Geology topics

Casey A. Pennock

Publications and source records attributed to Casey A. Pennock.

At least 19 recordsLinked to original sources

USGS2026

How tough a passage? Nonnative fish entrainment survivability at a major dam in the western USA

The Colorado River downstream of Glen Canyon Dam (GCD) in the southwest USA demonstrates anthropogenic modification including altered flow and temperature regimes and established nonnative species in upstream reservoirs. Water overallocation and megadrought are contributing to new potential invasion pathways for nonnative fishes through dam releases. In this study, we quantified factors that influence survival of fish entrained at GCD, a pathway for nonnative fishes to leave Lake Powell and enter the Grand Canyon as reservoir levels remain near historic lows since the reservoir initially filled. We measured conditions of entrainment through GCD using a mechanical Sensor Fish device and evaluated potential barotrauma by comparing our results to previous studies. Under measured conditions, a fish's gas bladder would expand by a factor of 2.7 due to entrainment, potentially resulting in injury from barotrauma. We estimated members of Centrarchidae may survive entrainment at problematically high rates across all water levels where entrainment is expected. Minimizing nonnative fish entrainment through GCD is best achieved through maintenance of water levels higher than those in recent years, potentially decreasing both entrainment likelihood and survival.

Arizona↗

USGS2026

Ecological stability of zooplankton communities in changing arctic lakes over three decades

In the Arctic, rapid warming is altering disturbance regimes, creating an urgent need to assess the ecological stability of these relatively pristine ecosystems. The Arctic landscape has thousands of lakes that are experiencing both gradual, persistent disturbances (e.g., warming) and increased frequency and intensity of pulse disturbances (e.g., land-surface failures). To assess the ecological stability of lake communities in a changing environment, we analyzed long-term data (14–39 years) on zooplankton communities and environmental variables from 25 lakes in arctic Alaska. We characterized long-term change and evaluated the resistance and recovery of communities to natural or experimental disturbances. We hypothesized that reference lakes would demonstrate resistance to background environmental change, represented by variable but consistent relative abundances over time, while manipulated or disturbed lakes would show recovery after the disturbance ceased, as represented by directional changes in relative abundance with return to pre-disturbance conditions. Environmental variables including water conductivity, chlorophyll- a , NO 3 − , pH, dissolved oxygen, light extinction, and soluble reactive phosphorus changed directionally through time, suggesting press effects due to climate change. Zooplankton communities were resistant to environmental change in undisturbed lakes, but, despite monitoring lakes for up to 28 years after disturbance, few lakes showed evidence of recovery to pre-disturbance conditions. This lack of recovery may indicate shifting baselines due to environmental change. The variation we did observe was largely due to shifts in species relative abundance rather than in species turnover, and this variability was related to lake depth. Zooplankton communities and environmental variables displayed more interannual variation in shallow (depth ≤ 6 m) lakes than in deep lakes. Overall, our results demonstrate that arctic zooplankton communities have high resistance to the effects of climate change but potentially low recovery or long recovery times to pulse disturbances.

Alaska↗

USGS2026

Bank-dwelling beavers contribute to the wood regime in a dryland river

Our findings suggest that the influence of dwelling beavers on large wood budgets may substantially determine the structure and function of medium to large rivers, particularly in dryland systems such as the White River. Recognizing the role of bank-dwelling beavers expands our understanding of biotic drivers of riverscape complexity and provides new opportunities to integrate ecology with fluvial geomorphology in managing and restoring dynamic river corridors.

Utah↗

USGS2026

Invasion potential of nonnative fishes through a large western dam into an iconic and vulnerable ecosystem

Native fishes face imperilment due to habitat loss and fragmentation, megadrought, invasive species, and synergies amongst threats. These threats coalesce at Glen Canyon Dam (GCD), which impounds the Colorado River to create Lake Powell, a reservoir inhabited by a suite of nonnative fishes. Water level in Lake Powell has declined precipitously due to water overallocation and megadrought, increasing the risk of nonnative fish entrainment through the dam, as the penstocks draw water from the epilimnion. Our goal was to characterize the distribution and abundance of nonnative fishes in the vicinity of GCD and identify environmental conditions that may increase entrainment. We used active and passive sampling, hydroacoustics, and acoustic telemetry to assess the distribution and abundance of fishes. When the reservoir was stratified fish remained in the epilimnion, were present near the GCD penstocks, and thus at high risk of entrainment. When the reservoir was mixed, fish generally occupied shallower, warmer areas further from GCD, but this varied among species. The distribution of nonnative fish and accompanying risk of entrainment varied by season and epilimnion depth. Retaining more water in Lake Powell to draw water from below the thermocline would reduce the risk of nonnative fish entrainment into Grand Canyon.

Arizona, Utah↗

USGS2025

Rapid recovery of an arctic lake ecosystem from a pulse disturbance caused by thermokarst failure

Due to rapid climate change, arctic ecosystems are experiencing an increase in disturbances including localized land-surface failures caused by melting ground ice (thermokarst failures). These failures result in the mass transport of sediment and organic materials into surface waters, with the potential to dramatically alter aquatic ecosystem function and biotic interactions. We coupled direct comparisons and long-term data of a suite of abiotic and biotic variables in a thermokarst-impacted lake and nearby reference lake to assess the impacts of thermokarst failure. After the thermokarst failure and relative to long-term averages, water transparency was substantially reduced. We hypothesized there would be subsequent changes to lower trophic levels and profound declines in fish foraging efficiency. However, these characteristics were within the range of natural variability and/or rapidly recovered to values within the range of long-term variability. In addition, although there was limited evidence of taxa-specific changes, we did not observe any strong changes in the total relative densities, growth rates, or composition of the bacterioplankton and zooplankton communities, benthic macroinvertebrates, or changes in fish diet, that could be attributed statistically to the thermokarst event. In sum, the thermokarst disturbance had substantial effects on water transparency and some lower trophic levels, which surprisingly were not manifested in higher trophic levels. Overall, the lake ecosystem appeared resistant to thermokarst disturbance with rapid recovery within two years after the disturbance.

Alaska↗

USGS2025

The effect of turbidity on foraging by prerostrum juvenile Paddlefish

Objective A previous study evaluating restoration success of Paddlefish Polyodon spathula suggested that excessive turbidity in lakes and rivers may inhibit foraging by juveniles prior to the development of the rostrum. Although a Paddlefish's rostrum, which contains electroreceptors, helps the fish to locate zooplankton prey, the prerostrum stage lacks many of these electroreceptors, potentially affecting foraging in highly turbid waters. To evaluate this hypothesis, we conducted a series of laboratory experiments in small aquaria by varying the level of turbidity measured as Secchi tube depth: >95 (clear tap water), 40, 20, 10, or 5 cm. Methods For each foraging trial, approximately 300 zooplankton were added to the aquaria, followed by three postlarval Paddlefish, which were allowed to feed on the zooplankton for approximately 18 h. After the 18-h period, Paddlefish were removed and dissected and the zooplankton in the gut were counted to quantify foraging success. Results From three experimental trials, we observed statistically significant nonlinear relationships for two trials, which included the largest fish tested. The smallest Paddlefish size-class showed no significant trend in foraging among turbidity treatments. However, the subsequent trials, which contained larger Paddlefish, both showed unimodal responses, with the highest foraging observed at moderate (30–50-cm Secchi tube depth) turbidity levels. Conclusions High turbidity and excessively clear water both appear to affect foraging by juvenile Paddlefish before the rostrum is fully developed, depending on fish size. Understanding this relationship can further influence management and restoration projects directed at this species.

Transaction of the American Fisheries Society↗

USGS2025

Shrinking channels, growing threats: Habitat degradation from channel narrowing and invasive vegetation in three dryland rivers

Water development and the proliferation of invasive riparian vegetation have led to widespread habitat loss and simplification of rivers in the western United States, contributing to the imperilment of native fishes. Here, we quantify channel narrowing and vegetation encroachment, which are conspicuous indicators of riverine habitat alteration, along ∼400 km of three dryland tributaries of the upper Colorado River. We conducted a comparative analysis of aerial photographs between the 1930s and 2010s/2020s time periods using visual interpretation and used Light Detection and Ranging (LiDAR) data along with Object-Based Image Analysis (OBIA) to quantify canopy cover of woody riparian species. All three rivers underwent substantial channel narrowing, coinciding with a general decrease in spring floods over time. However, the extent of narrowing varied among the rivers (78 %, 73 %, and 29 %) with greater narrowing corresponding to larger reductions in spring flows. In contrast, contemporary woody cover was similarly high among all three rivers (39 %, 41 %, and 36 %), and a woody vegetation analysis we conducted for one river indicated a substantial increase in vegetation along the active channel (4 %–74 %). These findings underscore a common pattern observed in rivers throughout the basin, where river channels often undergo narrowing and encroachment by invasive vegetation following dam construction and/or decreases in flows, ultimately leading to habitat simplification, with negative implications for native fishes and other riparian biota. Our findings also emphasize that, even in the presence of nonnative vegetation establishment, preserving or restoring large magnitude and long duration floods can help conserve diverse habitat in dryland rivers.

Arizona, Colorado, New Mexico, Utah↗

USGS2025

Using distribution and movement of invasive channel catfish Ictalurus punctatus in the San Juan River to inform removal efforts

Invasive channel catfish ( Ictalurus punctatus ) pose threats to native fishes through predation and competition. A channel catfish removal program in the San Juan River was implemented in the 1990s, but exploitation was insufficient to cause population declines. Because knowledge of movement timing and spatial aggregations of channel catfish could inform future removal efforts and increase exploitation rates, we used active and passive radio telemetry in 2023 and 2024 and mark-recapture data from 2011–2015 to assess seasonal patterns in their distribution and movement. Additionally, spawning periodicity was identified through water temperature and activity data recovered from adults implanted with archival transmitters and by estimating back-calculated spawning dates obtained via otoliths pulled from young-of-year fish. We observed limited upstream movement ( n = 25/528 radio-tagged fish > 10 river kilometers) prior to daily mean water temperatures reaching 21°C followed by upstream movement after water temperatures exceeded 21°C. Generalized linear mixed models suggested a positive relationship between movement rates and total length and a modal relationship between movement rates and water temperature with a predicted maximum movement at 20.8°C. Movement rates of females were predicted to be 2.4x greater than males. Mark-recapture data confirmed limited movement with an upstream bias during summer. Modal activity patterns observed in 10 of 12 archive-tagged fish peaked at an average of 21.3°C. We found little evidence of aggregations, but synchronized upstream movement likely cued by water temperatures ~ 21°C may provide the opportunity for resource managers to use passive sampling techniques to target channel catfish during upstream spawning movements.

Research Square↗

USGS2024

Movement patterns of a small-bodied minnow suggest nomadism in a fragmented, desert river

Background Unfettered movement among habitats is crucial for fish to access patchily distributed resources and complete their life cycle, but many riverscapes in the American Southwest are fragmented by dams. The federally endangered Rio Grande silvery minnow ( Hybognathus amarus , RGSM) persists in a fragmented remnant of its former range (ca. 95% range reduction), and its movement ecology is understudied. Methods We tracked movements of hatchery-reared RGSM, tagged with passive integrated transponder tags, using stationary and mobile antennas from 2019–2022. We quantified probability of movement and total distance moved by RGSM released above and below a dam. We then assessed how well two prevailing riverine movement theories (i.e., restricted movement paradigm [RMP] and colonization cycle hypothesis [CCH]) explained RGSM movement patterns. Results We detected 36.8% of released RGSM (n = 37,215) making at least one movement. Movements were leptokurtic and substantially greater than expected based on the RMP for both stationary (1.7–5.9 m) and mobile (30.3–77.8 m) individuals. On average, RGSM were detected at large for 75 days and moved a total of 12.2 rkm within a year. The maximum total distance moved by RGSM was 103 rkm. Similarly, we observed a multimodal distribution of detected range sizes with a mean detected range of 2.4 rkm and a maximum detected range of 78.2 rkm. We found little support for an upstream movement bias, as expected under the CCH, and most movements (74%) were directed downstream. Conclusions Our data suggest RGSM are highly mobile, with the ability to make long-distance movements. Neither movement theory adequately described movement patterns of RGSM; instead, our findings support a nomadic movement pattern and an apparent drift paradox matching recent studies of other pelagic-broadcast spawning minnows who persist upstream despite experiencing downstream drift as larvae. Resolution of the drift paradox may be achieved through further, targeted studies into different aspects of the species’ life history. Quantification of RGSM movement provides crucial insights into the species’ movement ecology and may help define the appropriate scale of recovery efforts.

New Mexico↗

USGS2024

Wood you believe it? Experimental addition of nonnative wood enhances instream habitat for native dryland fishes

Flow alteration and riparian vegetation encroachment are causing habitat simplification with severe consequences for native fishes. To assess the effectiveness of enhancing simplified habitat in a large dryland river, we experimentally added invasive wood at 19 paired treatment and reference (no wood added) subreaches (50–100 m) within the main channel of the San Juan River. Using a before-after-control-impact design, we sampled fishes and macroinvertebrates, and quantified habitat complexity. After wood addition, total native fish densities were 2.2× higher in treatments compared with references, whereas total nonnative fish densities exhibited no response. Macroinvertebrate densities were 6.8× higher, and habitat complexity increased in treatments. Counts of geomorphic features in treatments increased from 1 to a maximum of 11 following wood addition, while the number of features in references remained unchanged. Wood addition has potential to instigate natural riverine processes, ultimately enhancing native fish habitat by increasing macroinvertebrate densities and habitat complexity in dryland rivers. Water overallocation and increasing aridity will continue to challenge efforts to improve habitat conditions with environmental flows alone, and managers might consider integrating non-flow alternatives like addition of abundant, invasive wood to reduce habitat simplification.

New Mexico, Utah↗

USGS2024

Translocation in a fragmented river provides demographic benefits for imperiled fishes

Fragmentation isolates individuals and restricts access to valuable habitat with severe consequences for populations, such as reduced gene flow, disruption of recolonization dynamics, reduced resiliency to disturbance, and changes in aquatic community structure. Translocations to mitigate the effects of fragmentation and habitat loss are common, but few are rigorously evaluated, particularly for fishes. Over six years, we translocated 1215 individuals of four species of imperiled fish isolated below a barrier on the San Juan River, Utah, USA, that restricts access to upstream habitat. We used re-encounter data (both passive integrated transponder tag and telemetry detections and physical recaptures) collected between 2016 and 2023, to inform a spatially explicit multistate mark–recapture model that estimated survival and transition probabilities of translocated and non-translocated individuals, both below and above the barrier. Individuals of all four species moved large (>200 km) distances upstream following translocation, with the maximum upstream encounter distance varying by species. Results from the multistate mark–recapture model suggested translocated fish survived at a higher rate compared with non-translocated fish below the barrier for three of the four species. Above the barrier, translocated individuals survived at similar rates as non-translocated fish for bluehead sucker ( Catostomus discobolus ) and flannelmouth sucker ( Catostomus latipinnis ), while survival rates of translocated endangered Colorado pikeminnow ( Ptychocheilus lucius ; mean, 95% CI: 0.75, 0.55–0.88) and endangered razorback sucker ( Xyrauchen texanus ; 0.86, 0.75–0.92) were higher relative to non-translocated individuals (Colorado pikeminnow: 0.52, 0.51–0.54; razorback sucker: 0.75, 0.74–0.75). Transition probabilities from above the barrier to below the barrier were generally low for three of the four species (all upper 95% CI ≤ 0.23), but they were substantially higher for razorback sucker. Our results suggest translocation to mitigate fragmentation and habitat loss can have demographic benefits for large-river fish species by allowing movements necessary to complete their life history in heterogeneous riverscapes. Further, given the costs or delays in providing engineered fish passage structures or in achieving dam removal, we suggest translocations may provide an alternative conservation strategy in fragmented river systems.

Colorado, New Mexico, Utah↗

USGS2023

Density-dependent processes and population dynamics of native sculpin in a mountain river

Understanding the processes governing population dynamics is important for effective conservation and environmental management. Disentangling the relative role of density-dependent versus density-independent processes on population dynamics is often made difficult by the inability to control for abiotic or biotic factors, but long-term datasets are invaluable in this pursuit. We used a 14-year dataset from the Logan River, Utah, to assess long-term trends in abundance and evidence of density-dependent and density-independent effects on population dynamics of Paiute sculpin ( Cottus beldingii ) across six sites. Additionally, we evaluated the feeding ecology of sculpin over 4 years. Sculpin densities generally increased from upstream to downstream, and the annual per capita rate of increase was negatively and significantly correlated with sculpin density at four of six sites. We observed a negative relationship between total gut content and sculpin density but did not observe a negative relationship between relative condition and density. Sculpin displayed a generalist feeding strategy, and interannual differences in diet composition appeared to be influenced by interannual differences in flow, particularly years with higher magnitude flow. The observed spatial patterns in sculpin abundance throughout the watershed matched those of invasive brown trout ( Salmo trutta ), the top piscivore in the Logan River, and likely represent affinities for the suite of ecological conditions associated with downstream sections of the Logan River. Our results suggest that sculpin populations are regulated largely by density-dependent processes and match those from other studies on sculpin population dynamics including a range of species and habitats that differ vastly in abiotic conditions.

Ecology of Freshwater Fish↗

USGS2023

A perched culvert and natural obstructions limit fish dispersal in an intermittent prairie stream

Poorly constructed road crossings block upstream movement of fish into stream reaches that provide critical habitat or connect isolated populations. Although removing these barriers is often a conservation priority, quantifying fish passage following removal has not been well studied, particularly in intermittent streams. In this study, we sought to understand how barriers influence the dispersal of fishes in intermittent prairie streams. We used passive integrated transponder tags and antenna stations to quantify fish movement of 3 prairie-stream fishes (Central Stoneroller Campostoma anomalum [Rafinesque, 1820], Southern Redbelly Dace Chrosomus erythrogaster [Rafinesque, 1820], and Creek Chub Semotilis atromaculatus [Mitchill, 1818]) through a road crossing in an intermittent prairie stream for 3 y before and 3 y after removing a perched culvert. We verified that no upstream fish movement occurred through the culvert, despite large aggregations of tagged fish in the intermittent reach below the culvert. In contrast, tagged individuals of all 3 species were detected above the road crossing in each of the 3 y following removal. We also tracked the recovery of these species, plus the Orangethroat Darter Etheostoma spectabile (Agassiz, 1854), following a severe drought in 2018 in 2 spring-fed tributary reaches, 1 without a downstream barrier and 1 with the removed downstream culvert. Surveys of the tributary reaches showed rapid recovery of fish densities following drought in the tributary without a downstream barrier. However, recovery at the site above the removed culvert appeared to be limited by a natural waterfall. Our observations suggest barrier removal allowed access to spawning habitat within the intermittent reach immediately upstream of the site, but other obstacles or shear distance to perennial spring-fed tributary reaches also limited recolonization following severe drought. Increased connectivity between perennial and intermittent reaches in prairie streams likely benefits fishes by increasing their resilience following disturbance and providing habitat during critical life stages.

Kansas↗

USGS2022

Failure to achieve recommended environmental flows coincides with declining fish populations: Long-term trends in regulated and unregulated rivers

Dams can be operated to mimic components of the natural flow regime to minimise impacts on downstream ecosystems. However, infrastructure, societal needs, water management, and catchment runoff constrain which and when flow regime attributes can be mimicked. We compared fish assemblage responses, including native and non-native species, over 2 decades of managed environmental flows to those in a river retaining a relatively unaltered flow regime. Both of these arid-land rivers are within the overallocated Colorado River basin and have experienced declines in catchment runoff over the past 20 years. We predicted that fish–flow relationships would be conserved across time and between managed and unmanaged rivers. Declines in flow in both rivers coincided with declines in some native fishes, and more native and non-native fish species exhibited declines in the managed river than in the unmanaged river. Our ability to detect previously documented native fish–flow relationships diminished in the managed river system because established environmental flow targets were not met due to water management, but we detected these fish–flow relationships in the unmanaged river. Our results suggest declining catchment runoff and increased consumptive water use could reduce the effectiveness of environmental flows that have lower priority in most years.

Freshwater Biology↗

USGS2022

Effective conservation of desert riverscapes requires protection and restoration of in-stream flows with rehabilitation approaches tailored to water availability

Desert riverscape rehabilitation practitioners must contend with compounding effects of increasing human water demand, persistent drought, non-native species establishment, and climate change, which further stress desert riverine ecosystems such as rivers in the Colorado River basin, United States. Herein, we provide our perspective on the importance of natural flows, large floods in particular, for successful conservation and rehabilitation of riverscapes. We present ideas developed from our experience with rehabilitation projects across multiple desert tributary rivers with varying levels of habitat degradation and water abstraction. We propose spatially extensive measures such as protection of in-stream flows, tailoring rehabilitation efforts to available annual water availability, and working with nature using low-tech process-based techniques to more completely address the mechanisms of habitat degradation, such as flow reduction and vegetation-induced channel narrowing. Traditionally, rehabilitation efforts in the Colorado River basin take place at relatively small spatial extents, at convenient locations and, largely focus on reducing non-native plant and fish species. We suggest that we need to think more broadly and creatively, and that conservation or recovery of natural flow regimes is crucial to long-term success of almost all management efforts for both in-stream and riparian communities.

Frontiers in Environmental Science↗

USGS2022

The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States and Mexico

Aim Migratory freshwater fishes are those that must access discrete habitats to complete their life cycles. Freshwater fish migrations occur around the world and provide numerous ecosystem services for humans and natural systems; however, many migratory species are in decline globally. A limiting factor to successfully conserve freshwater migratory fishes is that the migratory life histories of many species are unknown or only partially described. To provide researchers with critical and comprehensive information to conserve migratory fishes, we developed the North American Freshwater Migratory Fish Database (NAFMFD). Location Canada, Mexico and the United States. Taxon Freshwater fish. Methods To develop this database, we assigned migratory status, pattern and behaviour to a comprehensive list of freshwater fish species found throughout North America. We assembled the database which included assignments (i.e. migratory status, pattern and behaviour) as well as the sources used to make the assignments. Researchers and managers from across North America reviewed the database for completeness and accuracy on the migratory life histories of fishes. Results The database synthesizes current knowledge of migratory status, pattern and behaviour of native and non-native freshwater fishes throughout North America, including 1250 species representing 79 families and 325 genera. Results showcase the diversity of migratory life histories of freshwater fishes on the continent, including that at least 25% of North American freshwater fishes are migratory, 23% are non-migratory and 44% have undetermined migratory status. Main conclusions NAFMFD improves the quality of migratory data accessible to researchers, which supports a more holistic understanding of the threats encountered by migratory fishes, including habitat fragmentation. The approach we used in developing NAFMFD can provide guidance for developing similar databases in other regions. Collectively, our work offers new insights into the range of freshwater fish migratory life histories, stimulating a need to better understand this diversity globally.

Journal of Biogeography↗

USGS2022

High densities of conspecifics buffer native fish from negative interactions with an ecologically similar invasive

Invasive species are a leading cause for native species declines, but it remains unclear whether maintenance of high native densities influence native persistence in freshwater systems. We designed complementary laboratory and field experiments to test whether high native cutthroat trout (Oncorhynchus clarkii) densities affect competition with invasive brown trout (Salmo trutta). We manipulated density of native cutthroat trout while holding brown trout density constant. Interspecific aggressive interactions towards native cutthroat trout were 66% fewer in the highest cutthroat trout density treatment compared to the lowest density treatment. At high densities of conspecifics, cutthroat trout, sympatric with brown trout, lost 33-81% less weight and demonstrated 1.5 times greater survival in both experiments than at low conspecific densities. Cutthroat trout held at low densities in laboratory experiments experienced greater stress-related disease virulence and negative effects of harassment, apparently owing to more frequent aggressive interactions with brown trout. These results support the hypothesis that reduced per capita effects of invaders on native fish at high densities buffers native species from negative effects of invaders, reducing the potential for invasive fish to displace high density native fish populations.

Utah↗

USGS2022

Native fish need a natural flow regime

Water development has threatened the ecological integrity of riverine ecosystems. Increasing water demand, persistent drought, and climate change exacerbate the effects of habitat degradation and loss in altered systems such as the Colorado River basin. Today, biologists are challenged to identify management actions that benefit native fishes while not hindering water development or management. Herein, we discuss the importance of the natural flow regime for functioning riverine ecosystems and provide examples from four tributaries to the Green River, a major headwater branch of the Colorado River. These tributaries represent a gradient of impacts ranging from water abstraction to the point of complete seasonal desiccation to a relatively natural flow regime, and consequently have maintained different levels of instream habitat complexity and native fish persistence. Despite decades of management, endangered species lack self-sustaining populations and other native species have been extirpated from over half their ranges, which begs the question: can water development and fish conservation be balanced under current water laws and climate change-driven declines in runoff? Given the continued decline in freshwater biodiversity and abundance occurring across the globe, we contend that immediate designation of rivers with natural flow regimes as freshwater conservation areas will enhance native species recovery.

Arizona, Colorado, Utah, Wyoming↗