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Joan Hagar

Publications and source records attributed to Joan Hagar.

At least 19 recordsLinked to original sources

Response of corvid nest predators to thinning: implications for balancing short- and long-term goals for restoration of forest habitat

Forest thinning on public lands in the Pacific Northwest USA is an important tool for restoring diversity in forest stands with a legacy of simplified structure from decades of intensive management for timber production. A primary application of thinning in young (< 50-year-old) stands is to accelerate forest development to mitigate loss of late-seral habitat to decades of logging. However, thinning may have short-term negative effects for some species associated with mature forest that are expected to benefit from the practice over the long term. An increased risk of nest predation is a primary concern to managers charged with stewardship of habitat for the federally threatened Marbled Murrelet ( Brachyramphus marmoratus ), a species that nests in older forests. Predation by corvids is the greatest cause of nest failure for the Marbled Murrelet, and corvids are known to respond positively to forest disturbance, but quantitative information is lacking on the potential impacts of thinning on risk of nest predation. We investigated the response of two common corvid nest predators, Steller’s Jay ( Cyanocitta stelleri ) and Canada Jay ( Perisoreus canadensis ), to variation in thinning intensity in young forest (< 50 years old) using data from a long-term silviculture experiment. We used a Before-After-Control-Impact (BACI) design, linear mixed modeling, and occupancy modeling to quantify differences in corvid observation rates among varying levels of thinning intensity, and to assess changes in jay response over more than a decade following thinning. We found an increase in observation rates of both species in the heavily thinned treatment during the first 5 to 7 years following thinning, and some evidence of a short-term increase in Steller’s Jay activity in the thinning-with-gaps treatment. Neither jay species responded to the least intensive thinning treatment, which reduced average canopy cover by < 30%. By approximately a decade after thinning, observation rates of jays did not differ between unthinned controls and any of the thinning treatments. Incorporating our quantitative information into landscape-level planning can help managers balance short- and long-term conservation goals.

Oregon

Annual migratory movement, apparent molt-migration, migration schedule, and diffuse migratory connectivity of Hermit Warblers

Quantifying migratory connectivity and annual movement is key to sound conservation planning for migratory species. Hermit Warblers ( Setophaga occidentalis ) are an endemic-breeding species in the Pacific Northwest that winters in Mexico and the Central Americas. This species faces threats from mature forest loss and climate change throughout its range, but we know little about its migration ecology. To understand the annual movements and migratory connectivity of Hermit Warblers, we tracked 22 adult male Hermit Warblers from six breeding sites across the species’ breeding range using geolocators to examine migratory connectivity, spatiotemporal patterns, and migration routes. We found a high degree of mixing on the wintering grounds among birds from different breeding locations, indicating low migratory connectivity. However, birds breeding in Yosemite, the southernmost breeding location in our study, wintered farther east and south than birds from more northern breeding locations, providing weak evidence for potential chain migration. All birds showed much shorter and faster movements during spring migration than during fall migration. Birds arrived at breeding grounds from late April to mid-May and left breeding ranges from late June to mid-July. In fall, birds moved slowly from the breeding locations to montane regions in southern Oregon and California, which may indicate post-breeding molt before swiftly migrating to wintering grounds. Low migratory connectivity in this species implies that habitat and climate change across the broad wintering range may affect breeding populations throughout the species’ breeding range. A particularly compressed breeding schedule and departure of birds from the breeding grounds in early July may indicate that breeding is limited by a short window of favorable climatic conditions for breeding, which could signal heightened vulnerability under future climatic scenarios.

Avian Conservation and Ecology

Level and pattern of overstory retention shape the abundance and long-term dynamics of natural and created snags

Standing dead trees, or snags, serve myriad functions in natural forests, but are often scarce in forests managed for timber production. Variable retention (VR), the retention of live and dead trees through harvest, has been adopted globally as a less intensive form of regeneration harvest. In this study, we explore how two key elements of VR systems — level (amount) and spatial pattern of live-tree retention — affect the carryover and post-harvest dynamics of natural and artificially created snags. We present nearly two decades of data from the DEMO Study, a regional-scale experiment in VR harvests of Douglas-fir-dominated forests in the Pacific Northwest. Snag losses to harvest were greater at 15 than at 40% retention (67 vs. 47% declines in density) and greater in dispersed than in aggregated treatments (64 vs. 50% declines). Densities of hard and tall (≥5 m) snags were particularly sensitive to low-level dispersed retention, declining by 76 and 81%, respectively. Despite these losses, post-harvest densities correlated with pre-harvest densities for most snag size and decay classes. In contrast to initial harvest effects, snag densities changed minimally over the post-harvest period (years 1 to 18 or 19), with low rates of recruitment offsetting low rates of loss. Post-harvest survival of snags was greater at 15 than at 40% retention (79 vs. 69%), as were rates of decay (68 vs. 52% of hard snags transitioned to soft). However, pattern had no effect on either process. Snag recruitment did not vary with retention level or pattern at the scale of the 13-ha harvest unit, but was several-fold greater in the 1-ha aggregates (14.3–27.8 snags ha −1 ) than in the corresponding dispersed treatments (4.2–5.3 snags ha −1 ). Snag size (diameter) distributions showed greater change in dispersed than in aggregated treatments, reflecting greater loss of smaller snags and recruitment biased toward larger snags. Created snags showed uniformly high survival (97%), irrespective of treatment, but rates of decay were greater at lower retention. If a goal of VR is to sustain snag abundance and diversity through harvest, emphasis should be placed on minimizing initial losses, either by reducing the intensity of felling in areas of dispersed retention or locating forest aggregates in areas of greater initial snag density, diversity, or incipient decay.

Forest Ecology and Management

The snag’s the limit: Habitat selection modeling for the western purple martin in a managed forest landscape

The western purple martin ( Progne subis arboricola ), an avian insectivore, is a species of conservation concern throughout the Pacific Northwest. Compared to the well-studied eastern subspecies ( Progne subis subis ), little is known of the life history and biology of the western subspecies. Availability of breeding habitat is believed to be a major limiting factor for western purple martins in forested habitat, but fundamental information on their current distribution and selection of nesting habitat is deficient. To fill this gap, we compared habitat characteristics at three spatial scales (snag-level, stand-level [48.6 ha], landscape-level [314 ha]) surrounding nest snags occupied by purple martins in western Oregon to unoccupied sites. We found habitat for nesting purple martins was defined by the presence of moderately decayed snags with nest cavities, located well away from closed-canopy forest in sufficiently large (>15 ha) open areas. Our modeling efforts suggested suitable habitat was rare within the study region because: 1) snags were scarce on private industrial forest lands and 2) large disturbed patches were uncommon on federal lands. We conclude that a disturbance regime characterized by infrequent but major stand-replacing events, such as fire or timber harvest, is likely the key to maintaining breeding habitat for purple martins in upland forests in western Oregon.

Oregon

Evaluating the role of active management in mature Douglas-fir (Pseudotsuga menziesii) stands for songbird conservation

Forest birds, particularly those associated with late-successional forests, are of widespread conservation interest. Although birds are among the more widely studied taxa of forest wildlife, relatively few studies have examined the long-term effects of active management (i.e., intentional stand density reduction) on the forest bird assemblage. This is an important omission, as changes in stand structure and composition over the decades following harvesting may influence wildlife utilization of forest stands. We developed an observational study to evaluate forest bird response to stand structural development multiple decades following harvesting with differing levels of overstory density reduction, and in unmanaged stands. We focused, in particular, on songbirds and cavity excavating species associated with old-growth Douglas-fir forests, and the active management strategies that we examined – thinning and structural retention harvesting – were selected based on their potential to accelerate stand structural development. Our 18 mature stands (age range: 107–187 years) were located in the western hemlock zone of western Oregon, and bird surveys were conducted an average of 41 years and 22 years post-harvest in thinned and retention harvest stands, respectively. Poisson generalized linear models were formulated to evaluate the effects of management condition on forest birds. Although five species were associated with specific management conditions, relative abundance was not statistically different between unmanaged, thinned and retention harvest stands for a majority of the species in our analysis. Species richness was also relatively invariant across management conditions. Our results do suggest that retention harvesting adversely affects some songbird species associated with old-growth forests – Pacific-slope flycatcher ( Empidonax difficilis ) and Pacific wren ( Troglodytes pacificus ) for example - but our findings also indicate that retention harvesting has long-term benefits for birds associated with early-successional forests. This includes migrant species that have experienced significant population declines in Western North America over recent decades, such as Wilson’s and MacGillivray’s warblers ( Cardellina pusilla and Geothlypis tolmiei , respectively). Overall, our results imply that for many species of forest birds, including those associated with old-growth forests, managers have some flexibility in overstory density management where long-term species persistence is an objective. Equally significant, our results provide strong support for the application of variable retention harvesting as a tool for the conservation of bird species associated with early-successional forests, while also reinforcing the value of mature structural legacies for bird species associated with late-successional forests. While also reinforcing the value of mature structural legacies for birds associated with late-successional forests.

Oregon

Diet composition of Fishers (Pekania pennanti) reintroduced on the Olympic Peninsula, Washington

Knowledge of diet composition can inform management strategies and efforts to recover endangered carnivore populations in vacant portions of their historic ranges. One such species, the Fisher ( Pekania pennanti ), was extirpated in Washington State prior to any formal documentation of its food habits in the coastal coniferous forests of western Washington. Fisher recovery efforts in Washington, based on translocating Fishers from extant populations, have been ongoing since 2008, beginning with the release of 90 Fishers on Washington's Olympic Peninsula from 2008 to 2010. We collected fecal samples or digestive tracts from 13 Fishers opportunistically on the Olympic Peninsula from 2009 through 2013. Subsequently, we identified the species composition of each sample's contents to determine the primary foods consumed by the reintroduced Fishers. Fisher diets were diverse and dominated by mammalian prey. Contents of feces and digestive tracts of Fishers were composed primarily of Snowshoe Hare ( Lepus americanus ) remains, followed by lesser proportions of Mountain Beavers ( Aplodontia rufa ), Northern Flying Squirrels ( Glaucomys sabrinus ), Douglas Squirrels ( Tamiasciurus douglasii ), Southern Red-backed Voles ( Myodes gapperi ), shrews ( Sorex spp.), and unidentified ungulate species. The diet of Fishers comprised species that occur across a wide range of land uses and management prescriptions, including previously logged forests and mature forests that have been set aside for retention of old-growth forest characteristics. Additional study of prey abundance and Fisher foraging behaviors related to structural habitat characteristics across a gradient of land uses would provide useful insights for enhancing the effectiveness of conservation efforts to benefit Fishers in Pacific Northwest coastal forests.

Washington

Western purple martin (Progne subis arboricola) occurrence on the Siuslaw National Forest, Summer 2019

The western subspecies of the purple martin ( Progne subis arboricola ) is currently listed as a “critically” sensitive species in four ecoregions of western Oregon: Coast Range, Klamath Mountains, West Cascades, and Willamette Valley (Oregon Department of Fish and Wildlife, 2019). Importantly distinct from the abundant and widespread eastern subspecies ( Progne subis subis ), the western subspecies is of particular concern to Federal forest managers. Whereas the eastern subspecies is almost entirely dependent on artificial human-provided housing, the western subspecies continues to rely on natural cavities for nesting habitat (Bettinger, 2003). Accurate estimates of the regional abundance of the western purple martin are difficult to obtain; the most recent statewide census for Oregon, conducted in 2005, estimated the population at 1,100 pairs (Western Purple Martin Working Group, 2010). Several factors, including a small population size, loss of breeding habitat, and reductions in the number of suitable nesting sites have put populations of the western purple martin at risk throughout much of the Pacific Northwest region (Rockwell, 2019).

Oregon

Removal of cattle grazing correlates with increases in vegetation productivity and in abundance of imperiled breeding birds

Livestock grazing is the most prevalent land use practice in the western United States and a widespread cause of degradation of riparian vegetation. Riparian areas provide high-quality habitat for many species of declining migratory breeding birds. We analyzed changes in vegetation and bird abundance at a wildlife refuge in southeastern Oregon over 24 years, following cessation of 120 years of livestock grazing. We quantified long-term changes in overall avian abundance and species richness and, specifically, in the abundances of 20 focal species. We then compared the local responses of the focal species to population-scale trends of the same species at three different large spatial scales. Overall avian abundance increased 23% during the 12 years after removal and remained consistent from then through year 24. Three times as many species colonized the survey sites as dropped out. Of the focal species, most riparian woodland-tree or shrub dependent, sagebrush obligate, and grassland or meadow taxa increased in abundance or remained stable locally. As these species were generally of conservation concern, the population increases contradicted regionally declining or stable trends. In contrast, most riparian woodland-cavity nester species decreased in abundance locally, reflecting disruption of aspen stand dynamics by decades of grazing. Avian nest parasites and competitors of native species declined in abundance locally, matching regional trends. Restoring riparian ecosystems by removing livestock appeared to be beneficial to the conservation of many of these declining populations of migratory birds.

Oregon

Herbicides and herbivory interact to drive plant community and crop‐tree establishment

Land management practices often directly alter vegetation structure and composition, but the degree to which ecological processes such as herbivory interact with management to influence biodiversity is less well understood. We hypothesized that large herbivores compound the effects of intensive forest management on early seral plant communities and plantation establishment (i.e., tree survival and growth), and the degree of such effects is dependent on the intensity of management practices. We established 225 m 2 wild‐ungulate (deer and elk) exclosures, nested within a manipulated gradient of management intensity (no‐herbicide Control, Light herbicide, Moderate herbicide and Intensive herbicide treatments), replicated at the scale of whole harvest units (10‐19 ha). Vegetation structure, composition and crop‐tree responses to herbivory varied across the gradient of herbicide application during the first two years of stand establishment, with herbivory effects most evident at intermediate herbicide treatments. In the Moderate herbicide treatment – which approximates treatments applied to > 2.5 million hectares in Pacific Northwest U.S.A. – foraging by deer and elk resulted in simplified, low‐cover plant communities more closely resembling the Intensive herbicide treatment. Herbivory further suppressed the growth of competing vegetation in the Light herbicide treatment, improving crop‐tree survival, and providing early evidence of an ecosystem service. By changing community composition and vegetation structure, intensive forest management alters foraging selectivity and subsequent plant‐herbivore interactions; initial shifts in early seral communities are likely to influence understory plant communities and tree growth in later stages of forest development.

Oregon

Use of created snags by cavity‐nesting birds across 25 years

Snags are important habitat features for many forest‐dwelling species, so reductions in the number of snags can lead to the loss of biodiversity in forest ecosystems. Intentional snag creation is often used in managed forests to mitigate the long‐term declines of naturally created snags, yet information regarding the use of snags by wildlife across long timescales (>20 yr) is lacking and prevents a complete understanding of how the value of created snags change through time. We used a long‐term experiment to assess how harvest treatment (i.e., small‐patch group selection, 2‐story, and clearcut) and snag configuration (i.e., scattered and clustered) influenced nesting in and foraging on 25–27‐year‐old Douglas‐fir ( Pseudotsuga menziesii ) snags by cavity‐nesting birds. In addition, we compared our contemporary measures of bird use to estimates obtained from historical surveys conducted on the same group of snags to quantify how bird use changed over time. Despite observing created snags for >750 hours across 2 consecutive breeding seasons, we found limited evidence of nesting activity. Only 11% of created snags were used for breeding, with nesting attempts by 4 bird species ( n = 36 nests); however, we detected 12 cavity‐nesting species present on our study sites. Furthermore, nearly all nests (94%) belonged to the chestnut‐backed chickadee ( Poecile rufescens ), a weak cavity‐excavating species that requires well‐decayed wood for creating nest cavities. Our surveys also recorded few observations of birds using created snags as foraging substrates, with only 1 foraging event recorded for every 20 hours of observation. We detected 82% fewer nests and recorded 7% fewer foraging observations during contemporary field work despite spending >7.5 times more effort observing created snags relative to historical surveys. We conclude that 25–27‐year‐old created Douglas‐fir snags provided limited opportunities for nesting and foraging by most cavity‐nesting birds, and that the period of greatest use by this group occurred within 5–15 years of creation.

Journal of Wildlife Management

Long-term dynamics and characteristics of snags created for wildlife habitat

Snags provide essential habitat for numerous organisms and are therefore critical to the long-term maintenance of forest biodiversity. Resource managers often use snag creation to mitigate the purposeful removal of snags at the time of harvest, but information regarding how created snags change over long timescales (>20 y) is absent from the literature. In this study, we evaluated the extent to which characteristics of large (>30 cm diameter at breast height [DBH]) Douglas-fir ( Pseudotsuga menziesii ) snags created by topping had changed after 25–27 y. We also tested whether different harvest treatments and snag configurations influenced present-day snag characteristics. Of 690 snags created in 1989–1991, 91% remained standing during contemporary surveys and 65% remained unbroken along the bole. Although most snags were standing, we detected increased bark loss and breaking along the bole relative to prior surveys conducted on the same pool of snags. Although snag characteristics were not strongly influenced by snag configuration, we found that snags in one harvest treatment (group selection) experienced less bark loss and had lower evidence of use by cavity-nesting birds (as measured by total cavity cover) relative to snags created with clearcut and two-story harvest treatments. Our results indicate that Douglas-fir snags created by topping can remain standing for long time-periods (≥25 y) in managed forests, and that the influence of harvest treatment on decay patterns and subsequent use by wildlife is an important consideration when intentionally creating snags for wildlife habitat.

Forest Ecology and Management

Oak mistletoe ( Phoradendron villosum ) is linked to microhabitat availability and avian diversity in Oregon white oak ( Quercus garryana ) woodlands

Mistletoes are parasitic or hemi-parasitic flowering plants that parasitize woody plants around the globe. Important food and cover resources provided by mistletoes have been related to strong patterns of positive association between wildlife diversity and mistletoe density. Mistletoes also create microhabitat features known to be important to wildlife by causing deformations in their host trees. However, links between availability of mistletoe-formed microhabitat and wildlife diversity has not been well studied. We investigated this relationship by quantifying microhabitat features and avian abundance and diversity related to infection by Oak Mistletoe (Phoradendron villosum) in Oregon White Oak (Quercus garryana). Q. garryana woodlands support several avian species of conservation concern, so an understanding of the influence of mistletoe on wildlife habitat is critical. Our results suggest that 1) structural heterogeneity within tree crowns; 2) avian species richness and abundance are positively associated with mistletoe load; and 3) P. villosum fruit, available is an important food for western bluebird (Sialia mexicana) and other wildlife in late autumn and early winter. If a goal of restoration is to maintain habitat for oak-associated bird species, managers should consider the retention of some oaks hosting mistletoe.

Oregon

Mount Rainier National Park

Natural Resource Condition Assessments (NRCAs) evaluate current conditions for a subset of natural resources and resource indicators in national parks. NRCAs also report on trends in resource condition (when possible), identify critical data gaps, and characterize a general level of confidence for study findings. The resources and indicators emphasized in a given project depend on the park’s resource setting, status of resource stewardship planning and science in identifying high-priority indicators, and availability of data and expertise to assess current conditions for a variety of potential study resources and indicators. Although the primary objective of NRCAs is to report on current conditions relative to logical forms of reference conditions and values, NRCAs also report on trends, when appropriate (i.e., when the underlying data and methods support such reporting), as well as influences on resource conditions. These influences may include past activities or conditions that provide a helpful context for understanding current conditions and present-day threats and stressors that are best interpreted at park, watershed, or landscape scales (though NRCAs do not report on condition status for land areas and natural resources beyond park boundaries). Intensive cause-andeffect analyses of threats and stressors, and development of detailed treatment options, are outside the scope of NRCAs. It is also important to note that NRCAs do not address resources that lack sufficient data for assessment. For Mount Rainier National Park, this includes most invertebrate species and many other animal species that are subject to significant stressors from climate change and other anthropogenic sources such as air pollutants and recreational use. In addition, we did not include an analysis of the physical hydrology associated with streams (such as riverine landforms, erosion and aggradation which is significant in MORA streams), due to a loss of staff expertise from the USGS-BRD staff conducting the work, and human disturbance landcover issues such as the effects of roads, trails, and other anthropogenic developments due to lack of funds.

Report

Disentangling the effects of climate and landscape change on bird population trends in the western U.S. and Canada

Changes in climate are often assumed result in changes to species’ ranges, with potential impacts on natural system functioning and ecosystem services. ‘Climate envelope models’, which rely on correlations between climate and species distributions, have been used to predict the future of biodiversity under these assumptions. However, other factors including land-cover, dispersal ability and interspecific competition and facilitation may play an important role in driving species distributions and population trends either alone or in combination with climate. In an ongoing project, we used long-term data on bird distributions and abundance to develop climate envelope and land-use models for 161 species in order to provide a multi-species test of the degree to which climate envelope versus land-use models are useful in predicting species distributions and population trends of birds in forest ecosystems of the western U.S. and Canada. Our results suggest that models describing associations between climatic variables and abundance patterns can be used for some species to predict changes through time, and that changes in climate have already driven shifts in the geographic patterns of abundance of bird populations in western North America. For other species, models using land-use variables including raw remote-sensing variables may provide the best predictions for abundance change. The results of this research showing the reliability of models across multiple species will aid managers in understanding which species are most vulnerable to changes from climate, land-use change and their interaction.

Final Report

Bird-vegetation associations in thinned and unthinned young Douglas-fir forests 10 years after thinning

Quantitative associations between animals and vegetation have long been used as a basis for conservation and management, as well as in formulating predictions about the influence of resource management and climate change on populations. A fundamental assumption embedded in the use of such correlations is that they remain relatively consistent over time. However, this assumption of stationarity has been rarely tested – even for forest birds, which are frequently considered to be 'indicator species' in management operations. We investigated the temporal dynamics of bird-vegetation relationships in young Douglas-fir (Pseudotsuga menziesii) forests over more than a decade following initial anthropogenic disturbance (commercial thinning). We modeled bird occurrence or abundance as a function of vegetation characteristics for eight common bird species for each of six breeding seasons following forest thinning. Generally, vegetation relationships were highly inconsistent in magnitude across years, but remained positive or negative within species. For 3 species, relationships that were initially strong dampened over time. For other species, strength of vegetation association was apparently stochastic. These findings indicate that caution should be used when interpreting weak bird-vegetation relationships found in short-term studies and parameterizing predictive models with data collected over the short term.

Oregon

Summary of bird-survey and banding results at W.L. Finley National Wildlife Refuge, 1998-2008

With some of the best remaining examples of oak habitats in the Willamette Valley, the Willamette Valley National Wildlife Refuge Complex (WVNWRC) has been implementing restoration efforts to reverse the successional trend towards Douglas-fir and maple that is threatening existing oak woodlands. The restoration work has been considered a model for other public and private efforts within the Willamette Valley, and has been showcased through the Oregon Oak Communities Working Group (http://www.oregonoaks.org). Although many oak restoration projects have been initiated over the last several years, and grant recipients typically identify wildlife species that are likely to benefit from their project, measures of success have not included the actual response of wildlife, such as a change in the probability of species occurrence or abundance. Monitoring in the WVNWRC has so far been limited to vegetative and structural changes within the plant community. Hagar and Stern (2001) identified bird species occurring in Willamette Valley oak woodlands that might be expected to benefit from such restoration efforts, including an endemic subspecies of the White-breasted Nuthatch (see Appendix 1 for scientific names of bird and plant species listed in this document), and the Acorn Woodpecker, both of which are species of concern in Oregon. However, empirical data documenting responses of bird assemblages to restoration actions are needed. The goal of this study was to document the effects of a restoration project in an Oregon White Oak woodland on Pigeon Butte in the W.L. Finley National Wildlife Refuge. Restoration treatments on Pigeon Butte include the removal of shade-tolerant tree species (primarily big-leaf maple and Douglas-fir) to reduce competition with oak trees and to return the stand to a more open structure. The objectives of this ongoing study are to compare abundance, survival, and productivity of diurnal songbird species before and after application of these restoration treatments. Monitoring these vital rates will provide crucial information about the effects of management on survival and productivity (DeSante and Rosenberg, 1998). Therefore, a constant-effort mist-netting project was continued in 2007 and 2008 that had previously collected songbird demographic data at Pigeon Butte from 1998 to 2002. Point-count surveys were conducted in the woodland to build on historical data available for the site (Anderson, 1970; Hagar and Stern, 2001). The data reported here represent 5 years of point count surveys and 6 years of banding before restoration treatment, but only one post-treatment sampling season. Continued monitoring of the bird population is recommended to determine both short-term effects and long-term trends following the habitat alterations that result from restoration treatment.

Oregon

Young Stand Thinning and Diversity Study: Response of Songbird Community One Decade Post-Treatment

The response of songbird assemblages to commercial thinning is likely to change as vegetation develops over time after thinning. The influence of thinning intensity and pattern on the timing of transitions in bird community composition following thinning is of interest to managers when a goal is to maintain diversity and accelerate the development of late-seral forest structure. We investigated changes in the composition of songbird assemblages and density of individual species from 2 years before to 12 years after experimental thinning of 40-year-old stands dominated by Douglas-fir (Pseudotsuga menziesii) in the Oregon Cascades. Species richness, and density for five species and the neotropical migrant group were greater in thinned than in unthinned control stands over all post-treatment years of the study. Similarly, three species maintained a negative response to thinning over the post-treatment period. The initial positive influence of thinning was no longer in evidence a decade after harvest for five species. Of seven species with an initial negative response to thinning, three indicated recovery towards pre-treatment densities by the end of the most recent post-treatment survey. Our study is one of the first to document long-term effects of commercial thinning on forest songbird assemblages in the Pacific Northwest.

Open-File Report