches did not directly result in colonization by aquatic life. Grassland management actions did not address stream connectivity issues or overcome land use influences elsewhere in the riverscape that may govern the responses of aquatic macroinvertebrates and fish. Stream fragmentation and continuing, damaging land use patterns seemed to exceed the positive effects of restoring isolated stream reaches in these heavily degraded catchments. Catchment-scale management strategies that combine reach-level restoration actions with efforts to improve connectivity are likely to be more successful in degraded riverscapes.

South Dakota

Extreme climatic variability during migration invokes physiological and dietary plasticity among spring migrating ducks

Environmental stochasticity encountered during migration can have negative consequences for individuals and population demographics through direct reductions in survival or cross-seasonal impacts. We took advantage of substantial interannual variation in spring migration conditions over a 4 year field study to examine physiological and dietary variation among two species of migrant ducks. We collected female Lesser Scaup ( Aythya affinis (Eyton, 1838)) and Blue-winged Teal ( Spatula discors (Linnaeus, 1766)) during spring migration and measured lipid and protein reserves, an index of recent lipid metabolism based on concentrations of lipid metabolites in plasma, and diets. We documented systematic interannual variation among these metrics in both species, contrasting primarily the warmest, earliest spring and the coldest, latest spring. Lesser Scaup had reduced lipid and protein reserves and consumed less energy-rich prey during the coldest and latest spring but showed no interannual variation in the index of lipid metabolism. Blue-winged Teal similarly had reduced protein reserves in the cold, late spring but maintained constant lipid reserves among years, likely facilitated by increased consumption of energy-rich seeds reflected in diets and lipid metabolism. Our results reveal impacts of environmental stochasticity on migrants and suggest that recruitment may be impacted by variable conditions encountered during migration during extreme weather events.

South Dakota

Application strategy for an anthraquinone-based repellent and the protection of soybeans from Canada goose depredation

Agricultural crops can sustain extensive damage caused by Canada geese ( Branta canadensis ) when these crops are planted near wetlands or brood-rearing sites. From 2000 to 2015, South Dakota Game, Fish and Parks spent >$5.6 million to manage damages caused by Canada geese to agricultural crops (primarily soybeans) in South Dakota, USA. For the purpose of developing a repellent application strategy for nonlethal goose damage management, we comparatively evaluated the width of anthraquinone applications (i.e., 9.4 L Flight Control ® Plus goose repellent/ha [active ingredient: 50% 9,10-anthraquinone] at 0–36 m versus 0–73 m perpendicular to the edge of wetlands in 2014), the timing of the first repellent application (i.e., 9.4 L Flight Control Plus goose repellent/ha at 50% versus 75% seedling emergence in 2015), the yield of soybeans ( Glycine max ) within repellent-treated and untreated subplots, and anthraquinone chemical residues in Day County, South Dakota. Soybean yield was greater in subplots 73 m from the water’s edge than that in the 36-m subplots ( P < 0.02). Among subplots first sprayed at 50% seedling emergence, soybean yield was greater at 73 m and 82 m than that at 36 m ( P < 0.005). In contrast, we observed no difference in yield at 36 m, 73 m, or 82 m in the subplots first sprayed at 72% seedling emergence ( P > 0.09). We therefore conclude that goose damages were effectively managed in subplots first sprayed at 72% seedling emergence. Anthraquinone residues averaged 674 and 629 ppm anthraquinone upon the first application of the repellent (June to July), 22 and 35 ppm anthraquinone in the mid-season hay (August to September), and 36 and 28 ppb anthraquinone in the harvested seed (October to November) in 2014 and 2015, respectively. Our results suggest that a 73-m bandwidth of anthraquinone-based repellents first applied at approximately 72% or 65–85% seedling emergence can protect soybeans from Canada goose depredation.

South Dakota

Long-term spatial heterogeneity in mallard distribution in the Prairie pothole region

The Prairie Pothole Region (PPR) of north-central United States and south-central Canada supports greater than half of all breeding mallards ( Anas platyrhynchos ) annually counted in North America and is the focus of widespread conservation and research efforts. Allocation of conservation resources for this socioeconomically important population would benefit from an understanding of the nature of spatiotemporal variation in distribution of breeding mallards throughout the 850,000 km 2 landscape. We used mallard counts from the Waterfowl Breeding Population and Habitat Survey to test for spatial heterogeneity and identify high- and low-abundance regions of breeding mallards over a 50-year time series. We found strong annual spatial heterogeneity in all years: 90% of mallards counted annually were on an average of only 15% of surveyed segments. Using a local indicator of spatial autocorrelation, we found a relatively static distribution of low-count clusters in northern Montana, USA, and southern Alberta, Canada, and a dynamic distribution of high-count clusters throughout the study period. Distribution of high-count clusters shifted southeast from northwestern portions of the PPR in Alberta and western Saskatchewan, Canada, to North and South Dakota, USA, during the latter half of the study period. This spatial redistribution of core mallard breeding populations was likely driven by interactions between environmental variation that created favorable hydrological conditions for wetlands in the eastern PPR and dynamic land-use patterns related to upland cropping practices and government land-retirement programs. Our results highlight an opportunity for prioritizing relatively small regions within the PPR for allocation of wetland and grassland conservation for mallard populations. However, the extensive spatial heterogeneity in core distributions over our study period suggests such spatial prioritization will have to overcome challenges presented by dynamic land-use and climate patterns in the region, and thus merits additional monitoring and empirical research to anticipate future population distribution. Published 2017. This article is a U.S. Government work and is in the public domain in the USA.

Wildlife Society Bulletin

Three responses of wetland conditions to climatic extremes in the Prairie Pothole Region

Wetlands in central North Dakota were revisited after 50 years to assess changes following extreme drought and a prolonged wet period. We compared data collected during 1961–1966 to current (2013–2014) wetland conditions. We revisited 80 wetlands in 2013 and 2014 across three study areas and measured wetland area, ponded-water depth, and specific conductance. Wetlands at the three study areas responded to prolonged wet conditions in one of three ways. Wetlands at Crystal Springs became larger, and had deeper ponds of lower specific conductance in 2013–14 compared to the 1960s. Wetlands at Cottonwood were larger with deeper ponds of slightly higher specific conductance in 2013–2014. Wetlands at Mt. Moriah had only subtle changes in size, pond depth, and specific conductance between periods. Prolonged wet conditions led to merging of most wetlands (defined as the outer edge of wet-meadow vegetation) at Crystal Springs and a few wetlands at Cottonwood. Low topographic relief at Crystal Springs and Cottonwood contributed to storage of excess water in wetlands with associated responses to prolonged wet conditions. In contrast, higher topographic relief and natural outlets into two intermittent streams at Mt. Moriah resulted in wetlands being less impacted by prolonged wet conditions.

North Dakota

Assessing spring direct mortality to avifauna from wind energy facilities in the Dakotas

T he Northern Great Plains (NGP) contains much of the remaining temperate grasslands, an ecosystem that is one of the most converted and least protected in the world. Within the NGP, the Prairie Pothole Region (PPR) provides important habitat for >50% of North America's breeding waterfowl and many species of shorebirds, waterbirds, and grassland songbirds. This region also has high wind energy potential, but the effects of wind energy developments on migratory and resident bird and bat populations in the NGP remains understudied. This is troubling considering >2,200 wind turbines are actively generating power in the region and numerous wind energy projects have been proposed for development in the future. Our objectives were to estimate avian and bat fatality rates for wind turbines situated in cropland- and grassland-dominated landscapes, document species at high risk to direct mortality, and assess the influence of habitat variables on waterfowl mortality at 2 wind farms in the NGP. From 10 March to 7 June 2013&ndash;2014, we completed 2,398 searches around turbines for carcasses at the Tatanka Wind Farm (TAWF) and the Edgeley-Kulm Wind Farm (EKWF) in South Dakota and North Dakota. During spring, we found 92 turbine-related mortalities comprising 33 species and documented a greater diversity of species ( n &thinsp;=&thinsp;30) killed at TAWF (predominately grassland) than at EKWF ( n &thinsp;=&thinsp;9; predominately agricultural fields). After accounting for detection rates, we estimated spring mortality of 1.86 (SE&thinsp;=&thinsp;0.22) deaths/megawatt (MW) at TAWF and 2.55 (SE&thinsp;=&thinsp;0.51) deaths/MW at EKWF. Waterfowl spring (Mar&ndash;Jun) fatality rates were 0.79 (SE&thinsp;=&thinsp;0.11) and 0.91 (SE&thinsp;=&thinsp;0.10) deaths/MW at TAWF and EKWF, respectively. Our results suggest that future wind facility siting decisions consider avoiding grassland habitats and locate turbines in pre-existing fragmented and converted habitat outside of high densities of breeding waterfowl and major migration corridors.

North Dakota, South Dakota

Invertebrates in managed waterfowl marshes

Invertebrates are an important food for breeding, migrating, and wintering waterfowl. Sparse study has been devoted to understanding the influence of waterfowl and wetland management on production of invertebrates for waterfowl foods; however, manipulation of hydrology and soils may change or enhance production. Fish can compete with waterfowl for invertebrate forage in wetlands and harm aquatic macrophytes; biomanipulation (e.g., stocking piscivores) may improve waterfowl habitat quality. Similarly, some terrestrial vertebrates (e.g., beaver ( Castor canadensis )) may positively or negatively impact invertebrate communities in waterfowl habitats. Various challenges exist to wetland management for invertebrates for waterfowl, but the lack of data on factors influencing production may be the most limiting.

Book chapter