Effects of hunting on survival and habitat use by American Woodcock on breeding and migration areas
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Geology topics
Publications and source records attributed to J. R. Longcore.
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Records of predaceous diving beetles (Coleoptera: Dytiscidae) collected in Maine are summarized. These records are augmented by field surveys of beetles in Aroostook Co., Maine during 1993-95. Keys to subfamilies are presented with color plates for selected species. A list of diving beetles that have been collected near Maine (state or province) is presented so that investigators will know what additional species might be expected in Maine. Basic taxonomy is presented to facilitate use of keys.
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We alternately marked 80 juvenile, female mallards (Anas platyrhynchos), half with transmitters and bands and half with only bands in 1990 and 1991. Survival of radio-marked ducks was monitored daily and then summarized weekly for the periods 31 August - 6 December, 1990 (weeks 5-18) and 11 September - 6 December, 1991) (weeks 6-18). The pattern of survival throughout the staging period did not differ (P = 0.785) between 1990 and 1991. Survival at the end of the staging period (week 18) was low in both years (0.379 in 1990; 0.282 in 1991; and 0.337 for a combined estimate). Only 2.5% (1990) and 7.5% (1991) of the bands of the banded-only sample were recovered. Fates of the ducks were associated with hunter opportunity and are evaluated as related to body mass, flock size and duck behavior as determined from hunter questionnaires.
A standard method for characterizing nest sites and concealment (visibility of orange decoy and percent overhead cover measured by densiometer) was used to obtain characteristics of 36 nest sites of black ducks in Maine, 1978-89. Nest locations were represented by cutover areas (10), islands (6), bogmat (5), emergent meadow (5), emergent wetland (3), stream floodplain (3), hardwood forest (1), conifer forest (1), mixed forest (1) and ephemeral pond (1). Within these locations nests were found in shrub clumps (1), under conifers (6), on hummocks (6), on ericaceous mats (4), under a clump of hardwood trees (4), under woody slash (3), on an emergent herbaceous clump (1), on a boulder (1) and on a muskrat house (1). After excluding 7 nests disturbed by investigators, 22 (76%) of the 29 remaining nests were successful nests. Nests in upland cuts were especially successful (9 of 9) and success was 75 - 100% at most locations, but both nests along stream floodplain were abandoned because of human disturbance. Unsuccessful nests were usually closer to ponds (2.5 times) or streams (6.6 times) and often at land water interfaces, i.e., islands and bogmat. Nests under conifers (5 of 6) and woody slash (3 of 3) usually were successful. The combination of low nesting density and isolation of nests in upland cutover areas (successful nests averaged nearly 3 times farther from roads) seem to influence black duck nest success.
Numbers of wetlands and amount of surface water (ha) within a 105 km2 area increased substantially between 1958-60 and 1978-80 in southcentral Maine. Conversely, in 1978-80 mean numbers of black duck pairs (11.9/100 ha surface water) and broods (10.8/100 ha surface water) were lower than for 1958-60 (pairs = 23.3; broods = 13.3) for wetlands surveyed in both periods. Differences in pairs are discussed in terms of wetland changes and in terms of differences in numbers of black ducks counted in the Atlantic Flyway mid-winter inventory for comparable years.
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The authors investigated the survival of breeding male American woodcock in Washington County, Maine. A total of 150 birds were radio-arked during 1987-1989 and relationships were evaluated between survival, body mass, predation, weather, and habitat use. Survival varied from 0.690 (1989) to 0.924 (1988), with a 3-year mean (95% confidence interval) of 0.789 (0.693-0.885)
Locations, habitat descriptions, and collection dates are listed for new records of 4 genera and 12 species of predaceous diving beetles (Coleoptera: Dytiscidae) in Maine. Previously, 17 genera and 53 species of the aquatic beetle were reported from Maine.
We used radiotelemetry to determine habitat use and movements of 38 female juvenile American black ducks (Anas rubripes) on the north shore of the St. Lawrence estuary, Quebec, from 28 August to 15 November 1991. Ducks separated into three groups based on habitat use: inland, estuarine, and those using a mixture of habitats. Ducks using mixed habitats used the greatest variety of habitat types, flew the greatest distances and most often between night roosts and day foraging areas, and were unlikely to be shot. The mean distance flown between night-use and day-use areas for all ducks increased nearly 50% after the hunting season began (overall mean = 6104 m, range 1500 - 26384 m). Mean home range size was 27.6 t 6.5 (SE) km2. Ducks exhibited stronger fidelity to wetlands used at night than to those used during the day. Ducks that were shot spent a high percentage of their time on the estuary (90.1 t 7.4 %) and exhibited high fidelity to a day-use area before the hunting season (73 t 7.3 %).
We used activity traps as designed by Riley and Bookhout (1990. Wetlands) to sample aquatic invertebrates as part of a study to characterize wetlands on a forested and an agricultural landscape (ca. 1,000 mi'2) in northern. Maine. Eight wetlands (5 from agricultural and 3 from forested landscapes) were sampled at random from 50 wetlands surveyed for waterfowl broods. At the landscape level, insect abundance (mean no./ trap), fish abundance (mean no./trap), percent vegetation, and water chemistry variables (pH, ANC, SPCOND, Ca, Mg, K, Na, Cl) were different between landscapes. Furthermore, nearly as many fish (2,112) were caught as were insects (2,443); 47% of the 332 traps contained fish, but 84 traps accounted for 94% of the fish caught. When >4 fish were in a trap fewer insects were in the trap. Differences in water temperature among wetlands and differences in rates of escape among insect orders affected the number of different taxa caught. Until capture success of activity traps is better understood, results from activity traps should be used with care.
We critiqued previous work on the biology and management of the American woodcock (Scolopax minor) on the breeding grounds. We determined that little is known about habitat variables and weather extremes that may limit the population. Most investigators who attempted to define habitat requirements of the woodcock used inadequate sample sizes, limited the duration of their studies, did not account for effects of weather, or failed to adequately measure habitat variables. Furthermore, the effects of hunting on local or regional populations has never been adequately studied. We concluded that obtaining data to understand the biology of the woodcock and the effects of hunting is essential before managers can reverse the long-term decline of woodcock numbers.
We describe methods for the study of the American woodcock (Scolopax minor) at the Moosehorn National Wildlife Refuge. Described are techniques for the capture of particular age-sex cohorts during different times of the year, the aging and sexing, the evaluation of habitat components, and the estimation of population size. We describe successful and tried but rejected techniques with radio-telemetry for the study of survival and habitat use.
During spring 1986-89, we equipped 175 male and 89 female American woodcocks (Scolopax minor) with radio transmitters. Radio-marking had little effect on behavior; within 1 day of marking, 37 of 64 (58%) displaying males were dominant and within 7 days, 138 of 157 (88%) were dominant. All females marked before nesting proceeded to nest, and marked females with broods remained with broods after release. Dominance of males declined from 73% in April to 69% in May and to 26% in June as breeding activity waned. In all years, after-second-year (ASY) males were dominant more often than second-year (SY) males (67.5% vs. 58.9%). ;Most males displayed at more than one (range = 2-12) site. Distances that males moved between the primary singing ground and subsequent singing grounds averaged 775 m in all years and ranged from 618 m (1986) to 966 m (1988). Females visited males at singing grounds throughout the breeding cycle; some females visited more than one site. During prenesting, females remained in daytime covers during the crepuscular period (55%), flew to different feeding covers (22%), visited singing grounds (14%), or flew from daytime covers to unknown locations (9%). During nesting, females left nests during the crepuscular period (72%) and moved to singing grounds (5%), to feeding areas (59%), and to undetermined locations (7%). Females with broods remained with their broods during the crepuscular period (62%), flew to feeding areas (30%), visited singing grounds (1%), or flew to undetermined locations (6%). The woodcock mating system is similar to a resource-based polygyny. Males compete for singing grounds near high-quality nesting areas. The fitness of males is expressed by dominance at more than one singing ground. Woodcocks are not monogamous; females may visit more than three different males during a single courtship period but do not visit males regularly. Most females leave nests during the crepuscular period to feed in a different cover.
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We examined the use of 87 palustrine and lacustrine wetlands by nongame water birds in central and eastern Maine using 3,527 h of observation (1,501 visits) made during April-August, 1977-85. Wetlands used by 15 species of water birds were distinguished from those not used, according to 20 habitat features. The species were the common loon (Gavia immer) , pied-billed grebe (Podilymbus podiceps), double-crested cormorant (Phalacrocorax auritus), American bittern (Botaurus lentiginosus), great blue heron (Ardea herodias), green-backed heron (Butorides striatus), osprey (Pandion haliaetus), bald eagle (Haliaeetus leucocephalus), northern harrier (Circus cyaneus), Virgima rail (Rallus limicola), sora (Porzana carolina), spotted sandpiper (Actitis macularia), common snipe (Gallinago gallinago), herring gull (Larus argentatus), and belted kingfisher (Ceryle alcyon). Predictive models of habitat use were developed for each species. Water birds were classified by similarity of habitats used, and species use was contrasted by wetland type. Smaller, isolated wetlands were used by fewer (P < 0.05) species than larger wetlands in complexes; many species had large area-requirements (pied-billed grebe, common loon, herring gull, double-crested cormorant, bald eagle) or preferred to use wetlands near other wetlands (common loon, herring gull, great blue heron, spotted sandpiper, osprey, bald eagle). Wetland area contributed more to overall variation in species richness on wetlands than wetland isolation, although on small wetlands (<3.6 ha) isolation was a better predictor of species richness than wetland area. Wetlands with intermediate amounts (33-66%) of emergent vegetation supported more species (P< 0.05) than closed (>66%) or open (<33%) wetlands. Low pH typified wetlands used by large-bodied piscivores (common loon, cormorant, osprey). Other water birds were associated with more densely vegetated, chemically buffered wetlands. Habitat features associated with wetland use by each waterbird species are reported, as are numerical responses of waterbird populations to wetland features and estimates of annual variation in habitat occupancy. Lacustrine wetlands supported a distinct, low diversity community of water birds, including most fish-eating species. Waterbird diversity at forested palustrine wetlands was intermediate between lacustrine communities and more species-rich assemblages at palustrine emergent and scrub-shrub wetlands. Regional variation in wetland characteristics and water bird use was associated with surficial geology, soils, and management practices. Management for nongame water birds in Maine should consider providing emergent and aquatic-bed vegetation with variable cover-to-water ratios, accommodating species-specific habitat needs, focusing on species of restricted distribution and low abundance, and maintaining wetland complexes. Bird use and habitat information from 87 wetlands and models of habitat selection for each species are provided in appendixes.
We monitored the movements of 97 female and 15 male juvenile American black ducks (Anas rubripes) in the vicinity of Moosehorn National Wildlife Refuge in eastern Maine and southwestern New Brunswick from September through mid- December, 1985-1987.. Movements were described by estimating home ranges and radial movements from the primary roost marsh. Overall home range sizes averaged 4987 ha (range 54 - 28 070 ha), and maximum distances moved from the roost averaged 9.9 km (range 0.9-42.8 km). Home ranges were linear (linearity index 2.8), and home range area and distance of movements from the roost both increased monthly. Ducks that used >15 % riverine habitat had larger home ranges and moved greater distances from the roost than ducks using <15% riverine wetlands. Movements did not differ between ducks usually alone and those usually in flocks. Juvenile black ducks moved in small flocks and were often alone (34% of 355 observations). Most ducks showed fidelity to one roost-marsh complex from September until migration in late November. Daily and seasonal movement patterns were similar to those predicted for refuging systems.